Khao Phrik, Rat Buri: Wordian, Thailand
collected by Waterhouse, Komalarjun, Nakinbodi, Sripatanavat 1964

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Gastropoda
Gastropoda informal uncoiled sinistral sp.
Ketwetsuriya et al. 2020
Gastropoda - Murchisoniina - Soleniscidae
Leptoptygma sp. Knight 1936
Ketwetsuriya et al. 2020
Soleniscus informal sp. 1 Meek and Worthen 1860
Ketwetsuriya et al. 2020
? Soleniscus informal sp. 2 Meek and Worthen 1860
Ketwetsuriya et al. 2020
Gastropoda - Murchisoniina - Palaeozygopleuridae
Palaeozygopleura cf. obesa Yoo 1994
Ketwetsuriya et al. 2020
(1 measurement)
Gastropoda - Murchisoniina - Lophospiridae
? Worthenia waterhousei n. sp. Ketwetsuriya et al. 2020
Ketwetsuriya et al. 2020
recombined as Yiningicus waterhousei
(1 measurement)
Gastropoda - Murchisoniina - Phymatopleuridae
Worthenia cf. crenulunula (Yoo 1994)
Ketwetsuriya et al. 2020
original and current combination Worthenia crenilunula
(1 measurement)
Gastropoda - Murchisoniina - Gosseletinidae
Ananias sp. Knight 1945
Ketwetsuriya et al. 2020
Peruvispira sp. Chronic 1949
Ketwetsuriya et al. 2020
Eoplatyzona ratchaburiensis n. sp. Ketwetsuriya et al. 2020
Ketwetsuriya et al. 2020
(1 measurement)
Gastropoda - Murchisoniina - Porcelliidae
Hesperiella cyrtocostata n. sp. Ketwetsuriya et al. 2020
Ketwetsuriya et al. 2020
(1 measurement)
Gastropoda - Trochidae
Coeloconulus panae Nützel and Nakazawa 2012
Ketwetsuriya et al. 2020
Gastropoda - Eucyclidae
? Eucycloscala sp. Cossmann 1895
Ketwetsuriya et al. 2020
Gastropoda - Bellerophontida - Bellerophontidae
Retispira lyelli (Gemmellaro 1889)
Ketwetsuriya et al. 2020
Retispira khaophrikensis n. sp. Ketwetsuriya et al. 2020
Ketwetsuriya et al. 2020
(1 measurement)
? Bellerophon sp. de Montfort 1808
Ketwetsuriya et al. 2020
Gastropoda - Bellerophontida - Euphemitidae
Euphemites sp. Warthin 1930
Ketwetsuriya et al. 2020
Warthia cf. saundersi Yochelson 1960
Ketwetsuriya et al. 2020
(1 measurement)
Gastropoda - Euomphalina - Elasmonematidae
Kimina sp. Yoo 1994
Ketwetsuriya et al. 2020
Gastropoda - Euomphalina - Microdomatidae
? Amaurotoma multispirata n. sp. Ketwetsuriya et al. 2020
Ketwetsuriya et al. 2020
(1 measurement)
Gastropoda - Meekospiridae
? Meekospira informal sp. 1 Ulrich and Scofield 1897
Ketwetsuriya et al. 2020
? Meekospira informal sp. 2 Ulrich and Scofield 1897
Ketwetsuriya et al. 2020
Gastropoda - Pseudozygopleuridae
? Hyphantozyga khaophrikensis n. sp. Ketwetsuriya et al. 2020
Ketwetsuriya et al. 2020
(1 measurement)
Gastropoda - Goniasmatidae
Goniasma sp. Tomlin 1930
Ketwetsuriya et al. 2020
? Stegocoelia informal sp. 1 (Donald 1889)
Ketwetsuriya et al. 2020
? Stegocoelia informal sp. 2 (Donald 1889)
Ketwetsuriya et al. 2020
Stegocoelia centrosinuata n. sp. Ketwetsuriya et al. 2020
Ketwetsuriya et al. 2020
(1 measurement)
Gastropoda - Coelostylinidae
Protostylus sp.
Ketwetsuriya et al. 2020
Gastropoda - Imoglobidae
Chlorozyga asiatica n. sp. Ketwetsuriya et al. 2020
Ketwetsuriya et al. 2020
(1 measurement)
Gastropoda - Procerithiidae
? Procerithium inaequetuberculata n. sp. Ketwetsuriya et al. 2020
Ketwetsuriya et al. 2020
(1 measurement)
Gastropoda - Heterostropha - Streptacididae
? Streptacis sp. Meek 1871
Ketwetsuriya et al. 2020
Gastropoda - Heterostropha - Donaldinidae
? Yoospira informal sp. 1 Bandel 2002
Ketwetsuriya et al. 2020
? Yoospira informal sp. 2 Bandel 2002
Ketwetsuriya et al. 2020
Gastropoda - Naticopsidae
Naticopsis sp. M'Coy 1844
Ketwetsuriya et al. 2020
Strophomenata - Orthotetida - Meekellidae
? Meekella sp.
2 individuals
2VV, 2DV
    = Meekella colpata Grant 1976
Grant 1976
Strophomenata - Orthotetida - Schuchertellidae
Streptorhynchus sp. King 1850
9 individuals
7VV, 2C (4 measurements)
    = Streptorhynchus khwaense Grant 1976
Grant 1976
Strophomenata - Orthotetida - Derbyiidae
? Derbyia sp. Waagen 1884
2 individuals
1DV, 1C (2 measurements)
    = Derbyia sp. Waagen 1884
Grant 1976
? Orthotetes sp.
1 individual
1 VV
    = Derbyia sp. Waagen 1884
Grant 1976
Diplanus (Miniplanus) minuta n. sp. Waterhouse and Piyasin 1970
34 individuals
21VV, 29DV, 5C (33 measurements)
Strophomenata - Strophomenida - Rugosochonetidae
Waagenites speciosus Waterhouse and Piyasin 1970
115 individuals
recombined as Waterhouseiella speciosa
36VV, 36DV, 79C (58 measurements)
Striochonetes scutella n. gen., n. sp. Waterhouse and Piyasin 1970
104 individuals
42VV, 42DV, 62C (66 measurements)
Linshuichonetes elfinis Campi and Shi 2002
Campi and Shi 2002
Waagenites speciosus Partim - Pl 17, figs 12-25 Only (7 measurements)
Strophomenata - Productida
? Marginifera aff. elongata (Huang 1932)
5 individuals
4VV, 1DV, 1C (2 measurements)
    = Marginifera drastica Grant 1976
Grant 1976
recombined as Marginifera (Marginifera) drastica
Marginifera typica Waagen 1884
13 individuals
8VV, 5DV, 5C (7 measurements)
    = Marginifera otaria Grant 1976
Grant 1976
recombined as Marginifera (Otariella) otaria
Strophomenata - Productida - Echinoconchidae
Septasteges acanthus n. gen., n. sp. Waterhouse and Piyasin 1970
7 individuals
recombined as Bilotina acantha
1VV, 6DV, 1C (2 measurements)
Strophomenata - Productida - Productellidae
Echinauris sp. Muir-Wood and Cooper 1960
2 individuals
2DV
Strophomenata - Productida - Linoproductidae
Linoproductus sp. (Chao 1927)
5 individuals
5 VV (1 measurement)
    = Linoproductus kaseti Grant 1976
Grant 1976
Strophomenata - Productida - Monticuliferidae
? Compressoproductus sp. Sarytcheva 1960
1 individual
1VV (1 measurement)
Asperlinus asperulus (Waagen 1884)
10 individuals
10VV, 5DV (4 measurements)
Strophomenata - Productida - Aulostegidae
? Xenosteges sp. Muir-Wood and Cooper 1960
2 individuals
1VV, 1C (1 measurement)
Chonosteges thailandica n. sp. Waterhouse and Piyasin 1970
5 individuals
nomen dubium belonging to Urushtenia
2VV, 2DV, 3C (4 measurements)
Ramavectus sp. Stehli 1954
11 individuals
11DV
    = Ramavectus pumwarni Grant 1976
Grant 1976
Strophomenata - Productida - Richthofeniidae
Globosobucina scopae n. gen., n. sp. Waterhouse and Piyasin 1970
1 individual
1C (1 measurement)
Richthofenia sp.
5 individuals
4VV, 1C (2 measurements)
Strophomenata - Productida - Lyttoniidae
Leptodus sp. Kayser 1883
9 individuals
5VV, 9DV (4 measurements)
Rhynchonellata - Rhynchonellida
Rhynchonellida indet. Kuhn 1949
6 individuals
1VV, 5C
Rhynchonellata - Rhynchonellida - Allorhynchidae
Wellerella naliensis (Reed 1944)
8 individuals
recombined as Allorhynchus naliensis
2VV, 2DV, 6C (5 measurements)
Rhynchonellata - Orthida - Rhipidomellidae
? Rhipidomella sp.
2 individuals
1 Dorsal Valve, 1 Conjoined (1 measurement)
    = Rhipidomella cordialis Grant 1976
Grant 1976
Rhynchonellata - Orthida - Schizophoriidae
Orthotichia indica (Waagen 1884)
14 individuals
10VV, 7DV, 4 C (7 measurements)
    = Orthotichia waterhousei Grant 1976
Grant 1976
recombined as Kotlaia waterhousei
Rhynchonellata - Spiriferinida - Spiriferellinidae
Spiriferellina aduncata n. sp. Waterhouse and Piyasin 1970
165 individuals
84VV, 71DV, 81C (77 measurements)
Rhynchonellata - Spiriferinida - Punctospiriferidae
? Punctospirifer sp. North 1920
1 individual
1 VV
Rhynchonellata - Spiriferida - Ambocoeliidae
Orbicoelia fraterculus n. gen., n. sp. Waterhouse and Piyasin 1970
12 individuals
11VV, 4DV, 1C (11 measurements)
Rhynchonellata - Spiriferida - Crassumbidae
Rorespirifer ruinosus n. gen., n. sp. Waterhouse and Piyasin 1970
4 individuals
3VV, 1DV, 1C (2 measurements)
Rhynchonellata - Spiriferida - Trigonotretidae
Neospirifer moosakhailensis (Davidson 1862)
4 individuals
3VV, 1DV
    = Neospirifer sp. Fredericks 1919
Grant 1976
Rhynchonellata - Spiriferida - Elythidae
Anomaloria glomerosa Grant 1976
Grant 1976
Phricodothyris attenuata Reed Partim - Pl 26, figs 9,14,15,18 Only (1 measurement)
Phricodothyris attenuata
14 individuals
13VV, 6DV, 1C (7 measurements)
    = Permophricodothyris notialasiatica Grant 1976
Grant 1976
Partim - Pl 26, figs 7,8,10-13,16,17 Only
Rhynchonellata - Athyridida - Neoretziidae
Hustedia ratburiensis n. sp. Waterhouse and Piyasin 1970
16 individuals
1VV, 5DV, 15C (14 measurements)
Rhynchonellata - Athyridida - Athyrididae
Cleiothyridina pectinifera (Sowerby 1841)
9 individuals
7VV, 2DV, 2C (7 measurements)
    = Cleiothyridina seriata Grant 1976
Grant 1976
Rhynchonellata - Athyridida - Diplospirellidae
Uncinella cf. indica Waagen 1883
1 individual
1C (1 measurement)
    = Uncinella siamestris Grant 1976
Grant 1976
Rhynchonellata - Terebratulida - Gillediidae
Hemiptychina sp. Waagen 1882
3 individuals
3C
    = Hemiptychina mintrita Grant 1976
Grant 1976
Rhynchonellata - Terebratulida - Cranaenidae
Pontielasma praeundatum n. gen., n. sp. Waterhouse and Piyasin 1970
13 individuals
3VV, 4DV, 9C (7 measurements)
Rhynchonellata - Terebratulida - Notothyrididae
Notothyris praelecta Reed 1944
39 individuals
20VV, 19DV, 19C (12 measurements)
    = Notothyris triplax n. sp. Grant 1976
Grant 1976
see common names

Geography
Country:Thailand State/province:Ratchaburi
Coordinates: 13.5° North, 99.7° East (view map)
Paleocoordinates:28.9° South, 98.5° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Permian Epoch:Guadalupian
Stage:Wordian 10 m.y. bin:Permian 3
Key time interval:Wordian
Age range of interval:268.80000 - 265.10000 m.y. ago
Stratigraphy
Formation:Rat Buri
Stratigraphic resolution:group of beds
Stratigraphy comments: The age of the Rat Buri fauna is problematic. "Rat Buri" limestones likely range in age from Kungurian through Capitanian to perhaps Wuchiapingian. Wordian fusulinids appear most common in other collections, but are not found in association with the brachiopod localities. The brachopod fauna is most similar to the Wordian Amb Formation of the Salt Range, in Pakistan, tentatively indicating a Wordian age.
Lithology and environment
Primary lithology: lithified argillaceous "limestone"
Secondary lithology:black lithified argillaceous,carbonaceous "limestone"
Includes fossils?Y
Includes fossils?Y
Lithology description: Two kinds of rock: muddy pale limestone and a black limestone that left a large residue of carbonaceous clay
Environment:offshore shelf
Geology comments: Fore-reef that is relatively open to the sea
Taphonomy
Modes of preservation:body,replaced with silica
Size of fossils:macrofossils
Collection methods and comments
Collection excludes:some macrofossils
Collection methods:bulk,chemical,field collection
Reason for describing collection:taxonomic analysis
Collectors:Waterhouse, Komalarjun, Nakinbodi, Sripatanavat Collection dates:November, 1964
Metadata
Database number:40954
Authorizer:D. Bottjer, M. Clapham Enterer:M. Clapham
Modifier:M. Clapham Research group:marine invertebrate
Created:2004-07-04 10:58:41 Last modified:2020-04-14 22:41:06
Access level:the public Released:2004-07-04 10:58:41
Creative Commons license:CC BY
Reference information

Primary reference:

11040. J. B. Waterhouse and S. Piyasin. 1970. Mid-Permian brachiopods from Khao Phrik, Thailand. Palaeontographica Abteilung A 135(3-6):83-197 [D. Bottjer/M. Clapham/M. Clapham]

Secondary references:

16868 N. W. Archbold. 1983. Permian marine invertebrate provinces of the Gondwana Realm. Alcheringa 7:59-73 [D. Bottjer/M. Clapham]
27583 M. J. Campi and G. R. Shi. 2002. Linshuichonetes gen. nov., a new rugosochonetid (Brachiopoda) genus from the Liangshan Formation (Early Permian) in Sichuan, China, and its ecology. Acta Palaeontologica Sinica 41(1):105-118 [M. Clapham/M. Clapham/M. Clapham]
11031 R. E. Grant. 1976. Permian brachiopods from southern Thailand. Paleontological Society Memoir 9:1-269 [D. Bottjer/M. Clapham/M. Clapham]
72089 C. Ketwetsuriya, A. G. Cook, and A. Nützel. 2020. Permian gastropods from the Ratburi Limestone, Khao Phrik, central Thailand. PalZ 94:53-77 [M. Clapham/M. Clapham/M. Clapham]