Gour Lazib HGL50: Ypresian - Lutetian, Algeria
collected 2003

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Scoleophidea indet.
Rage et al. 2021
Five trunk vertebrae (UM/HGL50-566, 567)
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
Rage et al. 2021
Twenty-three fragments (UM/HGL50-530) and more than 200 tiny fragments (UM/HGL50-531) of nonornamented maxillae; one incomplete atlas (UM/HGL50-532); one procoelous complex atlas+V2 represented by fused centra (UM/HGL50-533); one almost complete presacral vertebra (UM/HGL50-534); 27 incomplete or fragmentary presacral vertebrae (UM/HGL50-535); one almost complete humerus (UM/HGL50-536); 56 distal extremities of humeri (UM/ HGL50-537); 69 radioulna (UM/HGL50-538); one relatively complete ilium (UM/HGL50-539); 10 incomplete ilia (UM/ HGL50-540, 541); 25 tibiofibulae (UM/HGL50-542)
? Anura indet. (Fischer von Waldheim 1813)
Rage et al. 2021
Twenty-four presumed distal carpals 3-4-5 (UM/ HGL50-543-545)
Ranoidea indet. Rafinesque 1814
Rage et al. 2021
Six pre-8th vertebrae (UM/HGL50-524); eight 8th vertebrae (UM/HGL50-525); seven sacral vertebrae (UM/ HGL50-526); 21 humeri (UM/HGL50-527, 528); 14 ilia (UM/ HGL50-529).
Amphibia - Temnospondyli - Pipidae
Pipidae indet. Gray 1825
Rage et al. 2021
One ilium (UM/HGL50-487) and one humerus (UM/ HGL50-488)
Rocekophryne ornata n. gen., n. sp. Rage et al. 2021
Rage et al. 2021
Holotype. The posterior half of a frontoparietal (UM/HGL50- 489); Twenty-seven incomplete or fragmentary frontoparietals (UM/HGL50-490 to 493); 89 incomplete or fragmentary maxillae (UM/HGL50-494 to 503); three incomplete premaxillae (UM/HGL50-504, 505); seven fragmentary squamosals (UM/HGL50-506, 507); more than 260 fragments of ornamented cranial bones (UM/HGL50-508 to 512); five pre-8th presacral vertebrae (UM/HGL50-513, 514); five 8th presacral vertebrae (UM/HGL50-515, 516); one sacral vertebra (UM/HGL50-517); three humeri (UM/HGL50- 518 to 520); ten ilia (UM/HGL50-521 to 523).
Amphibia - Temnospondyli
? Caudata indet. (Scopoli 1777)
Rage et al. 2021
One incomplete dorsal vertebra (UM/HGL50-486)
Apoda indet. (Oppel 1810)
Rage et al. 2021
33 vertebrae (UM/HGL50-481 to 485).
Reptilia
Ophidia indet. (Latreille 1804)
Rage et al. 2021
Five fragmentary trunk vertebrae (UM/HGL50-580) and four caudal vertebrae (UM/HGL50-581)
Alethinophidia indet. (Nopcsa 1923)
Rage et al. 2021
Colubroidea indet. Oppel 1811
Rage et al. 2021
One incomplete mid-trunk vertebra (UM/HGL50- 579)
Afrotortrix draaensis n. gen., n. sp. Rage et al. 2021
Rage et al. 2021
Holotype. One mid-trunk vertebra (UM/HGL50-568).; Twenty trunk vertebrae (HGL50-569 to 573) and perhaps four poorly preserved trunk vertebrae (HGL50- 574).
Amphisbaenia indet. (Gray 1844)
Rage et al. 2021
One incomplete premaxilla (UM/HGL50-557) and five trunk vertebrae (UM/HGL50-558 and 559)
Scincomorpha informal indet. A (Camp 1923)
Rage et al. 2021
One fragment of dentary bearing four teeth (UM/ HGL50-550) and perhaps a fragment of maxilla bearing three teeth (UM/HGL50-551).
Scincomorpha informal indet. B (Camp 1923)
Rage et al. 2021
One fragment of bone bearing teeth (UM/HGL50- 552) and perhaps three other fragments bearing teeth (UM/ HGL50-553).
Scincomorpha informal indet. C (Camp 1923)
Rage et al. 2021
One fragment of bone bearing two teeth (UM/ HGL50-554), three fragments of maxillae bearing teeth (UM/ HGL50-555), and two larger fragments bearing one tooth each (UM/HGL50-556)
? Varanoidea indet. Gray 1827
Rage et al. 2021
One incomplete dorsal (UM/HGL50-560), one incomplete caudal (UM/HGL50-561) and perhaps another caudal vertebrae (UM/HGL50-562)
Reptilia - Agamidae
"Agamidae" indet. Gray 1827
Rage et al. 2021
One anterior portion of a left dentary (UM/HGL50- 546)
Reptilia - Gekkonidae
Gekkonidae indet. Gray 1825
Rage et al. 2021
Seven incomplete dentaries (UM/HGL50-547, 548) and four fragments bearing teeth (UM/HGL50-549)
Mammalia
Mammalia indet. Linnaeus 1758
Helioseus insolitus Sudre 1979
Mammalia - Chiroptera
Chiroptera indet. Blumenbach 1779
2 new genera and species
Mammalia - Chiroptera - Philisidae
Witwatia sigei Ravel et al. 2012
Ravel et al. 2014
Mammalia - Chiroptera
Drakonycteris glibzegdouensis n. gen., n. sp. Ravel et al. 2016
Ravel et al. 2016
Mammalia - Chiroptera - Emballonuridae
Pseudovespertiliavus parva n. gen., n. sp. Ravel et al. 2016
Ravel et al. 2016
Mammalia - Hyracoidea
Hyracoidea informal n. gen.
n. gen. n. sp.
Hyracoidea indet. Huxley 1869
Mammalia - Hyracoidea - Pliohyracidae
Microhyrax lavocati Sudre 1979
? Thyrohyrax sp. Meyer 1973
Mammalia - Proboscidea - Proboscidea
? Proboscidea indet. Illiger 1811
Mammalia - Condylarthra
Condylarthra indet. Cope 1881
n. gen. n. sp.
Mammalia - Creodonta
Creodonta indet. (Cope 1875)
Furodon crocheti n. gen., n. sp. Solé et al. 2014
Solé et al. 2014
HGL 50bis-56 - holotype; referred material: HGL 50-404, 405, 407 and 410
Glibzegdouia tabelbalaensis Crochet et al. 2001
GZC 35 - holotype; referred specimens: HGL 50-406, 408 and 411
Mammalia
Booidea informal indet. A
Rage et al. 2021
One mid- or posterior trunk vertebra (UM/HGL50- 575).
Booidea informal indet. B
Rage et al. 2021
One anterior trunk vertebra (UM/HGL50-576) and perhaps two caudal vertebrae (UM/HGL50-577).
Booidea indet.
Rage et al. 2021
Four poorly preserved vertebrae (UM/HGL50-578).
Mammalia - Lipotyphla
Lipotyphla indet. Haeckel 1866
synonym of Eulipotyphla
Mammalia - Rodentia
Rodentia indet. Bowdich 1821
Rodentia informal n. gen.
n. gen. n. sp.
Mammalia - Rodentia - Zegdoumyidae
Glibia pentalopha Vianey-Liaud et al. 1994
HGL50 specimens
Zegdoumys lavocati Vianey-Liaud et al. 1994
HGL50-01-16, 73 and 74
Lazibemys zegdouensis n. gen., n. sp. Marivaux et al. 2011
Marivaux et al. 2011
HGL50-70 - holotype; paratypes: HGL50-71 and 72
Mammalia - Primates
Primates indet. Linnaeus 1758
3 new genera and species
Mammalia - Primates - Azibiidae
Algeripithecus minutus Godinot and Mahboubi 1992
Azibius trerki Sudre 1975
Tabelia hammadae Godinot and Mahboubi 1994
synonym of Azibius trerki
Dralestes hammadaensis Tabuce et al. 2004
synonym of Azibius trerki
Dipnomorpha - Protopteridae
Protopteridae indet.
Actinopteri - Perciformes
Perciformes indet. Bleeker 1859
Actinopteri - Characiformes
Characiformes indet. Regan 1911
Actinopteri - Siluriformes
Siluriformes indet.
Actinopteri - Siluriformes - Clariidae
? Clariidae indet. Bonaparte 1846
Cladistia - Polypteriformes - Polypteridae
Polypteridae indet.
unclassified
Charophyta indet. Migula 1890
see common names

Geography
Country:Algeria State/province:Bechar region
Coordinates: 30.0° North, 4.0° West (view map)
Paleocoordinates:24.4° North, 6.5° West
Basis of coordinate:based on nearby landmark
Geographic resolution:small collection
Time
Period:Paleogene Epoch:Eocene
10 m.y. bin:Cenozoic 2
Key time interval:Ypresian - Lutetian
Age range of interval:56.00000 - 41.20000 m.y. ago
Stratigraphy
Formation:Glib Zegdou Member:Intermediate
Stratigraphic resolution:bed
Stratigraphy comments: 1 meter below HGL51; late Early or early Middle Eocene
Lithology and environment
Primary lithology:lenticular sandstone
Includes fossils?Y
Lithology description: Il renferme un horizon lenticulaire de grès jaune à rouge, à boules d’argile, très riche en charophytes. [yellow to red lenticular sandstone with clay balls, rich in charophytes] The fossiliferous level is a lenticular bed of carbonate cemented sandstone.
Environment:terrestrial indet.
Geology comments: Fluvatile origin
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,microfossils
Disassociated major elements:some
Disassociated minor elements:some
Collection methods and comments
Collection methods:chemical,field collection
Reason for describing collection:general faunal/floral analysis
Collection dates:2003 onwards
Collection method comments: Approximately 450 kg of sediment acid washed.
Taxonomic list comments:some mammals are new
Metadata
Also known as:Draa Hammada; Glib Zegdou outlier; Hammada du Dra
Database number:77014
Authorizer:P. Holroyd, P. Mannion, K. Beard, M. Uhen, J. Head Enterer:P. Holroyd, M. Jones, P. Mannion, G. Varnham, M. Uhen
Modifier:G. Varnham Research group:vertebrate
Created:2007-12-16 14:04:58 Last modified:2023-06-29 08:04:35
Access level:the public Released:2007-12-16 14:04:58
Creative Commons license:CC BY
Reference information

Primary reference:
26064. M. Adaci, R. Tabuce, F. Mebrouk, M. Bensalah, P.-H. Fabre, L. Hautier, J.-J. Jaeger, V. Lazzari, M. Mahboubi, L. Marivaux, O. Otero, S. Peigné, and H. Tong. 2007. Nouveaux sites à vertébrés paléogènes dans la région des Gour Lazib (Sahara nord-occidental, Algérie). Comptes Rendus Palevol 6:535-544 [P. Holroyd/P. Holroyd]

Secondary references:

45283 L. Marivaux, M. Adaci, M. Bensalah, H. G. Rodrigues, L. Hautier, M. Mahboubi, F. Mebrouk, R. Tabuce, and M. Vianey-Liaud. 2011. Zegdoumyidae (Rodentia, Mammalia), stem anomaluroid rodents from the Early to Middle Eocene of Algeria (Gour Lazib, Western Sahara): new dental evidence. Journal of Systematic Palaeontology 9(4):563-588 [P. Wagner/P. Wagner/P. Mannion]
85139 J. -C. Rage, M. Adaci, M. Bensalah, M. Mahboubi, L. Marivaux, F. Mebrouk, and R. Tabuce. 2021. Latest Early-early Middle Eocene deposits of Algeria (Glib Zegdou, HGL50), yield the richest and most diverse fauna of amphibians and squamate reptiles from the Palaeogene of Africa. Palaeovertebrata 44(1):e1 [J. Head/G. Varnham]
66475 A. Ravel, M. Adaci, M. Bensalah, M. Mahboubi, F. Mebrouk, E. M. Essid, W. Marzougui, H. K. Ammar, A.-L. Charruault, R. Lebrun, R. Tabuce, M. Vianey-Liaud, and L. Marivaux. 2014. New philisids (Mammalia, Chiroptera) from the Early-Middle Eocene of Algeria and Tunisia: new insight into the phylogeny, palaeobiogeography and palaeoecology of the Philisidae. [K. Beard/M. Jones]
64103 A. Ravel, M. Adaci, M. Bensalah, A.-L. Charruault, E. M. Essid, H. K. Ammar, W. Marzougui, M. Mahboubi, F. Merbrouk, G. Merzeraud, M. Vianey-Liaud, R. Tabuce, and L. Marvaux. 2016. Origine et radiation initiale des chauves-souris modernes : nouvelles découvertes dans l’Éocène d’Afrique du Nord. Geodiversitas 38(3):355-434 [M. Uhen/M. Uhen/M. Uhen]
59480 F. Solé, J. Lhuillier, M. Adaci, M. Bensalah, M. Mahboubi and R. Tabuce. 2014. The hyaenodontidans from the Gour Lazib area (?Early Eocene, Algeria): implications concerning the systematics and the origin of the Hyainailourinae and Teratodontinae. Journal of Systematic Palaeontology 12(3):303-322 [P. Mannion/P. Mannion]