Soyana (Iva Gora, PIN collection 94): Roadian, Russian Federation
collected by M. Edemskij 1934

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Bivalvia - Pectinida - Pseudomonotidae
Pseudomonotis speluncaria (von Schlotheim 1816)
recombined as Pseudomonotis (Pseudomonotis) speluncaria
Bivalvia - Pectinida - Streblochondriidae
Pecten sericeum (de Verneuil 1845)
recombined as Streblochondria sericea
Bivalvia - Ostreida - Pinnidae
Aviculopinna ivanisciana
Strophomenata - Productida - Linoproductidae
Productus cancrini de Verneuil 1845
recombined as Cancrinella cancrini
Rhynchonellata - Rhynchonellida - Rhynchoporidae
Rhynchopora geinitziana (de Verneuil 1845)
Lingulata - Lingulida - Lingulidae
Lingula orientalis
Lingula credneri Geinitz 1848
Insecta - Eoblattida - Bardapteridae
Olgaephilus emendatus Storozhenko 1990
Storozhenko 1990 2 specimens
PIN 94/120(121), 133(134) (1 measurement)
Insecta - Eoblattida - Megakhosaridae
Megakhosarina minuscula Aristov 2008
Aristov 2008 2 specimens
recombined as Blattokhosara minuscula
PIN 94/131, 338 (and/or 238)
Insecta - Reculida - Ivapteridae
Ivaptera sharovi Aristov 2009
Aristov 2009 8 specimens
PIN 94/50,132, 580, 660, 748, 834,1044,1048
Insecta - Palaeodictyoptera - Eukulojidae
Eukuloja cubitalis (Martynov 1932)
Sinitshenkova 1981 10 specimens
PIN 94/3, 7, 8, 10, 11, 13, 20, 710, 737, 1164
Eukuloja atrosumma Sinitshenkova 1981
Sinitshenkova 1981 2 specimens
PIN 94/935, 1158
Insecta - Permosialidae
Permosialis cauleoides n. sp. Martynova 1952
Martynova 1952 1 specimen
synonym of Permosialis paucinervis
PIN 94/798 (1 measurement)
Permosialis bifasciata Martynov 1932
Martynova 1952 3 specimens
synonym of Permosialis paucinervis
PIN 94/497, 576, 1212
Permosialis marmorata n. sp. Martynova 1952
Martynova 1952 2 specimens
synonym of Permosialis paucinervis
PIN 94/186(192), 721(676) (1 measurement)
Permosialis latiformis Martynova 1952
Martynova 1952 1 specimen
PIN 94/536
Insecta - Glosselytrodea - Jurinidae
Jurina reducta Martynov 1938
Martynov 1938 1 specimen
PIN 94/549
Jurina sojanensis Martynov 1938
Martynov 1938 4 specimens
PIN 94/555, 865; 864, 868 form radialis
Jurina marginata Martynov 1938
Martynov 1938 9 specimens
PIN 94/352, 369, 403, 409, 432, 549, 807, 937, 961
Insecta - Glosselytrodea - Glosselytridae
Glosselytron multivenosum Martynov 1938
Martynov 1938 1 specimen
PIN 94/276
Insecta - Paoliida - Paoliidae
Stenaropodites sojanensis Storozhenko 1992
Storozhenko 1992 1 specimen
PIN 94/1126
Sojanidelia striata Storozhenko 1992
Storozhenko 1992 2 specimens
PIN 94/559(814), 780(1132)
Sojanidelia makarkini Storozhenko 1992
Storozhenko 1992 2 specimens
PIN 94/370, 377
Sojanidelia nana n. sp. Storozhenko 1992
Storozhenko 1992 2 specimens
PIN 94/1122(1128), 1133 (1 measurement)
Sojanidelia fluctuosa Storozhenko 1992
Storozhenko 1992 1 specimen
PIN 94/675(934)
Permotermopsis pectinata Martynov 1937
Storozhenko 1992 1 specimen
PIN 94/115(6)
Mezenalicula connata n. gen., n. sp. Aristov and Storozhenko 2011
Aristov and Storozhenko 2011 1 specimen
PIN 94/883 (1 measurement)
Insecta - Grylloblattodea - Camptoneuritidae
Camptoneurites soyanensis Aristov et al. 2010
Aristov et al. 2010 1 specimen
PIN 94/29 (1 measurement)
Insecta - Cnemidolestodea - Sylvabestiidae
Sojanopermula lucida n. gen., n. sp. Storozhenko 1992
Storozhenko 1992 1 specimen
PIN 94/117 (1 measurement)
Permula edemskii n. sp. Aristov and Storozhenko 2011
Aristov and Storozhenko 2011 1 specimen
recombined as Sojanopermula edemskii
PIN 94/1027 (1 measurement)
Insecta - Hypoperlida - Hypoperlidae
Boreopsocus danksae Shcherbakov 1994
Shcherbakov 1994 3 specimens
PIN 94/197, 1299, 662
Insecta - Hemiptera - Prosbolidae
Sojanoneura triangularis Martynov 1935
Becker-Migdisova 1940 3 specimens
PIN 94/953, 1194, 640 (3 measurements)
Sojanoneura furcatula Martynov 1935
Becker-Migdisova 1940 3 specimens
PIN 94/624(586), 579, 810 (6 measurements)
Sojanoneura bimaculata Martynov 1935
Becker-Migdisova 1940 4 specimens
PIN 94/458, 639ab, 921, 862 (4 measurements)
Sojanoneura kazanensis Zalessky 1929
Becker-Migdisova 1940 2 specimens
PIN 94/466(452), 933 (2 measurements)
Prosbole indistincta n. sp. Becker-Migdisova 1940
Becker-Migdisova 1940 1 specimen
PIN 94/942 (1 measurement)
Prosbole permocicadoides Becker-Migdisova 1940
Becker-Migdisova 1940 2 specimens
PIN 94/943, 253 (2 measurements)
Prosbole sojanensis Martynov 1935
Becker-Migdisova 1940 1 specimen
PIN 94/1174(467) (1 measurement)
Prosbole longialata n. sp. Becker-Migdisova 1940
Becker-Migdisova 1940 1 specimen
PIN 94/1176 (1 measurement)
Prosbole ivagorae Becker-Migdisova 1940
Becker-Migdisova 1940 1 specimen
PIN 94/255(475) (1 measurement)
Prosbole brevialata n. sp. Becker-Migdisova 1940
Becker-Migdisova 1940 1 specimen
PIN 94/763(1190) (1 measurement)
Prosbole incerta n. sp. Becker-Migdisova 1940
Becker-Migdisova 1940 1 specimen
PIN 94/902 (1 measurement)
Permocicada simulatrix Martynov 1935
Becker-Migdisova 1940 2 specimens
PIN 94/465, 1189 (2 measurements)
Permocicada idelensis Zalessky 1929
Becker-Migdisova 1940 1 specimen
PIN 94/250(451) (1 measurement)
Permocicada pusilla n. sp. Becker-Migdisova 1940
Becker-Migdisova 1940 1 specimen
PIN 94/688 (1 measurement)
Permocicada integra n. sp. Becker-Migdisova 1940
Becker-Migdisova 1940 1 specimen
PIN 94/906ab (1 measurement)
Permocicada nigronervosa Martynov 1928
Becker-Migdisova 1940 1 specimen
PIN 94/249
Permocicada sublata n. sp. Becker-Migdisova 1940
Becker-Migdisova 1940 2 specimens
PIN 94/1055(1043), 179 (1 measurement)
Permocicada triramosa n. sp. Becker-Migdisova 1940
Becker-Migdisova 1940 1 specimen
PIN 94/1196 (1 measurement)
Permocicadopsis aberrans (Martynov 1935)
Becker-Migdisova 1940 2 specimens
PIN 94/462, 8885 (2 measurements)
Permocicadopsis angustata (Martynov 1935)
Becker-Migdisova 1940 6 specimens
PIN 94/645, 964; 468, 1047, 389, 886(var. typica) (4 measurements)
Insecta - Hemiptera - Prosbolopseidae
Prosbolopsis ovalis Martynov 1935
Becker-Migdisova 1960 5 specimens
PIN 94/420, 254, 251, 1184, 171
? Scytoneura sp. Martynov 1935
Becker-Migdisova 1960 1 specimen
PIN 94/641
Scytoneura elliptica Martynov 1935
Becker-Migdisova 1960 10 specimens
PIN 94/243, 450, 473, 480, 483, 577, 596, 1191, 1192, 1283 (7 measurements)
Anaprosbole ivensis Becker-Migdisova 1960
Becker-Migdisova 1960 1 specimen
PIN 94/464 (1 measurement)
Insecta - Hemiptera - Ingruidae
Ingruo lanceolata n. gen., n. sp. Becker-Migdisova 1960
Becker-Migdisova 1960 1 specimen
PIN 94/455 (1 measurement)
Insecta - Hemiptera - Scytinopteridae
Scytinoptera reducta (Martynov 1928)
Becker-Migdisova 1948 20 specimens
PIN 94/1179, 665, 469, 975, 472, 461, 408, 493, 922, 638, 245, 1251, 395, 881, 257, 585, 431, 784, 731, 912 (12 measurements)
Scytinoptera kokeni Handlirsch 1904
Becker-Migdisova 1948 2 specimens
PIN 94/481, 875 (1 measurement)
Insecta - Hemiptera - Coleoscytidae
Coleoscyta occallata n. sp. Becker-Migdisova 1960
Becker-Migdisova 1960 1 specimen
PIN 94/1178(482) (1 measurement)
Insecta - Hemiptera - Archescytinidae
Sojanoscytina tetraclada (Martynov 1933)
Becker-Migdisova 1960 1 specimen
recombined as Archescytina tetraclada
PIN 94/1185 (1 measurement)
Archescytina fusca Becker-Migdisova 1960
Becker-Migdisova 1960 2 specimens
PIN 94/474, 107 (2 measurements)
Sojanoscytina ramosa n. sp. Becker-Migdisova 1960
Becker-Migdisova 1960 1 specimen
PIN 94/1182 (1 measurement)
Sojanoscytina ivagorae n. sp. Becker-Migdisova 1960
Becker-Migdisova 1960 1 specimen
PIN 94/887 (1 measurement)
Insecta - Hemiptera - Boreoscytidae
Archescytinopsis flexuosa n. sp. Becker-Migdisova 1949
Becker-Migdisova 1949 2 specimens
synonym of Boreoscyta nefasta
PIN 94/1017, 775 (2 measurements)
Insecta - Permembiidae
Letopaloptera carpenteriana n. sp. Martynova 1961
Martynova 1961 1 specimen
PIN 94/561 (1 measurement)
Insecta - Mecoptera - Permopanorpidae
Permopanorpa angustata Martynov 1932
Martynova 1948 4 specimens
recombined as Martynopanorpa angustata
PIN 94/169, 515, 553, 620
Permopanorpa tenuis Martynov 1932
Martynova 1948 1 specimen
replaced by Permopanorpa martynovi
PIN 94/544
Insecta - Mecoptera - Permochoristidae
Petrochorista lepida Martynova 1948
Martynova 1948 1 specimen
PIN 94/754
Petrochorista sojanensis n. sp. Martynova 1948
Martynova 1948 1 specimen
PIN 94/1151 (1 measurement)
Petrochorista dubia Martynov 1932
Martynova 1948 4 specimens
recombined as Petromantis dubia
PIN 94/681, 730, 978, 1198
Permochorista maculipennis Martynov 1932
Martynova 1948 7 specimens
recombined as Petromantis maculipennis
PIN 94/142, 167, 419, 592_589), 768, 1152, 1210
Permochorista venosa Martynov 1932
Martynova 1948 3 specimens
synonym of Petromantis maculipennis
PIN 94/900, 1060, 1205
Petromantis rossica Handlirsch 1904
Martynova 1948 8 specimens
PIN 94/439, 501, 531, 746, 758, 861, 911; 94/1230 ("Soyana")
Agetopanorpa sp. Carpenter 1930
Novokshonov 1994 1 specimen
PIN 94/175
Agetochorista dubia Martynov 1932
Martynova 1948 6 specimens
recombined as Agetopanorpa dubia
PIN 94/179, 185, 540, 705 (2 wings), 827
Agetochorista letopalae Martynova 1948
Martynova 1948 1 specimen
recombined as Agetopanorpa letopalae
PIN 94/1153
Agetochorista tillyardiana Martynova 1948
Martynova 1948 2 specimens
recombined as Agetopanorpa tillyardiana
PIN 94/893, 909
Agetochorista ornata Martynov 1932
Martynova 1948 15 specimens
recombined as Agetopanorpa ornata
PIN 94/149, 164, 168, 170, 174, 176, 516, 538, 778, 980, 1139, 1157, 1211, 1213, 1312
Agetochorista umbrata Martynov 1932
Martynova 1948 2 specimens
synonym of Agetopanorpa ornata
PIN 94/1204, 1213
Agetochorista splendida Martynov 1932
Martynova 1948 10 specimens
synonym of Agetopanorpa ornata
PIN 94/165, 193, 595, 598, 671, 708, 709, 968, 1138; var. furcata 94/238
Insecta - Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis Martynov 1932
Martynova 1948 1 specimen
PIN 94/771
Insecta - Raphidioptera - Parasialidae
Parasialis dissedens n. sp. Ponomarenko 1977
Ponomarenko 1977 2 specimens
PIN 94/1217, 820 (1 measurement)
Insecta - Permithonidae
Permithonopsis triramosa n. sp. Martynova 1952
Martynova 1952 1 specimen
synonym of Permegalomus maculipennis
PIN 94/172 (1 measurement)
Permithonopsis proxima Martynov 1932
Martynova 1952 1 specimen
synonym of Permithonopsis ivensis
PIN 94/152
Permithonopsis obscura n. sp. Martynova 1952
Martynova 1952 1 specimen
PIN 94/173(726) (1 measurement)
Insecta - Kennedyidae
Kennedya ivensis Nel et al. 2012
Felker 2023 1 specimen
PIN 94/2 (1 measurement)
Malacostraca - Isopoda - Palaeophreatoicidae
Palaeophreatoicus sojanensis n. gen., n. sp. Birshtein 1962
see common names

Geography
Country:Russian Federation
Coordinates: 65.6° North, 43.6° East (view map)
Paleocoordinates:33.2° North, 33.0° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Permian Epoch:Guadalupian
Stage:Roadian 10 m.y. bin:Permian 3
Key time interval:Roadian
Age range of interval:272.30000 - 268.80000 m.y. ago
Stratigraphy
Stratigraphic resolution:group of beds
Lithology and environment
Primary lithology: lithified "siliciclastic"
Environment:lagoonal Tectonic setting:foreland basin
Geology comments: Probable brackish lagoon with some access to sea
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Collection methods and comments
Collection methods:survey of museum collection
Collection size:798 specimens
Reason for describing collection:taxonomic analysis
Collectors:M. Edemskij Collection dates:1934
Metadata
Database number:96860
Authorizer:M. Clapham Enterer:M. Clapham, J. Karr, L. Edwards
Modifier:M. Clapham Research group:marine invertebrate,paleoentomology
Created:2010-07-05 14:35:33 Last modified:2014-02-14 17:22:13
Access level:the public Released:2010-07-05 14:35:33
Creative Commons license:CC BY
Reference information

Primary reference:

33065. F. R. Schram. 1980. Miscellaneous late Paleozoic Malacostraca of the Soviet Union. Journal of Paleontology 54(3):542-547 [M. Clapham/M. Clapham]

Secondary references:

34929 D. S. Aristov. 2008. New Grylloblattids of the Family Megakhosaridae (Insecta: Grylloblattida) from the Permian of Russia. Paleontological Journal 42(3):269-272 [M. Clapham/J. Karr]
35088 D. S. Aristov. 2009. A New Family of the Order Grylloblattida (Insecta) from the Middle Permian of Russia. Paleontological Journal 43(2):178-182 [M. Clapham/M. Clapham/M. Clapham]
39962 D. S. Aristov and A. P. Rasnitsyn. 2012. Revision of the family Idelinellidae, with a review of the Permian Eoblattida (Insecta). Paleontological Journal 46:49-60 [M. Clapham/M. Clapham]
36405 D. S. Aristov, S. Y. Storozhenko, and Y. Y. Cui. 2010. Review of the Permian Camptoneuritidae (Insecta: Grylloblattida). Acta Geologica Sinica 84:756-761 [M. Clapham/M. Clapham]
37840 D. S. Aristov and S. Y. Storozhenko. 2011. Review of the Permian family Permulidae nomen novum pro Aliculidae Storozhenko, 1997 (Grylloblattida). ZooKeys 130:111-130 [M. Clapham/M. Clapham]
39941 E. E. Becker-Migdisova. 1940. Iskopaemye permskie cikady semejstva Prosbolidae s reki sojany. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 11(2):1-95 [M. Clapham/J. Karr/M. Clapham]
48456 E. E. Becker-Migdisova. 1948. Permskie tsikady semeystva Scytinopteridae s reki Soyany. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 15:1-42 [M. Clapham/M. Clapham]
37431 E. E. Becker-Migdisova. 1949. Novoe permskoe semeistvo Boreoscytidae i vopros o filogenii predkov Homoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 20:171-182 [M. Clapham/J. Karr/M. Clapham]
37942 E. E. Becker-Migdisova. 1960. New Permian Homoptera from European USSR. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 76:1-111 [M. Clapham/J. Karr/M. Clapham]
87497 A. S. Felker. 2023. New taxa of damselflies (Protozygoptera) from the Permian of the East European Platform. Paleontological Journal 57:857-914 [M. Clapham/M. Clapham]
47307 V. D. Ivanov. 1988. The structure of the Paleozoic caddis flies of the family Microptysmatidae (Insecta). Paleontological Journal 22(3):63-69 [M. Clapham/J. Karr]
53604 A. V. Martynov. 1938. O novom Permskom otryade pryamokrylykh nasekomykh Glosselytrodea. Izvestiya Akademii Nauk SSSR 1938:187-206 [M. Clapham/M. Clapham]
50889 O. M. Martynova. 1948. Materialy po evolyutsii Mecoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 14:1-76 [M. Clapham/M. Clapham/M. Clapham]
50930 O. M. Martynova. 1952. Permskie setchatokrylye SSSR. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 40:197-237 [M. Clapham/M. Clapham/M. Clapham]
39074 O. M. Martynova. 1961. Sovremennye i vymershie verblyudki (Insecta, Raphidioptera). Paleontologicheskii Zhurnal 1961(3):73-83 [M. Clapham/M. Clapham/M. Clapham]
42045 V. G. Novokshonov. 1994. Scorpion Flies of the Family Permochoristidae, the Closest Common Ancestors of Extant Scorpion Flies (Insecta, Panorpida-Mecoptera). Zoologicheskii Zhurnal 73(7-8):58-70 [M. Clapham/L. Edwards]
39786 A. G. Ponomarenko. 1977. Paleozoic Members of the Megaloptera (Insecta). Paleontological Journal 11(1):73-81 [M. Clapham/L. Edwards]
39083 D. E. Shcherbakov. 1994. A new genus of the Paleozoic order Hypoperlida. Russian Entomological Journal 3:33-36 [M. Clapham/M. Clapham]
41068 N. D. Sinitshenkova. 1981. Svoeobraznye "dvukrylye" nasekomye iz Permi Arkhangel'skoy oblasti i Urala (Hypoperlida, Eukulojidae). Byulleten Moskovskogo Obshchestva Ispytatelei Prirody, Otdel Geologicheskiy 56:116-126 [M. Clapham/M. Clapham/M. Clapham]
50618 S. Storozhenko. 1992. Permian fossil insects of north-east Europe: new and little-known Ideliidae (Insecta, Plecopteroidea, Grylloblattida). Entomologica Fennica 3:21-39 [M. Clapham/M. Clapham/M. Clapham]
37722 S. Y. Storozhenko. 1990. Permian fossil insects of North-East Europa: revision of the family Atactophlebiidae. Deutsche Entomologische Zeitschrift 37(4-5):407-412 [M. Clapham/J. Karr]
47277 S. Y. Storozhenko. 1992. Permian fossil insects of north-east Europe: new Liomopterids (Grylloblattida, Liomopteridae and Phenopteridae). Deutsche Entomologische Zeitschrift 39:209-220 [M. Clapham/J. Karr/M. Clapham]
50179 S. Y. Storozhenko and D. S. Aristov. 2014. Review of the Paleozoic and Mesozoic families Megakhosaridae and Blattogryllidae (Insecta: Grylloblattida). Far Eastern Entomologist 271:1-28 [M. Clapham/M. Clapham]