Sandelzhausen gravel pit: Langhian, Germany
collected by BSPG 1969-1975, 1994-2001
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Actinopteri
- Cypriniformes
- Cyprinidae
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Palaeocarassius informal sp. A
Obrhelová 1970
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Böhme 2010 | 1755 elements | ||||||||
1659 isolated pharyngeal teeth, 1 pharyngeal bone, 13 serrated fin spines, 85 first vertebrae | ||||||||||
Palaeoleuciscus informal sp. A
Obrhelová 1969
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Böhme 2010 | 33 elements | ||||||||
= P. sp. A sensu Böhme (2002, Beitr. Paläont. 27); 9 isolated pharyngeal teeth, 17 fragments of pharyngeal bone with attached teeth, 1 operculum fragment, 6 first vertebrae | ||||||||||
Actinopteri
- Anabantiformes
- Channidae
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Channa elliptica
von Salis 1967
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Böhme 2010 | 150 elements | ||||||||
68 vertebrae, 5 pharyngobranchial fragments, 1 premaxilla, 1 dentary fragment, >70 isolated teeth | ||||||||||
Actinopteri
- Gobiiformes
- Gobiidae
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Gobius sp.
Linnaeus 1758
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Böhme 2010 | 3 elements | ||||||||
1 premaxilla, 2 jaw elements | ||||||||||
Reptilia
- Phasianidae
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cf. Palaeortyx intermedia
Ballman 1966
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Gohlich 2003 | |||||||||
Reptilia
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Diplocynodon cf. gracilis
Vaillant 1872
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Scherer 1973 | 408 elements | ||||||||
1 fragment of dentary ("Dentale-Fragment 1" BSPG 1959 II 417) | ||||||||||
= Diplocynodon cf. buetikonensis
von Meyer 1855
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Scherer 1981 | |||||||||
additional specimens (BSPG 1959 II 34, 420 to 426, 436, 437, 680, 681, 2036-2042): 11 skull elements/fragments, 12 jaw fragments, 43 isolated teeth | ||||||||||
= Diplocynodon styriacus
Hofmann 1887
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Böhme 2010 | |||||||||
synonym of Diplocynodon ungeri | ||||||||||
"Diplocynodon cf. bueticonensis is a synonym of Diplocynodon styriacus according to Ginsburg & Bulot (1997)"; Böhme (2010) reports further 331 small isolated teeth (BSPG 1959 II 6526, 7539, 16363) which are said to have accumulated in the sedimentary environment due to tooth replacement in juveniles | ||||||||||
Diplocynodon sp.
Pomel 1847
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Scherer 1973 | 98 elements | ||||||||
2 fragmentary dentaries ("Dentale-Fragment 2" BSPG 1959 II 418, "Dentale-Fragment 3" BSPG 1959 II 419) and 96 isolated teeth (BSPG 1959 II 427 to 435) | ||||||||||
= Diplocynodon cf. buetikonensis
von Meyer 1855
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Scherer 1981 | |||||||||
= Diplocynodon styriacus
Hofmann 1887
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Böhme 2010 | |||||||||
synonym of Diplocynodon ungeri | ||||||||||
Reptilia
- Scincidae
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Scincidae informal indet. 1
Gray 1825
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Böhme 2010 | 2 fragments | ||||||||
2 dentary fragments | ||||||||||
Scincidae informal indet. 2
Gray 1825
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Böhme 2010 | 2 fragments | ||||||||
2 dentary fragments | ||||||||||
Tropidophorus bavaricus n. sp.
Böhme 2010
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Böhme 2010 | 4 specimens | ||||||||
1 dentary with 12 tooth positions (= holotype, BSPG 1959 II 16395), 3 dentary fragments | ||||||||||
Reptilia
- Cordylidae
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Cordylidae indet.
Gray 1837
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Böhme 2010 | 2 fragments | ||||||||
2 jaw fragments | ||||||||||
Bavaricordylus cf. molassicus
Böhme 2010
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Böhme 2010 | 6 fragments | ||||||||
2 maxillary fragments, 2 dentary fragments, 2 isolated dentary teeth | ||||||||||
Reptilia
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Scincomorpha indet.
(Camp 1923)
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Böhme 2010 | 6 specimens | ||||||||
2 intermaxillae, 4 jaw fragments | ||||||||||
Reptilia
- Anguidae
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Anguidae informal indet. sp. nov.
Gray 1825
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Böhme 2010 | 3 fragments | ||||||||
2 dentary fragments and 1 maxilla fragment | ||||||||||
Ophisaurus sp.
Daudin 1803
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Böhme 2010 | 40 specimens | ||||||||
13 skull bones/bone fragments, 10 dentaries/fragments + 2 further jaw fragments, 4 vertebrae, several osteoderms | ||||||||||
Ophisaurus informal sp. I sensu Roček, 1984
Daudin 1803
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Böhme 2010 | 10 fragments | ||||||||
4(?) dentary fragments, 6 maxilla fragments | ||||||||||
cf. Ophisaurus cf. spinari
Klembara 1979
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Böhme 2010 | 3 fragments | ||||||||
"sensu Roček, 1984"; 3 maxilla fragments | ||||||||||
Pseudopus laurillardi
(Lartet 1851)
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Böhme 2010 | 25 elements | ||||||||
5 skull bones, 4 dentaries/fragments, 7 vertebrae, 1 isolated tooth, several osteoderms | ||||||||||
Reptilia
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Palaeoblanus informal sp. nov.
Schleich 1988
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Böhme 2010 | 2 specimens | ||||||||
1 intermaxilla, 1 posterior dentary fragment | ||||||||||
Palaeoblanus tobieni
Schleich 1988
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Böhme 1999 | |||||||||
Reptilia
- Lacertidae
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Lacerta informal sp. 1
Linnaeus 1758
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Böhme 2010 | 1 element | ||||||||
"Lacerta sensu lato", left dentary | ||||||||||
Lacerta informal sp. 2
Linnaeus 1758
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Böhme 2010 | 10 specimens | ||||||||
"Lacerta sensu lato", 1 dentary + 7 fragments, 1 maxilla + 1 fragment | ||||||||||
Lacerta informal sp. 3
Linnaeus 1758
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Böhme 2010 | 21 specimens | ||||||||
"Lacerta sensu lato", 1 dentary + 14 fragments + 2 jaw fragments, 1 maxilla + 3 fragments | ||||||||||
cf. Miolacerta tenuis
Roček 1984
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Böhme 2010 | 2 fragments | ||||||||
2 dentary fragments | ||||||||||
Edlartetia sansaniensis
Augé and Rage 2000
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Böhme 2010 | 1 fragment | ||||||||
1 dentary fragment | ||||||||||
Reptilia
- Chamaeleonidae
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Chamaeleo aff. caroliquarti
Moody and Roček 1980
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Böhme 2010 | 5 elements | ||||||||
nomen dubium belonging to Chamaeleo | ||||||||||
5 skull bones/fragments | ||||||||||
Chamaeleo bavaricus n. sp.
Schleich 1983
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Schleich 1983 | 4 fragments | ||||||||
1 dentary fragment (=holotype, BSPG 1959 II 2066), 3 maxilla fragments (reported by Böhme, 2010) | ||||||||||
Reptilia
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Eoanilius sp.
Rage 1974
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Szyndlar 2009 | 10 elements | ||||||||
10 trunk vertebrae | ||||||||||
Reptilia
- Crotalidae
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Vipera sp.
Laurenti 1768
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Szyndlar 2009 | 70 elements | ||||||||
"Oriental viper" (contrastingly, Szyndlar & Rage, 2002, ref no. 43793, list the Vipera sp. from Sandelzhausen as a member of the "aspis complex"), 19 vertebrae and >50 isolated venom fangs | ||||||||||
Reptilia
- Elapidae
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Elapidae indet.
Boié 1827
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Szyndlar 2009 | 6 elements | ||||||||
6 trunk vertebrae | ||||||||||
Naja sp.
Laurenti 1768
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Szyndlar 2009 | 6 elements | ||||||||
6 trunk vertebrae | ||||||||||
Reptilia
- Colubridae
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"Natricinae" indet.
Bonaparte 1838
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Szyndlar 2009 | 50 elements | ||||||||
50 trunk vertebrae | ||||||||||
cf. Neonatrix sp.
Holman 1973
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Szyndlar 2009 | 4 elements | ||||||||
4 trunk vertebrae | ||||||||||
"Colubrinae" indet.
Oppel 1811
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Szyndlar 2009 | 12 elements | ||||||||
10 trunk vertebrae and 2 maxillary fragments, "tiny" | ||||||||||
"Coluber" sp.
Linnaeus 1758
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Szyndlar 2009 | 66 elements | ||||||||
65 trunk vertebrae + 1 maxillary fragment; the quotation marks are due to the fact that the author consciously uses the somewhat older concept of the genus (extant "colubrine" snakes by now assigned to the genera Hemorrhois, Hierophis, and Platyceps) | ||||||||||
? Telescopus sp.
Waner 1830
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Szyndlar 2009 | 2 elements | ||||||||
2 trunk vertebrae | ||||||||||
Natrix sp.
Laurenti 1768
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Szyndlar 2009 | 32 elements | ||||||||
32 trunk vertebrae | ||||||||||
cf. Natrix sp.
Laurenti 1768
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Szyndlar 2009 | 4 elements | ||||||||
4 trunk vertebrae, "large form" | ||||||||||
Reptilia
- Testudines
- Testudinidae
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Testudo rectogularis
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Schleich 1984 | |||||||||
Geochelone cf. perpiniana
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Schleich 1984 | |||||||||
= Titanochelon schleichi n. sp.
Pappa et al. 2023
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Pappa et al. 2023 | |||||||||
SNSB-BSPG 1959 II 2033, a partial skeleton comprising most of the carapace, the plastron, and some appendicu- lar elements (scapula, humerus, radius, ulna, carpal ele- ments, tibia, fibula, tarsal elements), mounted in anatomical position. | ||||||||||
Reptilia
- Testudines
- Geoemydidae
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Ocadia sophiae
(Ammon 1911)
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Hervet 2004 | |||||||||
recombined as Mauremys sophiae | ||||||||||
Clemmydopsis turnauensis
(Meyer 1847)
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Schleich 1984 | |||||||||
Reptilia
- Testudines
- Trionychidae
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Trionyx aff. bohemicus
Liebus 1930
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Schleich 1984 | |||||||||
= Trionychidae indet.
Gray 1825
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Georgalis and Joyce 2017 | |||||||||
Trionyx sp.
Geoffroy Saint-Hilaire 1809
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Schleich 1984 | |||||||||
= Trionychidae indet.
Gray 1825
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Georgalis and Joyce 2017 | |||||||||
Mammalia
- Proboscidea
- Mammutidae
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Zygolophodon turicensis
(Schintz 1824)
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Mammalia
- Proboscidea
- Gomphotheriidae
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Gomphotherium angustidens
Burmeister 1837
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subspecies G. a. subtapiroides | ||||||||||
Mammalia
- Talpidae
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Myxomygale hutchisoni
(Ziegler 1985)
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Hugueney and Maridet 2017 | |||||||||
Mammalia
- Rodentia
- Cricetidae
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Democricetodon aff. schaubi
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Fahlbusch 1964 | 15 elements | ||||||||
n. sp., 15 teeth | ||||||||||
Democricetodon minor
(Lartet 1851)
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Fahlbusch 1964 | 15 elements | ||||||||
n.ssp. gracilis, 15 teeth | ||||||||||
Democricetodon affinis
Schaub 1925
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Fahlbusch 1964 | 40 elements | ||||||||
n.ssp. mutilus, 40 teeth | ||||||||||
Cotimus bifidus n. sp.
Fahlbusch 1964
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Fahlbusch 1964 | 21 elements | ||||||||
recombined as Eumyarion bifidus | ||||||||||
21 teeth | ||||||||||
Amphibia
- Temnospondyli
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Latonia gigantea
Lartet 1851
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Böhme 2010 | 70 elements | ||||||||
42 maxillae and several other skull bones, 12 ilia and several further postcrania | ||||||||||
Amphibia
- Temnospondyli
- Pelobatidae
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Pelobates fahlbuschi n. sp.
Böhme 2010
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Böhme 2010 | 1080 elements | ||||||||
>1026 skull bones/bone fragments, 1 prearticular, 22 ilia, 29 vertebra + 1 urostyle | ||||||||||
Eopelobates sp.
Parker 1929
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Böhme 2010 | 10 elements | ||||||||
7 skull bones, 3 ilia | ||||||||||
Amphibia
- Temnospondyli
- Bufonidae
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Bufo sp.
Laurenti 1768
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Böhme 2010 | 1 fragment | ||||||||
single left ilium fragment | ||||||||||
Bufo cf. viridis
Laurenti 1768
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Böhme 2010 | 11 elements | ||||||||
1 braincase, 6 ilia, 3 sacral vertebrae + 1 urostyle | ||||||||||
Amphibia
- Temnospondyli
- Ranidae
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Ranidae informal indet. nov. gen. et sp.
Rafinesque 1814
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Böhme 2010 | 11 elements | ||||||||
1 maxilla fragment, 1 ilium, 9 terminal phalanges | ||||||||||
Pelophylax sp.
Fitzinger 1843
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Böhme 2010 | 40 elements | ||||||||
28 ilia and many further postcrania | ||||||||||
Amphibia
- Temnospondyli
- Urodela
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Triturus cf. vulgaris
(Linnaeus 1758)
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Böhme 2010 | 290 elements | ||||||||
285 vertebrae, several cranial and postcranial remains | ||||||||||
Triturus cf. marmoratus
(Latreille 1800)
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Böhme 2010 | 210 elements | ||||||||
3 occipital bones, 198 vertebrae, several cranial and postcranial remains | ||||||||||
Chelotriton sp.
Pomel 1853
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Böhme 2010 | 45 elements | ||||||||
11 skull bones/bone fragments, 6 dentaries, 26 vertebrae, several ribs | ||||||||||
Salamandra sansaniensis
Lartet 1851
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Böhme 2010 | 95 elements | ||||||||
3 occipital bones, 92 vertebrae | ||||||||||
Amphibia
- Temnospondyli
- Albanerpetontidae
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Albanerpeton inexpectatum
Estes and Hoffstetter 1976
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Böhme 2010 | 8 elements | ||||||||
4 skull bones, 2 dentaries, 2 vertebrae | ||||||||||
Gastropoda
- Pomatiidae
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Pomatias sp.
Stimpson 1860
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Salvador 2013 | |||||||||
Gastropoda
- Heterostropha
- Lymnaeidae
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Lymnaea dilatata
(Noulet 1854)
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Salvador and Rasser 2014 | |||||||||
Galba dupuyiana
(Noulet 1845)
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Salvador and Rasser 2014 | |||||||||
Gastropoda
- Heterostropha
- Planorbidae
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Planorbarius cornu
(Brongniart 1810)
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Salvador and Rasser 2014 | |||||||||
Ferrissia deperdita
(Desmarest 1814)
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Salvador and Rasser 2014 | |||||||||
Gyraulus albertanus
(Clessin 1877)
|
Salvador and Rasser 2014 | |||||||||
Gyraulus dealbatus
(Braun 1851)
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Salvador and Rasser 2014 | |||||||||
synonym of Gyraulus applanatus | ||||||||||
Gastropoda
- Heterostropha
- Pyramidulidae
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Pyramidula sp.
Fitzinger 1833
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Salvador 2015 | |||||||||
Trunactellina sp. in Moser et al. (2009) | ||||||||||
Gastropoda
- Heterostropha
- Gastrocoptidae
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Gastrocopta acuminata
(Klein 1846)
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Salvador 2015 | |||||||||
Gastrocopta nouletiana
(Dupuy 1850)
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Salvador 2015 | |||||||||
Gastropoda
- Heterostropha
- Chondrinidae
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Granaria sp.
Held 1837
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Salvador 2015 | |||||||||
Napaeus? sp. (in part) in Gall (1972); Granaria sp. 2 and ?Chondrinidae/?Enidae in Moser et al. (2009) | ||||||||||
Granaria cf. grossecostata
(Gottschick and Wenz 1919)
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Salvador 2015 | |||||||||
Napaeus? sp. (in part) in Gall (1972); Granaria sp. 1 in Moser et al. (2009) | ||||||||||
Gastropoda
- Heterostropha
- Strobilopsidae
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Strobilops sp.
Pilsbry 1893
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Salvador 2015 | |||||||||
Discus (Discus) pleuradra in Gall (1972); Strobilops (Strobilops) and S. (Discostrobilops) sp. in Moser et al. (2009) | ||||||||||
Gastropoda
- Heterostropha
- Vertiginidae
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Vertigo callosa
(Reuss 1849)
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Salvador 2015 | |||||||||
recombined as Vertigo (Vertigo) callosa | ||||||||||
Gastropoda
- Heterostropha
- Helicodiscidae
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Lucilla subteres
(Clessin 1877)
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Salvador and Rasser 2014 | |||||||||
Gastropoda
- Heterostropha
- Gastrodontidae
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Janulus supracostatus
(Sandberger 1873)
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Salvador and Rasser 2014 | |||||||||
Gastropoda
- Heterostropha
- Limacidae
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Limax sp.
Linnaeus 1758
|
Salvador and Rasser 2014 | |||||||||
Gastropoda
- Heterostropha
- Vitrinidae
|
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Vitrina sp.
Draparnaud 1801
|
Salvador and Rasser 2014 | |||||||||
Gastropoda
- Heterostropha
- Testacellidae
|
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Testacella zellii
Klein 1853
|
Salvador 2013 | |||||||||
Testacella larteti larteti in Gall (1972); Testacella sp. in Moser et al. (2009) | ||||||||||
Gastropoda
- Heterostropha
- Elonidae
|
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Tropidomphalus cf. incrassatus
(Klein 1853)
|
Salvador 2013 | |||||||||
recombined as Pseudochloritis incrassata | ||||||||||
Gastropoda
- Heterostropha
- Helicidae
|
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Helicidae indet.
Rafinesque 1815
|
Salvador 2013 | |||||||||
Cepaea cf. sylvestrina
(Schlotheim 1820)
|
Salvador 2013 | |||||||||
Cepaea cf. eversa
(Deshayes 1851)
|
Salvador 2013 | |||||||||
Gastropoda
- Heterostropha
- Hygromiidae
|
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Leucochroopsis kleinii
(Klein 1847)
|
Salvador 2013 | |||||||||
Gastropoda
- Heterostropha
- Spiraxidae
|
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Palaeoglandina sp.
Wenz 1914
|
Salvador 2013 | |||||||||
Palaeoglandina gracilis porrecta in Gall (1972) | ||||||||||
Gastropoda
- Heterostropha
- Clausiliidae
|
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Pseudidyla moersingensis
(Boettger 1877)
|
Salvador 2015 | |||||||||
Clausiliidae indet 1 and 2 in Moser et al. (2009) | ||||||||||
Gastropoda
- Heterostropha
- Filholiidae
|
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Triptychia sp.
Sandberger 1876
|
Salvador 2015 | |||||||||
Gastropoda
- Heterostropha
- Carychiidae
|
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Carychium (Carychium) galli n. sp.
Salvador 2015
|
Salvador 2015 | |||||||||
Carychium nouleti nouleti and gibbum in Gall (1972); C. sp. 1 and 2 in Moser et al. (2009) | ||||||||||
Carychium (Carychiella) eumicron
(Bourguignat 1857)
|
Salvador 2015 | |||||||||
Carychium eumicron peneckei in Gall (1972); Carychium sp. 3 in Moser et al. (2009) | ||||||||||
Gastropoda
- Heterostropha
- Valvatidae
|
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Valvata sp.
Müller 1774
|
Salvador 2013 | |||||||||
Gastropoda
- Discidae
|
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Discus pleuradrus
(Bourguignat 1881)
|
Salvador and Rasser 2014 | |||||||||
Gastropoda
- Neritidae
|
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Theodoxus sp.
Monfort 1810
|
Salvador 2013 | |||||||||
Bivalvia
- Cardiida
- Sphaeriidae
|
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Sphaerium sp.
Scopoli 1777
|
Salvador 2013 | |||||||||
see common names |
Geography
Country: | Germany | State/province: | Bavaria | County: | Kelheim |
Coordinates: | 48.6° North, 11.8° East (view map) | ||||
Paleocoordinates: | 48.9° North, 10.4° East | ||||
Basis of coordinate: | stated in text | ||||
Altitude: | 450 meters | ||||
Geographic resolution: | outcrop |
Time
Period: | Neogene | Epoch: | Miocene |
Stage: | Langhian | 10 m.y. bin: | Cenozoic 5 |
Key time interval: | Langhian | Mammal zone: | OSM C |
Age range of interval: | 15.98000 - 13.82000 m.y. ago | ||
Age estimate: | 16 Ma (graphic correlation) |
Stratigraphy
Geological group: | Molasse | Formation: | Upper Freshwater Molasse | ||
Stratigraphic resolution: | group of beds | ||||
Stratigraphy comments: The fossiliferous bed, which has a mean thickness of about 2.5 m, belongs to the Upper Freshwater Molasse (OSM) informal formation. Its age assigment has slightly changed several times over the past decades. Currently it is regarded as belonging to the OSM unit C (i.e. sedimentary cycle OSM 3), corresponding to the lower part of the Middle Miocene. Studies on several mammal groups have revealed that the locality biochronologically now is unequivocally referrable to the deeper part of MN 5 and not, as earlier thought, to MN6. The absolute age of the locality is about 16 Ma (see Fahlbusch, 2003 and references therein).
The fossiliferous bed itself is subdivided into 6 (sub)layers (A to F from base to top). However, in almost all cases no particular horizon is specified for a particular fossil or taxon in the paleontological literature. |
Lithology and environment
Primary lithology: | shelly/skeletal,black,brown,gray,green sandy,conglomeratic marl | ||
Secondary lithology: | unlithified silty mudstone | ||
Includes fossils? | Y | ||
Includes fossils? | Y | ||
Lithology description: The base of the fossiliferous bed rests on a paleorelief of about 30 cm. The (sub)layers A to D are composed of greenish-gray to brownish-black, marly conglomerates ("Geröllmergel") with decrease in quantity and size of pebbles from max. 10 cm in the lower part to few mm in the upper part of the fossiliferous bed. (Sub)layer A is partially well cemented by carbonate minerals. The pebbles mainly consist of quartz but also of various crystalline rock and hornfels, and some are flattened limestone pebbles. (Sub)layer C contains a thin lignite lens which, together with its lateral equivalents, served as guide bed dividing the "Untere Geröllmergel" below from the "Obere Geröllmergel" above. The (sub)layers A to D are roughly distinguishable from one another throughout the whole gravel pit, however, on a smaller scale rapid (often within few meters) facies changes occur. (Sub)layer E is a montmorillonite-rich bed of silty clay. The (sub)layers A to E all contain fossils: predominantly vertebrates and gastropod shell debris. (Sub)layer F is devoid of fossils. (Sub)layer B partially is characterized by structures which indicate synsedimentary movements such as slumps, but it cannot be ruled out that they may result from bioturbation. Almost all (sub)horizons of the fossiliferous bed are characterized by the presence of light grey to yellowish-white limestone concretions of various shape and size. These are the result of different diagenetic phenomena (e.g. subrecent precipitation from descendent meteoric water or early diagenetic processes related to fossil formation) (all informations on geology are from Fahlbusch 2003 and Böhme, 2010). | |||
Environment: | fluvial-lacustrine indet. | Tectonic setting: | foreland basin |
Geology comments: (Sub)bed A is interpreted as a gravel bed channel. (Sub)beds B to D (lower) may represent an alluvial plain in which, after seasonal floods, small, short-lived ponds existed under semiarid to semihumid conditions. (Sub)beds D (upper) and E are interpreted as permanent water body, a small lake or pond, under semihumid to humid conditions (Böhme, 2010). |
Taphonomy
Modes of preservation: | body |
Size of fossils: | macrofossils,mesofossils |
Spatial orientation: | random |
Preservation of anatomical detail: | variable |
Abundance in sediment: | abundant |
Disassociated minor elements: | all |
Size sorting: | well |
Fragmentation: | frequent |
Temporal resolution: | time-averaged |
Collection methods and comments
Collection excludes: | some macrofossils | ||
Collection methods: | bulk,surface (float),mechanical,acetic,sieve,survey of museum collection | ||
Rock censused: | 15000000 cm3 (volume) | ||
Reason for describing collection: | taxonomic analysis | ||
Museum repositories: | BSP | ||
Collectors: | BSPG | Collection dates: | 1969-1975, 1994-2001 |
Collection method comments: The fossil locality was discovered in 1959 by Richard Dehm (director of the BSPG at the time). After getting positive results from samples taken in 1962 systematic excavations including screenwashing took place from 1969 to 1975 (several weeks per year). In 1994 a second excavation period started which ended in 2001. During both periods a total of 1500 m3 has been quarried, accompanied by 150 t of screenwashed sediment (Fahlbusch, 2003). It should be noted, that amount of screenwashed samples varies between the (sub)layers and cannot be reconstructed (Böhme, 2010).
Almost all of the material is housed in the BSPG, bearing the catalogue signature BSPG 1959 II. | |||
Taxonomic list comments:Szyndlar (2009) lists 130 caudal vertebrae, 2 dentary fragments, and 1 unidentified dentigerous bone as Serpentes indet.
Note: Occurence of Eumyarion bifidus reported by Heißig (1989) has been removed from the list due to duplicate (alternative combination of Cotimus bifidus). (TL) |
Metadata
Also known as: | Sandelzhausen 1, Kiesgrube Bachmaier-Karber; Sandelshausen | ||
Database number: | 34615 | ||
Authorizer: | J. Hunter, T. Cleary, P. Mannion, J. Mueller, M. Clapham, E. Vlachos, P. Holroyd, D. Nicholson | Enterer: | M. Hall, P. Mannion, T. Cleary, E. Vlachos, M. Clapham, T. Liebrecht, D. Nicholson, P. Holroyd |
Modifier: | G. Varnham | Research group: | vertebrate |
Created: | 2003-09-12 07:54:45 | Last modified: | 2022-02-11 10:09:19 |
Access level: | the public | Released: | 2003-09-12 07:54:45 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
9144. | 5% 38460 | N. Schmidt-Kittler. 1972. Die obermiozäne Fossil-Lagerstätte Sandelzhausen. 6. Proboscidea (Mammalia). Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Histor. Geologie 12:83-95 [J. Hunter/M. Hall/T. Liebrecht] |
Secondary references:
67837 | M. Böhme. 1999. Doppelschleichen (Sauria, Amphisbaenia) aus dem Untermiozän von Stubersheim 3 (Süddeutschland). Mitteilungen der Bayerischen Staatssammlung für Paläontologie und historischen Geologie 39:85-90 [T. Cleary/T. Cleary] | |
43888 | M. Böhme. 2010. Ectothermic vertebrates (Actinopterygii, Allocaudata, Urodela, Anura, Crocodylia, Squamata) from the Miocene of Sandelzhausen (Germany, Bavaria) and their implications for environment reconstruction and palaeoclimate. Paläontologische Zeitschrift 84(1):3-41 [J. Mueller/T. Liebrecht] | |
12913 | V. Fahlbusch. 1964. Die Cricetiden (Mamm.) der Oberen Süßwasser-Molasse Bayerns (Cricetids of the Upper Freshwater Molasse of Bavaria). Bayerische Akademie der Wissenschaften; mathematisch-naturwissenschaftliche Klasse; Abhandlungen, neue Folge 118:1-136 [J. Alroy/S. Kuemmell/S. Kuemmell] | |
52039 | J. D. Gardner and M. Böhme. 2008. Review of the Albanerpetontidae (Lissamphibia), with comments on the paleoecological preferences of European Tertiary albanerpetontids. In J. T. Sankey & S. Baszio (ed.), Vertebrate Microfossil Assemblages: Their Role in Paleoecology and Paleobiogeography 178-218 [M. Carrano/M. Carrano] | |
77386 | J. D. Gardner, A. Villa, S. Colombero, M. Venczel, and M. Delfino. 2021. A Messinian (latest Miocene) occurrence for Albanerpeton Estes & Hoffstetter, 1976 (Lissamphibia: Albanerpetontidae) at Moncucco Torinese, Piedmont Basin, northwestern Italy, and a review of the European Cenozoic record for albanerpetontids. Geodiversitas 43(14):391-404 [M. Carrano/M. Carrano] | |
62147 | G. L. Georgalis and W. G. Joyce. 2017. A Review of the Fossil Record of Old World Turtles of the Clade Pan-Trionychidae. Bulletin of the Peabody Museum of Natural History 58(1):115-208 [P. Holroyd/P. Holroyd/P. Mannion] | |
40668 | U. B. Gohlich. 2003. The avifauna of the Grund Beds (Middle Miocene, Early Badenian, northern Austria). Annalen des Naturhistorischen Museums in Wien, Serie A 104:237-249 [M. Uhen/M. Uhen/T. Liebrecht] | |
58591 | S. Hervet. 2004. Systématique du groupe "Palaeochelys sensu lato – Mauremys" (Chelonii, Testudinoidea) du Tertiaire d'Europe occidentale: principaux résultats [Systematic of the "Palaeochelys sensu lato – Mauremys" group (Chelonii, Testudinoidea) from the Tertiary of Western Europe: principal results. Annales de Paléontologie 90(1):13-78 [D. Nicholson/D. Nicholson/D. Nicholson] | |
61807 | M. Hugueney and O. Maridet. 2017. Evolution of Oligo-Miocene talpids (Mammalia, Talpidae) in Europe: focus on the genera Myxomygale and Percymygale n. gen. Historical Biology [P. Mannion/P. Mannion] | |
70379 | M. Ivanov. 2001. Changes in the composition of the European snake fauna during the Early Miocene and at the Early/Middle Miocene transition. Palaeontologische Zeitschrift 74(4):563-573 [T. Cleary/T. Cleary] | |
85112 | I. Pappa, E. Vlachos, and M. Moser. 2023. A new species of a giant tortoise from Sandelzhausen (MN5, Burdigalian/Langhian boundary, Early/Middle Miocene, South Germany). The Anatomical Record 1-14 [E. Vlachos/E. Vlachos] | |
85582 | R. B. Salvador. 2013. The fossil land and freshwater mollusks of Sandelzhausen (Early/Middle Miocene, Germany): Caenogastropoda, Neritimorpha, lower Heterobranchia and Bivalvia. Strombus 20:19-26 [M. Clapham/M. Clapham] | |
85570 | R. B. Salvador. 2013. The fossil pulmonate snails of Sandelzhausen (Early/Middle Miocene, Germany): Succineidae, Testacelloidea and Helicoidea. Zootaxa 3721:157-171 [M. Clapham/M. Clapham] | |
85569 | R. B. Salvador. 2015. The fossil pulmonate snails of Sandelzhausen (Early/Middle Miocene, Germany): Ellobiidae, Pupilloidea, and Clausilioidea. Paläontologische Zeitschrift 89:37-50 [M. Clapham/M. Clapham] | |
85571 | R. B. Salvador and M. W. Rasser. 2014. The fossil pulmonate snails of Sandelzhausen (Early/Middle Miocene, Germany) (Hygrophila, Punctoidea and limacoids). Archiv für Molluskenkunde 143:187-202 [M. Clapham/M. Clapham] | |
43901 | E. Scherer. 1973. Die obermiozäne Fossil-Lagerstätte Sandelzhausen. 9. Crocodilia. Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie 13:103-114 [J. Mueller/T. Liebrecht] | |
43902 | E. Scherer. 1981. Die mittelmiozäne Fossil-Lagerstätte Sandelzhausen. 12. Crocodylia (abschließender Bericht). Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie 21:81-87 [J. Mueller/T. Liebrecht] | |
43939 | H.-H. Schleich. 1983. Die mittelmiozäne Fossil-Lagerstätte Sandelzhausen. 13. Chamaeleo bavaricus sp. nov., ein neuer Nachweis aus dem Jungtertiär Süddeutschlands. Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie 23:77-81 [J. Mueller/T. Liebrecht/T. Liebrecht] | |
66150 | H.H. Schleich. 1984. Neogene Testudines of Germany. Their stratigraphical and ecological evaluation. Studia Geologica Salmanticensia (Studia Palaeocheloniologica 1), volumen especial 1:249-267 [E. Vlachos/E. Vlachos] | |
43680 | Z. Szyndlar. 2009. Snake fauna (Reptilia: Serpentes) from the Early/Middle Miocene of Sandelzhausen and Rothenstein 13 (Germany). Paläontologische Zeitschrift 83(1):55-66 [J. Mueller/T. Liebrecht] |