Shanwang, Linqu County, collection SK (Shanwang Fossil Protection Post coll.) (Miocene of China)

Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)

• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone

• Two minor volcanic faulted basins, the Linqu and Shanwang Basins, are present in the study area. The former lies near the county town of Linqu, and the latter in the Shanwang National Geopark, east of this town. The faulted basins are the result of intense Mesozoic-Cenozoic tectonic extension and rifting; the development of these basins was closely related in time with the nearby NE-SW and NW-SE trending faults in the Yishu Fault Zone.

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Shanwang Fossil Protection Post. This collection contains specimens labeled with six-digit numbers prefaced by SK.

Primary reference: J. F. Zhang. 1993. New Miocene species of Bibionidae (Insecta: Diptera) with discussion on taxonomic position of Clothonopsis miocenica. Acta Palaeontologica Sinica 32:141-150 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 138945: authorized by Matthew Clapham, entered by Matthew Clapham on 03.02.2013

Creative Commons license: CC BY (attribution)

Taxonomic list

Arachnida
 Araneae - Thomisidae
Parvulus latissimus n. gen. n. sp.
Parvulus latissimus n. gen. n. sp. Zhang et al. 1994 crab spider
SK000650
 Araneae - Lycosidae
Dryadia acanthopoda n. gen. n. sp. Zhang et al. 1994 wolf spider
SK000548
Lycosa malleata n. sp. Zhang et al. 1994 wolf spider
SK000274
Lycosa subterranea n. sp. Zhang et al. 1994 wolf spider
SK000471
 Araneae - Salticidae
Evagoratus longicruris n. gen. n. sp.
Evagoratus longicruris n. gen. n. sp. Zhang et al. 1994 spider
SK000510
 Araneae - Araneidae
Araneus carbonaceus n. sp. Zhang et al. 1994 orb-weaving spider
SK000530
Araneus inelegans n. sp. Zhang et al. 1994 orb-weaving spider
SK000450
Araneus leptopoda n. sp. Zhang et al. 1994 orb-weaving spider
SK000560
 Araneae - Dysderidae
Dysdera dilatata n. sp. Zhang et al. 1994 woodlouse hunter spider
SK000348
Insecta
 Hemiptera - Reduviidae
Reduvius immitus n. sp. Zhang et al. 1994 assassin bug
SK000418
Reduvius diatomus Zhang and Zhang 1990 assassin bug
SK000338
Reduvius nicus n. sp. Zhang et al. 1994 assassin bug
SK000570
Oncocephalus astutus n. sp. Zhang et al. 1994 assassin bug
SK000631
 Hemiptera - Trisegmentatidae
Trisegmentatus onymus n. gen. n. sp.
Trisegmentatus onymus n. gen. n. sp. Zhang et al. 1994 true bug
SK000394
 Hemiptera - Coreidae
Notobitiella perpetis n. sp. Zhang et al. 1994 leaf-footed bug
SK000552
Paradasynus venustus n. sp. Zhang et al. 1994 leaf-footed bug
SK000460
Oxyproctus relictus n. gen. n. sp. Zhang et al. 1994 leaf-footed bug
SK000544
Ichnomnematus hadromastigus n. gen. n. sp. Zhang et al. 1994 leaf-footed bug
SK000323
 Hemiptera - Aphididae
"Cinara pastica n. sp." = Cinara (Cinara) pastica
"Cinara pastica n. sp." = Cinara (Cinara) pastica Zhang et al. 1994 aphid
SK000312
 Orthoptera - Acrididae
"Oedipodidae indet." = Oedipodinae, Oedemastopoda grammaca n. gen. n. sp.
"Oedipodidae indet." = Oedipodinae Walker 1871 band-winged grasshopper
SK000500
Oedemastopoda grammaca n. gen. n. sp. Zhang et al. 1994 band-winged grasshopper
SK000639
 Blattodea - Kalotermitidae
Kalotermes nigellus n. sp. Zhang et al. 1994 termite
SK000412
Kalotermes fossus n. sp. Zhang et al. 1994 termite
SK000643
 Hymenoptera - Tenthredinidae
Leptocampus amoenus n. gen. n. sp., Phthonocampus eos n. gen. n. sp., Emprionopsis doliabdominalis n. gen. n. sp.
Leptocampus amoenus n. gen. n. sp. Zhang et al. 1994 wood wasp
SK000320
Phthonocampus eos n. gen. n. sp. Zhang et al. 1994 wood wasp
SK000604
Emprionopsis doliabdominalis n. gen. n. sp. Zhang et al. 1994 wood wasp
SK000358
 Hymenoptera - Ichneumonidae
Epicharopimpla achaica n. gen. n. sp. Zhang et al. 1994 ichneumon wasp
SK000516
Neoparacryptus impavidus n. sp. Zhang et al. 1994 ichneumon wasp
SK000307
? Therion brachypodicum n. sp. Zhang et al. 1994 ichneumon wasp
SK000517
 Hymenoptera - Apidae
Xylocopa veta n. sp. Zhang et al. 1994 large carpenter bee
SK000586
Bombus dilectus n. sp. Zhang et al. 1994 bumble bee
SK000406
 Hymenoptera - Formicidae
Eurymyrmex geologicus n. gen. n. sp.
Eurymyrmex geologicus n. gen. n. sp. Zhang et al. 1994 ant
SK000268
 Hymenoptera - Tiphiidae
Tiphia rara n. sp.
Tiphia rara n. sp. Zhang et al. 1994 wasp
SK000462
 Lepidoptera - Sphingidae
Mioclanis shanwangiana n. gen. n. sp.
Mioclanis shanwangiana n. gen. n. sp. Zhang et al. 1994 moth
SK000361
 Coleoptera - Attelabidae
Eneuops variabilis n. gen. n. sp.
Eneuops variabilis n. gen. n. sp. Zhang et al. 1994 leaf rolling weevil
SK000281
 Coleoptera - Cerambycidae
Trosites naevius n. sp., Mesosa transversa n. sp., Miopyrestes doxa n. gen. n. sp., Perimesosa trigonia n. gen. n. sp., Anoplodera gigantea n. sp.
Trosites naevius n. sp. Zhang et al. 1994 longhorned beetle
SK000426
Mesosa transversa n. sp. Zhang et al. 1994 longhorned beetle
SK000565
Miopyrestes doxa n. gen. n. sp. Zhang et al. 1994 longhorned beetle
SK000574
Perimesosa trigonia n. gen. n. sp. Zhang et al. 1994 longhorned beetle
SK000554
Anoplodera gigantea n. sp. Zhang et al. 1994 longhorned beetle
SK000391
 Coleoptera - Chrysomelidae
Echinocallispa flavida n. gen. n. sp., Derorthaea curtiantenna n. gen. n. sp., "Haltica tholimorpha n. sp." = Altica tholimorpha, Disosebaethe pona n. gen. n. sp.
Echinocallispa flavida n. gen. n. sp. Zhang et al. 1994 tortoise beetle
SK000423
Derorthaea curtiantenna n. gen. n. sp. Zhang et al. 1994 leaf beetle
SK000417
"Haltica tholimorpha n. sp." = Altica tholimorpha Zhang et al. 1994 flea beetle
SK000569
Disosebaethe pona n. gen. n. sp. Zhang et al. 1994 leaf beetle
SK000374
 Coleoptera - Coccinellidae
Hippodamia olbia n. sp. Zhang et al. 1994 ladybird beetle
SK000334
 Coleoptera - Helotidae
Helota senilis n. sp., "Helota chinensis n. sp." = Helota zhangi
Helota senilis n. sp. Zhang et al. 1994 beetle
SK000476
"Helota chinensis n. sp." = Helota zhangi Wegrzynowicz 2007 beetle
SK000275
 Coleoptera - Elateridae
"Corymbites sincerus n. sp." = Ctenicera sincera
"Corymbites sincerus n. sp." = Ctenicera sincera Zhang et al. 1994 click beetle
SK000434
 Coleoptera - Staphylinidae
Lathrobium phthartum n. sp. Zhang et al. 1994 rove beetle
SK000397
 Coleoptera - Hydrophilidae
Penitus gemellus n. gen. n. sp.
Penitus gemellus n. gen. n. sp. Zhang et al. 1994 water scavenger beetle
SK000329
 Diptera - Asilidae
Promachus rapax n. sp. Zhang et al. 1994 robber fly
SK000330
 Diptera - Bibionidae
Bibio penisularis n. sp. Zhang 1993 march fly
SK000313
Bibio udus n. sp. Zhang et al. 1994 march fly
SK000388
Plecia platoptera n. sp. Zhang 1993 love bug
SK000444