Karatau-Mikhailovka (PIN collection 2384) (Jurassic to of Kazakhstan)

Where: Chimkent, Kazakhstan (42.9° N, 70.0° E: paleocoordinates 41.6° N, 73.7° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Karabastau Formation, Callovian to Callovian (166.1 - 157.3 Ma)

Environment/lithology: lacustrine - large; lithified siltstone

• "dark shales" Grimaldi & Engel, 2005

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Reposited in the PIN

Primary reference: A. G. Sharov. 1968. Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118:1-216 [C. Beard/C. Beard/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 106102: authorized by Matthew Clapham, entered by Matthew Clapham on 02.03.2011, edited by Andrew Spencer-Lee and Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Insecta
 Odonata - Aktassiidae
Aktassia magna n. gen. n. sp.37
Aktassia magna n. gen. n. sp.37 Pritykina 1968 true dragonfly
PIN 2384/4
 Odonata -
"Protolindenia aktassica" = Kazakhophlebiella aktassica37
"Protolindenia aktassica" = Kazakhophlebiella aktassica37 Pritykina 1968 true dragonfly
PIN 2384/2
 Odonata - Stenophlebiidae
Stenophlebia karatavica37
Stenophlebia karatavica37 Pritykina 1968 dragonfly
PIN 2384/3
 Odonata - Selenothemistidae
Turanothemis nodalis n. gen. n. sp.37
Turanothemis nodalis n. gen. n. sp.37 Pritykina 1968 dragonfly
PIN 2384/9
 Odonata - Isophlebiidae
Isophlebiodes obscurus n. gen. n. sp.37
Isophlebiodes obscurus n. gen. n. sp.37 Pritykina 1968 dragonfly
PIN 2384/18
 Blattaria - Caloblattinidae
Asioblatta punctata49
Asioblatta punctata49 Vishnyakova 1968 cockroach
PIN 2384/35,37
 Psocodea - Electrentomidae
Paramesopsocus adibi3
Paramesopsocus adibi3 Azar et al. 2008 book louse
PIN 2384/286, 291
 Hymenoptera - Anaxyelidae
? Anaxyelidae indet.17, Anaxyela destructa n. sp.40
? Anaxyelidae indet.17 Martynov 1925 winged insect
PIN 2384/1313
Anaxyela destructa n. sp.40 Rasnitsyn 1969 winged insect
PIN 2384/1314
 Hymenoptera - Xyelidae
Lydoxyela excellens n. gen. n. sp.38
Lydoxyela excellens n. gen. n. sp.38 Rasnitsyn 1966 sawfly
PIN 2384/1316
 Hymenoptera -
Prolyda karatavica n. gen. n. sp.39
Prolyda karatavica n. gen. n. sp.39 Rasnitsyn 1968 sawfly
PIN 2384/1311
 Hymenoptera - Blasticotomidae
Xyelocerus admirandus n. gen. n. sp.39
Xyelocerus admirandus n. gen. n. sp.39 Rasnitsyn 1968 wood wasp
PIN 2384/1315
 Hymenoptera - Bethylonymidae
Bethylonymus pedalis n. sp.42, Bethylonymus nigricornis n. sp.42
Bethylonymus pedalis n. sp.42 Rasnitsyn 1975 wasp
PIN 2384/1333
Bethylonymus nigricornis n. sp.42 Rasnitsyn 1975 wasp
PIN 2384/1946
 Hymenoptera - Mesoserphidae
Scoliuroserphus pallidulus44
Scoliuroserphus pallidulus44 Rasnitsyn 1986 wasp
PIN 2384/1367
 Hymenoptera - Ephialtitidae
Symphytopterus nigricornis42, ? Asiephialtites ventricosus n. sp.42, Stephanogaster pyriformis n. sp.42, Sessiliventer symphytus n. sp.42
Symphytopterus nigricornis42 Rasnitsyn 1975 wasp
PIN 2384/1323
? Asiephialtites ventricosus n. sp.42 Rasnitsyn 1975 wasp
PIN 2384/1332
Stephanogaster pyriformis n. sp.42 Rasnitsyn 1975 wasp
PIN 2384/1320
Sessiliventer symphytus n. sp.42 Rasnitsyn 1975 wasp
PIN 2384/1328
 Hymenoptera - Brachycleistogastridae
Brachycleistogaster nigritibialis42
Brachycleistogaster nigritibialis42 Rasnitsyn 1975 wasp
PIN 2384/1344
 Hymenoptera - Anomopterellidae
"Anomopterella stenocera n. sp." = Choristopterella stenocera42
"Anomopterella stenocera n. sp." = Choristopterella stenocera42 Rasnitsyn 1975 wasp
PIN 2384/1078
 Hymenoptera - Praeaulacidae
Praeaulacites subniger n. sp.41, Praeaulacites medius41
Praeaulacites subniger n. sp.41 Rasnitsyn 1972 wasp
PIN 2384/1945
Praeaulacites medius41 Rasnitsyn 1972 wasp
PIN 2384/1317, 1318, 1329
 Hymenoptera - Megalyridae
Mesaulacinus stenocerus n. sp.42, Leptocleistogaster alboscuta n. sp.42
Mesaulacinus stenocerus n. sp.42 Rasnitsyn 1975 wasp
PIN 2384/1325
Leptocleistogaster alboscuta n. sp.42 Rasnitsyn 1975 wasp
PIN 2384/1319
 Mecoptera - Bittacidae
Sibirobittacus kazakhstanensis n. sp.32
Sibirobittacus kazakhstanensis n. sp.32 Novokshonov 1993 hangingfly
PIN 2384/942
 Neuroptera - Chrysopidae
Baisochrysa pumila14
Baisochrysa pumila14 Khramov et al. 2015 green lacewing
PIN 2384/838
 Neuroptera - Ithonidae
Osmyloides distinctus n. gen. n. sp.34, "Kasachstania fasciata n. gen. n. sp." = Panfilovdvia fasciata34
Osmyloides distinctus n. gen. n. sp.34 Panfilov 1980 moth lacewing
PIN 2384/863
"Kasachstania fasciata n. gen. n. sp." = Panfilovdvia fasciata34 Panfilov 1980 moth lacewing
PIN 2384/814
 Neuroptera -
Chrysoleonites plexus n. sp.34
Chrysoleonites plexus n. sp.34 Panfilov 1980 lacewing
PIN 2384/842
 Neuroptera - Kalligrammatidae
Lithogramma oculatum n. gen. n. sp.33, Kalligramma flexuosum n. sp.33
Lithogramma oculatum n. gen. n. sp.33 Panfilov 1968 lacewing
PIN 2384/856
Kalligramma flexuosum n. sp.33 Panfilov 1968 lacewing
PIN 2384/857
 Neuroptera - Osmylidae
Jurosmylus parvulus13, Scapoptera recta n. gen. n. sp.34
Jurosmylus parvulus13 Khramov 2014 lance lacewing
PIN 2384/849
Scapoptera recta n. gen. n. sp.34 Panfilov 1980 lance lacewing
PIN 2384/862
 Neuroptera - Coniopterygidae
Juraconiopteryx zherichini n. gen. n. sp.27
Juraconiopteryx zherichini n. gen. n. sp.27 Meinander 1975 dustywing
PIN 2384/1300
 Coleoptera - Carabidae
Antemnacrassa albosulcata n. sp.30, Ovrabites ovalis2
Antemnacrassa albosulcata n. sp.30 Nikolajev 2004 ground beetle
PIN 2384/575
Ovrabites ovalis2 Ponomarenko 1977 ground beetle
PIN 2384/554
 Coleoptera - Triaplidae
Catinius pelta n. gen. n. sp.35
Catinius pelta n. gen. n. sp.35 Ponomarenko 1968 beetle
PIN 2384/634
 Coleoptera - Trachypachidae
Psacodromeus ovalis2, Karadromeus latus n. sp.2
Psacodromeus ovalis2 Ponomarenko 1977 false ground beetle
PIN 2384/757
Karadromeus latus n. sp.2 Ponomarenko 1977 false ground beetle
PIN 2384/567
 Coleoptera - Coptoclavidae
Charonoscaphidia elongata n. gen. n. sp.2
Charonoscaphidia elongata n. gen. n. sp.2 Ponomarenko 1977 beetle
PIN 2384/717, 707, 718
 Coleoptera -
Notocupes lapidarius n. sp.35
Notocupes lapidarius n. sp.35 Ponomarenko 1968 beetle
PIN 2384/404
 Coleoptera - Phoroschizidae
Tersus leptocerus35
Tersus leptocerus35 Ponomarenko 1968 beetle
PIN 2384/715
 Coleoptera - Cupedidae
Cupidium abavum n. gen. n. sp.35
Cupidium abavum n. gen. n. sp.35 Ponomarenko 1968 reticulated beetle
PIN 2384/398
 Coleoptera - Ommatidae
"Tetraphalerus maximus" = Allophalerus maximus35, Omma aberratum n. sp.35
"Tetraphalerus maximus" = Allophalerus maximus35 Ponomarenko 1968 beetle
PIN 2384/401
Omma aberratum n. sp.35 Ponomarenko 1968 beetle
PIN 2384/396
 Coleoptera -
"Notocupes nigrimonticola n. sp." = Brachilatus nigrimonticola35
"Notocupes nigrimonticola n. sp." = Brachilatus nigrimonticola35 Ponomarenko 1968 beetle
PIN 2384/397
 Coleoptera - Staphylinidae
Mesotachinus major n. sp.48, Archodromus comptus n. gen. n. sp.48, Abolescus glabratus48, Mesoxytelus mandibularis48
Mesotachinus major n. sp.48 Tikhomirova 1968 rove beetle
PIN 2384/417,419
Archodromus comptus n. gen. n. sp.48 Tikhomirova 1968 rove beetle
PIN 2384/418
Abolescus glabratus48 Tikhomirova 1968 rove beetle
PIN 2384/423,427
Mesoxytelus mandibularis48 Tikhomirova 1968 rove beetle
PIN 2384/444
 Coleoptera - Hybosoridae
Protohybosorus mesoasiaticus n. sp.31
Protohybosorus mesoasiaticus n. sp.31 Nikolajev 2010 scavenger scarab beetle
PIN 2384/760
 Coleoptera - Nemonychidae
Oxycorynoides (Oxycorynoides) similis10, Oxycorynoides (Pseudoxycorynoides) sukatshevae10, Eobelus longipes2, Belonotaris (Arnoldibelus) rasnitsyni10, Belonotaris rohdendorfi22, Belonotaris (Belonotaris) punctatissimus10, Ampliceps dubius10, "Scelocamptus curvipes n. sp." = Ampliceps tenuirostris2, Ampliceps dentitibia10, Ampliceps furcitibia10, "Brenthorrhinus brevirostris n. sp." = Abrenthorrhinus brevirostris11, Distenorrhinus (Distenorrhinus) pallidirostris12, "Distenorrhinus (Astenorrhinus) major n. sp." = Astenorrhinus major12, Procurculio fortipes12, "Pseudobrenthorrhinus crassicornis n. gen. n. sp." = Eccoptarthrus crassipes11
Oxycorynoides (Oxycorynoides) similis10 Arnoldi 1977 pine flower snout beetle
PIN 2384/513, 520, 532, 533, 538
Oxycorynoides (Pseudoxycorynoides) sukatshevae10 Legalov 2014 pine flower snout beetle
PIN 2384/514, 521, 534
Eobelus longipes2 Arnoldi 1977 pine flower snout beetle
PIN 2384/516
Belonotaris (Arnoldibelus) rasnitsyni10 Legalov 2014 pine flower snout beetle
PIN 2384/531
Belonotaris rohdendorfi22 Legalov 2014 pine flower snout beetle
PIN 2384/535
Belonotaris (Belonotaris) punctatissimus10 Arnoldi 1977 pine flower snout beetle
PIN 2384/516
Ampliceps dubius10 Arnoldi 1977 pine flower snout beetle
PIN 2384/511, 613
"Scelocamptus curvipes n. sp." = Ampliceps tenuirostris2 Arnoldi 1977 pine flower snout beetle
PIN 2384/512
Ampliceps dentitibia10 Arnoldi 1977 pine flower snout beetle
PIN 2384/530
Ampliceps furcitibia10 Arnoldi 1977 pine flower snout beetle
PIN 2384/507, 653
"Brenthorrhinus brevirostris n. sp." = Abrenthorrhinus brevirostris11 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2384/510
Distenorrhinus (Distenorrhinus) pallidirostris12 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2384/536
"Distenorrhinus (Astenorrhinus) major n. sp." = Astenorrhinus major12 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2384/526
Procurculio fortipes12 Arnoldi 1977 pine flower snout beetle
PIN 2384/517
"Pseudobrenthorrhinus crassicornis n. gen. n. sp." = Eccoptarthrus crassipes11 Arnoldi 1977 pine flower snout beetle
PIN 2384/537
 Coleoptera - Brentidae
Karacar contractus n. gen. n. sp.9
Karacar contractus n. gen. n. sp.9 Gratshev and Legalov 2011 weevil
 Coleoptera - Obrieniidae
Pseudobrienia rasnitsyni n. gen. n. sp.20
Pseudobrienia rasnitsyni n. gen. n. sp.20 Legalov 2012 weevil
PIN 2384/720
 Coleoptera - Anthribidae
Protoscelis medvedevi21, "Pseudomegamerus grandis" = Protoscelis longicornis26, Protoscelis jurassica26, Protoscelis parvula21
Protoscelis medvedevi21 Legalov 2013 fungus weevil
PIN 2384/569
"Pseudomegamerus grandis" = Protoscelis longicornis26 Medvedev 1968 fungus weevil
PIN 2384/415
Protoscelis jurassica26 Medvedev 1968 fungus weevil
PIN 2384/770; 655,725,726,733,751,752,781,789 (in Legalov, 2013)
Protoscelis parvula21 Medvedev 1968 fungus weevil
PIN 2384/509, 639
 Coleoptera - Parandrexidae
Parandrexis parvula15
Parandrexis parvula15 Martynov 1926 beetle
PIN 2384/701
 Coleoptera - Cerophytidae
"Idiomerus musculus" = Necromera muscula6, "Idiomerus intermedius n. sp." = Necromera intermedia6
"Idiomerus musculus" = Necromera muscula6 Dolin 1980 beetle
PIN 2384/475
"Idiomerus intermedius n. sp." = Necromera intermedia6 Dolin 1980 beetle
PIN 2384/497
 Coleoptera - Elateridae
Desmatus beckeri5, Lithomerus cockerelli n. gen. n. sp.6, Lithocoelus detrusus n. gen. n. sp.5, Acheonus abbreviatus n. gen. n. sp.6, Platyelater reflexicollis6, Hypnomorphus curtus6, Hypnomorphus rohdendorfi n. gen. n. sp.5, Hypnomorphus inventus n. sp.5, Codemus synaptoides6, Codemus sharovi n. sp.6, Codemus quadricolis6, Codemus carinatus n. sp.6, Adiagnostus minutulus n. sp.6, Hypnomorphoides latus6
Desmatus beckeri5 Dolin 1975 click beetle
PIN 2384/489
Lithomerus cockerelli n. gen. n. sp.6 Dolin 1980 click beetle
PIN 2384/454, 460, 492
Lithocoelus detrusus n. gen. n. sp.5 Dolin 1975 click beetle
PIN 2384/484
Acheonus abbreviatus n. gen. n. sp.6 Dolin 1980 click beetle
PIN 2384/462 (paratype also given as specimen 462 in text
Platyelater reflexicollis6 Dolin 1980 click beetle
PIN 2384/485
Hypnomorphus curtus6 Dolin 1980 click beetle
PIN 2384/472
Hypnomorphus rohdendorfi n. gen. n. sp.5 Dolin 1975 click beetle
PIN 2384/457
Hypnomorphus inventus n. sp.5 Dolin 1975 click beetle
PIN 2384/491, 450, 479
Codemus synaptoides6 Dolin 1980 click beetle
PIN 2384/481(483)
Codemus sharovi n. sp.6 Dolin 1980 click beetle
PIN 2384/474, 465
Codemus quadricolis6 Dolin 1980 click beetle
PIN 2384/500, 502
Codemus carinatus n. sp.6 Dolin 1980 click beetle
PIN 2384/464, 483
Adiagnostus minutulus n. sp.6 Dolin 1980 click beetle
PIN 2384/455
Hypnomorphoides latus6 Dolin 1980 click beetle
PIN 2384/449
 Permopsocida - Archipsyllidae
Archipsylla lata n. sp.50
Archipsylla lata n. sp.50 Vishnyakova 1976 winged insect
PIN 2384/335
 Hemiptera - Corixidae
Archaecorixa lata n. gen. n. sp.36
Archaecorixa lata n. gen. n. sp.36 Popov 1968 water boatman
PIN 2384/377
 Hemiptera - Naucoridae
Aidium pleurale n. gen. n. sp.36
Aidium pleurale n. gen. n. sp.36 Popov 1968 creeping water bug
PIN 2384/368
 Hemiptera - Protopsyllidiidae
Karatavopsyllidium martynovi n. sp.4
Karatavopsyllidium martynovi n. sp.4 Becker-Migdisova 1985 true bug
PIN 2384/293
 Hemiptera - Liadopsyllidae
Liadopsylla grandis4, Liadopsylla tenuicornis4
Liadopsylla grandis4 Becker-Migdisova 1985 jumping plant louse
PIN 2384/324, 328
Liadopsylla tenuicornis4 Martynov 1926 jumping plant louse
PIN 2384/318
 Orthoptera - Prophalangopsidae
Aboilus femineus7, Aboilus aulietus
Aboilus femineus7 Gorochov 1996 hump-winged cricket
PIN 2384/217
Aboilus aulietus Sharov 1968 hump-winged cricket
PIN 2384/248
 Orthoptera - Protogryllidae
"Protogryllus karatavicus n. sp." = Falsispeculum karatavicus
"Protogryllus karatavicus n. sp." = Falsispeculum karatavicus Sharov 1968 grasshopper
PIN 2384/255
 Dermaptera - Protodiplatyidae
Microdiplatys oculatus n. sp.51
Microdiplatys oculatus n. sp.51 Vishnyakova 1980 earwig
PIN 2384/196
 Dicondylia -
Blattodea indet.1
Blattodea indet.1 Latreille 1810 cockroach
PIN 2384/8 described as a paratype of Blattogryllus karatavicus
 Blattodea - Liberiblattinidae
Hydrokhoohydra aquabella52
Hydrokhoohydra aquabella52 Vršanský 2019 cockroach
PIN 2384/186, 194, 1274
 Blattodea - Mesoblattinidae
Latiblatta lativalvata49
Latiblatta lativalvata49 Vishnyakova 1968 cockroach
PIN 2384/192
 Plecoptera - Perlariopseidae
Karanemoura abrupta n. gen. n. sp.47
Karanemoura abrupta n. gen. n. sp.47 Sinitshenkova 1987 stonefly
PIN 2384/2367
 Embioptera - Brachyphyllophagidae
Brachyphyllophagus phantasus n. sp.45
Brachyphyllophagus phantasus n. sp.45 Rasnitsyn 2000 webspinners
PIN 2384/260
 Dicondylia - Susumaniidae
Phasmomimoides minutus n. sp.8
Phasmomimoides minutus n. sp.8 Gorochov 2000 winged insect
PIN 2384/261
 Eoblattida - Blattogryllidae
Blattogryllus karatavicus43
Blattogryllus karatavicus43 Rasnitsyn 1976 insect
PIN 2384/8, 205
 Diptera - Nemestrinidae
"Eohirmoneura carpenteri n. gen. n. sp." = Hirmoneura (Eohirmoneura) carpenteri46
"Eohirmoneura carpenteri n. gen. n. sp." = Hirmoneura (Eohirmoneura) carpenteri46 Rohdendorf 1968 tangle-veined fly
PIN 2384/1236
 Diptera - Rhagionidae
Protorhagio ruptus29, Protorhagio karataviensis29
Protorhagio ruptus29 Mostovski 2008 snipe fly
PIN 2384/1224 (also listed under Protorhagio capitatus)
Protorhagio karataviensis29 Mostovski 2008 snipe fly
PIN 2384/1146
 Diptera -
Apystomimus zaitzevi28
Apystomimus zaitzevi28 Mostovski 1999 fly
PIN 2384/1258
 Diptera - Procramptonomyiidae
Procramptonomyia kovalevi24
Procramptonomyia kovalevi24 Lukashevich 2012 fly
PIN 2384/1149,1173, 1281
 Diptera - Protorhyphidae
Archirhyphus asiatcus24
Archirhyphus asiatcus24 Rohdendorf 1964 fly
PIN 2384/1021
 Diptera - Ptychopteridae
Eoptychopterina karatavica23
Eoptychopterina karatavica23 Lukashevich 1993 phantom crane fly
PIN 2384/1238; 2323 in Lukashevich, 2004
 Diptera - Tipulidae
Tipunia completa16
Tipunia completa16 Kopeć et al. 2023 crane fly
PIN 2384/1160
 Diptera - Trichoceridae
Tanyochoreta (Trichokara) minuta18
Tanyochoreta (Trichokara) minuta18 Krzemińska et al. 2009 winter crane fly
PIN 2384/1201
 Diptera - Elliidae
Polyanka minuta19
Polyanka minuta19 Krzemińska et al. 1993 fly
PIN 2384/988, 87
 Diptera - Psychodidae
Praemacrochile ansorgei25
Praemacrochile ansorgei25 Lukashevich and Krzemiński 2009 moth fly
PIN 2384/1067, 1266
1Aristov et al. 2006; 2Arnoldi et al. 1977; 3Azar et al. 2008; 4Becker-Migdisova 1985; 5Dolin 1975; 6Dolin 1980; 7Gorochov 1996; 8Gorochov 2000; 9Gratshev and Legalov 2011; 10Gratshev and Legalov 2014; 11Gratshev and Zherikhin 1995; 12Gratshev and Zherikhin 1995; 13Khramov 2014; 14Khramov et al. 2016; 15Kirejtshuk 1994; 16Kopeć et al. 2023; 17Kopylov 2018; 18Krzemińska et al. 2009; 19Krzemińska et al. 1993; 20Legalov 2012; 21Legalov 2013; 22Legalov 2022; 23Lukashevich 1993; 24Lukashevich 2012; 25Lukashevich and Krzemiński 2009; 26Medvedev 1968; 27Meinander 1975; 28Mostovski 1999; 29Mostovski 2008; 30Nikolajev 2004; 31Nikolajev 2010; 32Novokshonov 1993; 33Panfilov 1968; 34Panfilov 1980; 35Ponomarenko 1968; 36Popov 1968; 37Pritykina 1968; 38Rasnitsyn 1966; 39Rasnitsyn 1968; 40Rasnitsyn 1969; 41Rasnitsyn 1972; 42Rasnitsyn 1975; 43Rasnitsyn 1976; 44Rasnitsyn 1986; 45Rasnitsyn and Krassilov 2000; 46Rohdendorf 1968; 47Sinitshenkova 1987; 48Tikhomirova 1968; 49Vishnyakova 1968; 50Vishnyakova 1976; 51Vishnyakova 1980; 52Vršanský et al. 2019