Bembridge Marls (Hooley coll) (Eocene of the United Kingdom)

Where: England, United Kingdom (50.7° N, 1.4° W: paleocoordinates 49.2° N, 5.2° W)

• coordinate based on nearby landmark

When: Bembridge Marls Member (Bouldnor Formation), Priabonian (38.0 - 33.9 Ma)

• "insect beds"

•Upper part of 13r chron just above 1n interval

Environment/lithology: lagoonal or restricted shallow subtidal; lithified, argillaceous lime mudstone

• The Bembridge Marls were deposited in fresh-brackish water lakes and lagoons

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by R. W. Hooley; reposited in the BMNH

• Specimens kept in British Natural History Museum

•Most specimens collected by Mr. R. W. Hooley.

•Some of A'Court Smiths collection was obtained by R. W. Hooley and is in his collection as well.

Primary reference: F. E. Zeuner. 1939. Fossil Orthoptera Ensifera 1-321 [M. Clapham/M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 123960: authorized by Matthew Clapham, entered by Jered Karr on 03.02.2012, edited by Jenna Sandlin and Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hemiptera -
Hastites muiri n. gen. n. sp.6
Hastites muiri n. gen. n. sp.6 Cockerell 1922 planthopper
H.353
 Hemiptera - Achilidae
Hooleya indecisa n. gen. n. sp.6
Hooleya indecisa n. gen. n. sp.6 Cockerell 1922 achilid planthopper
H.1145
 Hemiptera -
Aphis gurnetensis n. sp.3 Cockerell 1921 aphid
H.1124
 Hemiptera - Aphalaridae
"Livilla hooleyi n. sp." = Carsidarina hooleyi3
"Livilla hooleyi n. sp." = Carsidarina hooleyi3 Cockerell 1921 jumping plant louse
H.430/445, H.449
 Hemiptera - Aphrophoridae
? Aphrophora woodwardi n. sp.6 Cockerell 1922 spittle bug
H.619
 Hemiptera - Lygaeidae
Lygaeites amabilis n. sp.5 Cockerell 1921 milkweed bug
H.1398
 Hemiptera - Tingidae
? Celantia seposita n. sp.5 Cockerell 1921 lace bug
H.134, H.572
 Thysanoptera - Thripidae
Thripidae "indet. 1"14 Stevens 1829 thrips
BMNH In.25324, II.2854
Coccothrips hoffeinsorum14 Shmakov 2014 thrips
BMNH In.43474
 Thysanoptera - Aeolothripidae
Sinaeolothrips brodiei14 Cockerell 1917 thrips
BMNH In.24785, II.2855
Aeolothrips jarzembowskii n. sp.14 Shmakov 2014 thrips
BMNH In.25273
 Psocodea -
Wightipsocus intasi n. sp.2 Azar 2014 bark louse
BMNH In.24538
"Psocus acourti n. sp." = Wightipsocus acourti3 Cockerell 1921 bark louse
BMNH In.24366; In.20559, 25370, 25228, 64128, probably 42508/25677 in Azar 2014
Awightipsocus minimus2 Azar 2014 bark louse
BMNH In.25108
Awightipsocus megaloi2 Azar 2014 bark louse
BMNH In.24601, 24532
Awightipsocus jarzembowskii2 Azar 2014 bark louse
BMNH In.25436, 24579, 24788
 Blattodea -
Isoptera indet.12 termite
 Blattodea - Termitidae
Termitidae "sp.A"12 Latreille 1802 termite
NHML In.24603
 Orthoptera - Acrididae
Proschistocerca oligocaenica n. sp.16 Zeuner 1937 spur-throated grasshopper
BMNH In 24578, 24587, 25728, 24782; 20527 (collection uncertain)
 Orthoptera - Gryllidae
Gryllidae "indet. 13" Laicharting 1781 true crickets
BMNH In. 24779
Gryllidae "indet. 12" Laicharting 1781 true crickets
BMNH In. 24780
? Pteronemobius anglicus n. sp.16 Zeuner 1937 true crickets
BMNH In 24778
Gryllus schindewolfi n. sp.16 Zeuner 1937 field cricket
BMNH In 24774
Gryllus oligocenus n. sp. Cockerell 1921 field cricket
BMNH In. 24329
Gryllus cf. lineiceps Walker 1906 field cricket
NHM In. 24773
Gryllus vetus n. sp. Cockerell 1921 field cricket
BMNH In. 24328
 Orthoptera - Gryllotalpidae
"Gryllotalpa prima n. sp." = Pterotriamescaptor primus
"Gryllotalpa prima n. sp." = Pterotriamescaptor primus Cockerell 1921 mole cricket
BMNH In. 24327
 Orthoptera - Tettigoniidae
"Poekilloptera melanospila n. sp." = Archepseudophylla melanospila3
"Poekilloptera melanospila n. sp." = Archepseudophylla melanospila3 Cockerell 1921 katydid
BMNH In.24356 (H.48)
 Hymenoptera - Scoliidae
Palaeoscolia relicta n. gen. n. sp.1
Palaeoscolia relicta n. gen. n. sp.1 Antropov 2014 wasp
BMNH In.24527
 Hymenoptera - Vespidae
"? Polybia anglica n. sp." = Palaeopolybia anglica4, Eumeninae indet.1
"? Polybia anglica n. sp." = Palaeopolybia anglica4 Cockerell 1921 wasp
NHML In.24339 (H. 132)
Eumeninae indet.1 Leach 1815 potter wasp
BMNH In.43416
 Hymenoptera - Formicidae
Camponotus cockerelli1 Donisthorpe 1920 ant
BMNH In.24794, 24842, 24881, 24931, 25175
Oecophylla megarche9 Cockerell 1915 weaver ant
Leucotaphus donisthorpei1 Dlussky and Perfilieva 2014 ant
BMNH In.24920, 25124
Leucotaphus gurnetensis9 Cockerell 1915 ant
H.139, 168, 189, 377, 450; not In.24376, 24378
Emplastus hypolithus1 Cockerell 1915 ant
BMNH In.24368, 24373 (D. britannicus in Donisthorpe 1920)
Emplastus kozlovi1 Dlussky and Perfilieva 2014 ant
BMNH In.24372, 24936, 25112, 25141
"Dolichoderus britannicus" = Emplastus britannicus9 Cockerell 1915 ant
H.114, 262, 440, 497 not found. In.24461 (H.117); In.24367, 25083, 25085 in Antropov et al. 2014
"Dolichoderus ovigerus" = Emplastus britannicus9 Cockerell 1915 ant
H.374
"Emplastus emeryi n. gen. n. sp." = Emplastus britannicus9 Cockerell 1915 ant
BMNH In.24367 (H.129)
Dolichoderus vectensis1 Donisthorpe 1920 ant
BMNH In.24934
Paraphaenogaster hooleyana n. sp.1 Dlussky and Perfilieva 2014 ant
BMNH In.24963, 25082
Ponerites hooleyi n. sp.1 Dlussky and Perfilieva 2014 ant
BMNH In.24971, 24378
Taphopone aberrans n. sp.1 Dlussky and Perfilieva 2014 ant
BMNH In.24917, 24847, 24910, 24960
Taphopone microptera1 Dlussky and Perfilieva 2014 ant
BMNH In.24378
Taphopone petrosa n. sp.1 Dlussky and Perfilieva 2014 ant
BMNH In.24369, 24927, 25006
 Hymenoptera - Psenidae
Tytthopsen nanus n. gen. n. sp.1
Tytthopsen nanus n. gen. n. sp.1 Antropov 2014 apoid wasp
BMNH In.25568
 Hymenoptera - Pemphredonidae
Plisomena gigantea n. gen. n. sp.1
Plisomena gigantea n. gen. n. sp.1 Antropov 2014 aphid wasp
BMNH In.24789
 Hymenoptera - Braconidae
? Eubazus hooleyi n. sp.1, Dacnusites reductus n. sp.4, "Diospiloides hooleyi n. gen. n. sp." = Microtypus hooleyi4, Bassus quadrangularis1
? Eubazus hooleyi n. sp.1 Belokobylskij 2014 wasp
BMNH In.24790
Dacnusites reductus n. sp.4 Cockerell 1921 wasp
NHML In.24338 (H. 833)
"Diospiloides hooleyi n. gen. n. sp." = Microtypus hooleyi4 Cockerell 1921 wasp
NHML In.24337 (H. 882)
Bassus quadrangularis1 Brues 1933 wasp
BMNH In.24958
 Hymenoptera - Ichneumonidae
Acourtia perplexa n. gen. n. sp.4, Marjorietta gigantea n. sp.1, "Stilpnus" oligocenus n. sp.4, "Hemiteles" protervus1, ""Ichneumon" acourti n. sp." = Hemiteles acourti4, Lithapechtis fumosus n. gen. n. sp.4, Lithapechtis salmacidus n. sp.1, Eusterinx humalai n. sp.1, Pimplinae "indet. 1"1, "Itoplectis saxosus n. sp." = Itoplectis saxosa4, ? Polysphincta atavina n. sp.4
Acourtia perplexa n. gen. n. sp.4 Cockerell 1921 ichneumon wasp
NHML In.24336 (H. 858)
Marjorietta gigantea n. sp.1 Khalaim 2014 ichneumon wasp
NHML In.43477(1)
"Stilpnus" oligocenus n. sp.4 Cockerell 1921 ichneumon wasp
NHML In.24331 (H. 834)
"Hemiteles" protervus1 Khalaim 2014 ichneumon wasp
BMNH In.25765
""Ichneumon" acourti n. sp." = Hemiteles acourti4 Cockerell 1921 ichneumon wasp
NHML In.24332 (H. 1520)
Lithapechtis fumosus n. gen. n. sp.4 Cockerell 1921 ichneumon wasp
NHML In.24333 (H. 506)
Lithapechtis salmacidus n. sp.1 Khalaim 2014 ichneumon wasp
BMNH In.24839(24791)
Eusterinx humalai n. sp.1 Khalaim 2014 ichneumon wasp
BMNH In.25730
Pimplinae "indet. 1"1 Wesmael 1845 ichneumon wasp
BMNH In.24417(1)
"Itoplectis saxosus n. sp." = Itoplectis saxosa4 Cockerell 1921 ichneumon wasp
NHML In.24334 (H. 183)
? Polysphincta atavina n. sp.4 Cockerell 1921 ichneumon wasp
NHML In.24335 (H. 1036)
 Hymenoptera - Cynipidae
"Rhodites vetus n. sp." = Diplolepis vetus4
"Rhodites vetus n. sp." = Diplolepis vetus4 Cockerell 1921 wasp
NHML In.24341 (H. 701)
 Coleoptera - Carabidae
Pterostichus gurnetensis n. sp.3 Cockerell 1921 ground beetle
H.221
 Mecoptera - Bittacidae
Bittacus "sp.A"12, "Bittacus veternus n. sp." = Bittacus veternus12, "Panorpa veterna n. sp." = Bittacus veternus3
Bittacus "sp.A"12 Latreille 1802 hangingfly
NHML In.64537
"Bittacus veternus n. sp." = Bittacus veternus12 Cockerell 1921 hangingfly
NHML In.24330
"Panorpa veterna n. sp." = Bittacus veternus3 Cockerell 1921 hangingfly
H.1394
 Diptera - Culicidae
Culex protorhinus11 Cockerell 1915 mosquito
H.463, H.540, H.1014
Aedes protolepis11 Cockerell 1915 mosquito
H.468
 Diptera - Dixidae
Dixa priscula n. sp.3
Dixa priscula n. sp.3 Cockerell 1921 meniscus midge
H.708
 Diptera - Asilidae
Asilus gurnetensis n. sp.3 Cockerell 1921 robber fly
H.1174
Proctacanthus fractus n. sp.3 Cockerell 1921 robber fly
H.1536
 Diptera - Rhagionidae
Chrysopilus anglicus n. sp.3 Cockerell 1921 snipe fly
H.410
Chrysopilus stigmaticus n. sp.3 Cockerell 1921 snipe fly
H.1395
 Diptera - Tabanidae
"Chrysops vectensis n. sp." = Tabanus vectensis3
"Chrysops vectensis n. sp." = Tabanus vectensis3 Cockerell 1921 horse fly
H.46 (Branchopoda bed)
 Diptera - Anisopodidae
"Rhyphus hooleyi n. sp." = Sylvicola hooleyi3
"Rhyphus hooleyi n. sp." = Sylvicola hooleyi3 Cockerell 1921 wood gnat
H.456
 Diptera - Limoniidae
Gonomyia ferrea n. sp.3, Gonomyia grisea n. sp.3, Styringomyia extensa n. sp.8, "Empeda hyalina n. sp." = Cheilotrichia hyalina3
Gonomyia ferrea n. sp.3 Cockerell 1921 crane fly
H.835
Gonomyia grisea n. sp.3 Cockerell 1921 crane fly
H.1534
Styringomyia extensa n. sp.8 Cockerell and Haines 1921 crane fly
H.261
"Empeda hyalina n. sp." = Cheilotrichia hyalina3 Cockerell 1921 crane fly
H.1893.4
 Diptera - Tipulidae
Adelphomyia disjunctula n. sp.7, "Holorusia vasifera n. sp." = Ischnotoma vasifera7, Tipula hooleyi n. sp.7
Adelphomyia disjunctula n. sp.7 Cockerell and Haines 1921 crane fly
H.1398
"Holorusia vasifera n. sp." = Ischnotoma vasifera7 Cockerell and Haines 1921 crane fly
H.9
Tipula hooleyi n. sp.7 Cockerell and Haines 1921 crane fly
H.49
 Dicondylia -
Lepidoptera "sp.L"12 Linnaeus 1758 moth
NHML In.24506
Lepidoptera "sp.K"12 Linnaeus 1758 moth
NHML In.25219
Lepidoptera "sp.J"12 Linnaeus 1758 moth
NHML In.25157
Lepidoptera "sp.E"12 Linnaeus 1758 moth
NHML In.25251
Lepidoptera "sp.B"12 Linnaeus 1758 moth
NHML In.64540
 Lepidoptera - Hepialidae
Prohepialus sp.12 Piton 1940 moth
NHML In.17464 In.64528 In.64528 In.64539
 Lepidoptera - Nepticulidae
Nepticulidae indet.10 Stainton 1859 moth
NHML In.25512 In.25252
 Lepidoptera - Copromorphidae
Copromorpha fossilis n. sp.12 Jarzembowski 1980 moth
NHML In.25766
 Lepidoptera - Cossidae
Gurnetia durranti n. gen. n. sp.12
Gurnetia durranti n. gen. n. sp.12 Cockerell 1921 carpenter moth
NHML In.24324
 Trichoptera - Beraeidae
Beraeodes vectensis15 Cockerell 1921 caddisfly
BMNH In.24633, 25382, 25409, 26041
 Trichoptera - Polycentropodidae
? Plectrocnemia incompleta n. sp.15 Sukatsheva 2014 caddisfly
BMNH In.24595
 Trichoptera -
Paleodicella anomala n. gen. n. sp.15
Paleodicella anomala n. gen. n. sp.15 Sukatsheva 2014 caddisfly
BMNH In.24630
 Dicondylia -
"? Papilionidea indet." = Amphiesmenoptera12
"? Papilionidea indet." = Amphiesmenoptera12 Kiriakoff 1948 winged insect
NHML In.64545a,b
 Neuroptera - Hemerobiidae
Hemerobiidae "indet.A"12 Latreille 1802 brown lacewing
NHML In.43477
 Neuroptera - Mantispidae
"Promantispa relicta" = Vectispa relicta12
"Promantispa relicta" = Vectispa relicta12 Cockerell 1921 mantidfly
NHML In.24326, In.24597
 Odonata - Dysagrionidae
Dysagrionini "indet. A"13 Cockerell 1908 damselfly
BMNH I.9866/9718
 Odonata - Coenagrionidae
Coenagrionidae "indet. C"13 Kirby 1890 narrow-winged damselfly
BMNH In.24762
 Odonata - Lestidae
Lestes aff. regina13 Theobald 1937 spread-winged damselfly
BMNH In.24639
 Odonata - Megapodagrionidae
Oligoargiolestes oligocenum13 Kennedy 1925 flatwing
BMNH In.24645
 Odonata - Hypolestidae
Eohypolestes hooleyi n. gen. n. sp.13
Eohypolestes hooleyi n. gen. n. sp.13 Nel and Fleck 2014 damselfly
BMNH In.24653, 24769
 Odonata -
Calopterygiformia indet.13 Bechly 1996 damselfly
BMNH In.24759
 Odonata - Bolcathoridae
? Bolcathore "sp. A"13 Gentilini 2002 damselfly
BMNH In.24648
 Odonata - Aeshnidae
Anglogomphaeschna eocenica13 Nel and Fleck 2014 darner dragonfly
BMNH In.24649
"Gynacanthinae" "indet. A"13 Cockerell 1913 darner dragonfly
BMNH In.24638, 24751
 Odonata - Cordulephyidae
Neophya legrandi n. sp.13 Nel and Fleck 2014 true dragonfly
BMNH In.24606, 24641, 24757
 Odonata - Corduliidae
Corduliidae "indet. A"13 Selys-Longchamps 1850 green-eyed skimmer
BMNH In.24654/24760
 Odonata - Urothemistidae
Eomacrodiplax incompleta n. gen. n. sp.13
Eomacrodiplax incompleta n. gen. n. sp.13 Nel and Fleck 2014 true dragonfly
BMNH In.24770