Bore Lullingen 1, depth 42m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.5° N, 6.2° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Bislicher stage

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, fine-grained, silty sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1992

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 61904: authorized by Martin Aberhan, entered by Sabine Nürnberg on 07.07.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Foraminifera
 Lituolacea - Ataxophragmiidae
Martinottiella communis
Martinottiella communis Huang et al. 2019
 Rotalioidea - Elphidiidae
Elphidium inflatum
Globothalamea
 Rotaliida - Nonionidae
Nonion boueanum
 Rotaliida - Cibicididae
Cibicides lobatulus, Cibicides dutemplei
Cibicides dutemplei dOrbigny 1846
Tubothalamea
 Miliolida - Miliolidae
Spiroloculina badensis
Foraminifera
  - Spiroplectamminidae
Spiroplectammina carinata
Nodosariata
 Vaginulinida - Vaginulinidae
Lenticulina (Lenticulina) calcar, Lenticulina (Lenticulina) inornata, Lenticulina (Robulus) grandis
 Nodosariida - Nodosariidae
Dentalina bifurcata, Dentalina konincki, Nodosaria "sp. 1"
Dentalina konincki Reuss 1861
Nodosaria "sp. 1" Lamarck 1812
 Polymorphinida - Polymorphinidae
Globulina gibba, Guttulina problema, Pseudopolymorphina subnodosa, Pseudopolymorphina ovalis
Globulina gibba d'Orbigny 1832
incl. 1 specimen G. gibba for. punctata and 1 specimen of G. gibba for. rugosa
Guttulina problema d'Orbigny 1826
Foraminifera
 Lituolida - Lituolidae
Textularia gramen
Textularia gramen dOrbigny 1846
Polychaeta
  -
Polychaeta indet.1
Polychaeta indet.1 Grube 1850
Errantida indet. B
Scaphopoda
 Gadilida - Gadilidae
Gadila dingdenensis1
 Dentaliida - Dentaliidae
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1, Lentigodentalium pseudomutabile1
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
Gastropoda
 Epitonioidea - Epitoniidae
"Amaea (Scalina) subreticula" = Scalina2, sensu lato Turriscala straeleni2, Epitonium (Spiniscala) frondiculum2, Cirsotrema (Cirsotrema) sallomacense2, Cirsotrema (Cirsotrema) crassicostatum2
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
sensu lato Turriscala straeleni2 wentletrap
T. straeleni for. dingdensis for. nov.
Epitonium (Spiniscala) frondiculum2 Wood 1842 wentletrap
Cirsotrema (Cirsotrema) sallomacense2 wentletrap
C. sallomacense nordicica
Cirsotrema (Cirsotrema) crassicostatum2 Deshayes 1839 wentletrap
 Vanikoroidea - Eulimidae
Balcis (Balcis) alba2, Eulima (Eulima) bilineata2, Eulima (Eulima) glabra2
Balcis (Balcis) alba2 snail
incl. 6 specimens B. alba for. lactea
 Neotaenioglossa - Calyptraeidae
Calyptraea (Calyptraea) chinensis2
Calyptraea (Calyptraea) chinensis2 Linnaeus 1758 slipper shell
 Cerithioidea - Cerithiidae
"Bittium (Bittium) spina" = Bittium spina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Cerithioidea - Turritellidae
Turritella subangulata2, Turritella "sp. 2"2
Turritella "sp. 2"2 Lamarck 1799 turret shell
 Pseudomelanoidea - Pseudomelaniidae
sensu lato Inquisitor borealis3
 Sorbeoconcha - Naticidae
Naticidae indet.2, Sinum (Sinum) aquense2, "Natica neglecta" = Cochlis neglecta2, "Natica sensu lato tigrina" = Cochlis tigrina2, Euspira gottschei2, Euspira staringi2, Euspira helicina2, Tectonatica miopusilla2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
"Natica neglecta" = Cochlis neglecta2 Mayer 1858 moon snail
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
Euspira gottschei2 moon snail
Euspira staringi2 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
 Sorbeoconcha - Triforidae
Cerithiella (Cerithiella) genei2
 Sorbeoconcha - Triphoridae
Norephora (Norephora) fritschi2
 Sorbeoconcha - Cerithiopsidae
Cerithiopsis (Dizoniopsis) "sp. 2"2, Cerithiopsis (Cerithiopsis) vogeli2
 Sorbeoconcha - Rissoidae
sensu lato Alvania holsatica2, Ovirissoa westfalica2
 Sorbeoconcha - Cassidae
"Semicassis (Echinophoria) bicoronata" = Echinophoria2
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
 Sorbeoconcha - Ranellidae
"Charonia (Sassia) tarbelliana" = Sassia2
"Charonia (Sassia) tarbelliana" = Sassia2 triton shell
 Neogastropoda - Costellariidae
"Vexillum (Uromitra) aciculum" = Uromitra3, "Vexillum (Uromitra) gliberti" = Uromitra3, Orthosurcula steinvorthi3
"Vexillum (Uromitra) aciculum" = Uromitra3 ribbed mitre
"Vexillum (Uromitra) gliberti" = Uromitra3 ribbed mitre
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Olividae
Oliva (Strephona) dufresnei2
Oliva (Strephona) dufresnei2 Basterot 1825 olive snail
 Neogastropoda - Cancellariidae
Babylonella fusiformis2, Sveltia varicosa2, Cancellaria (Merica) contorta2
Sveltia varicosa2 snail
S. varicosa paucicostata
Cancellaria (Merica) contorta2 snail
C. contorta gelriana
 Neogastropoda - Mitridae
Mitra (Tiara) substriatula3
Mitra (Tiara) substriatula3 d'Orbigny 1852 mitre shell
 Neogastropoda - Fasciolariidae
Streptodictyon sexcostatus2, ? Exilifusus attenuatus2
 Neogastropoda - Nassariidae
Nassarius bocholtensis2, Nassarius tenuistriatus2, Nassarius voorthuyseni2, Nassarius woodwardi2, Nassarius karinae2
Nassarius karinae2 Wienrich 2001 snail
 Neogastropoda - Columbellidae
Mitrella (Macrurella) nassoides2, Anachis (Costoanachis) terebralis2, Anachis (Costoanachis) "sp. 1"2, Anachis (Costoanachis) "sp. 4"2, Anachis (Costoanachis) "sp. 6"2
 Neogastropoda - Volutidae
Scaphella (Scaphella) bolli2
 Neogastropoda - Muricidae
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2, "Murex (Haustellum) inornatus" = Haustellum2, Hadriania coelata2, Lyrotyphis (Eotyphis) fistulosus2, Typhis (Typhis) ? pungens2
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2 Bronn 1831 murex snail
"Murex (Haustellum) inornatus" = Haustellum2 murex snail
Hadriania coelata2 Dujardin 1837 murex snail
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
 Neogastropoda - Clavatulidae
Clavatula boreointerrupta3
 Neogastropoda - Terebridae
Strioterebrum hoernesi3
 Neogastropoda - Conidae
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Mangeliidae
sensu lato Mangelia luellingensis3, Benthomangelia aff. obtusangula3, sensu lato Benthomangelia aff. xeniae3, Sorgenfreispira tenella3
sensu lato Mangelia luellingensis3 Wienrich and Janssen 2007 snail
Sorgenfreispira tenella3 Conrad 1848 snail
 Neogastropoda - Raphitomidae
Teretia anceps3, sensu lato Teretia "sp. 2"3, Favriella sinuosula3, Metuonella grippi3, Raphitoma "sp. 2"3, Pleurotomella mioweberi3
sensu lato Teretia "sp. 2"3 Norman 1888 snail
Raphitoma "sp. 2"3 Monterosato 1884 snail
 Neogastropoda - Turridae
Perrona obliquiplicatula3, Boreodrillia hosiusi3, Nitidiclavus maitrejus3, sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula stoffelsii3, Gemmula (Gemmula) denticula3, Gemmula (Gemmula) zimmermanni3, Fusiturris aquensis3, Fusiturris duchastelii3
sensu lato Unedogemmula boreoturricula3 turrid
incl. 11 specimens U. (s. lat.) boreoturricula x hanseata ?
Gemmula (Gemmula) denticula3 turrid
G. denticula borealis
Fusiturris duchastelii3 turrid
F. duchastelii flexiplicata
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Clathurellidae
Pleurotomoides luisae3, Pleurotomoides simplex3
 Neogastropoda - Borsoniidae
Asthenotoma pannoides3, Asthenotoma pannus3, Bathytoma jugleri3
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Sorbeoconcha - Vitrinellidae
Circulus praecedens2, Circulus partimstriatus2, Circulus hennei2
Circulus praecedens2 snail
C. praecedens for. praecedens
Circulus hennei2 Glibert 1952 snail
 Heterostropha - Pyramidellidae
Chrysallida (Pyrgulina) pygmaea3, Chrysallida (Parthenina) cimbrica3, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3, Turbonilla (? Strioturbonilla) "sp. 8"3, Turbonilla (Turbonilla) gastaldi3, Turbonilla (Mormula) "sp. 5"3, Pyramidella (Voluspa) elata3, Pyramidella (Voluspa) plicosa3, Eulimella (Eulimella) acicula3, Eulimella (Eulimella) scillae3, Evalea sp.3, Odostomia (Brachystomia) cf. sallespissiensis3, Odostomia (Megastomia) cf. conoidea3, Odostomia (Megastomia) tuexeni3, Syrnola hoernesi3, Syrnola neumayri3, Syrnola "sp. 2"3
Chrysallida (Pyrgulina) pygmaea3 Grateloup 1838 snail
incl. 4 specimens Chrysallida pygmaea f. belgica
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
sensu JANSSEN
"Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3 snail
sensu JANSSEN
Turbonilla (Turbonilla) gastaldi3 snail
sensu JANSSEN
Turbonilla (Mormula) "sp. 5"3 Adams 1864 snail
Evalea sp.3 Adams 1860 snail
Syrnola "sp. 2"3 Adams 1860 snail
 Heterostropha - Murchisonellidae
Ebala (Ebala) vandervlerki3
 Heterostropha - Architectonicidae
Architectonica (Architectonica) simplex3
 Heterostropha - Acteonidae
Acteon (Acteon) semistriatus3
Acteon (Acteon) semistriatus3 snail
spelled as 'Actaeon'
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) buccinea3
 Trochoidea - Trochidae
Calliostoma (Ampullotrochus) ? laureatum2
 Trochoidea - Skeneidae
Skenea minuta2
 Lepetellida - Lepetellidae
Lepetella compressiuscala2
 Opisthobranchia - Retusidae
Retusa (Cylichnina) elongata3, Volvulella (Volvulella) acuminata3
 Opisthobranchia - Cylichnidae
Cylichna (Cylichna) pseudoconvoluta3
 Opisthobranchia -
Roxania utriculus3
 Thecosomata - Cavoliniidae
Vaginella austriaca3
 Thecosomata - Limacinidae
Limacina miorostralis3, Limacina valvatina3
Limacina valvatina3 Reuss 1867 snail
Bivalvia
 Nuculanida - Nuculanidae
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
 Nuculanida - Yoldiidae
Yoldia (Yoldia) glaberrima1, "Portlandia (Yoldiella) pygmaea" = Yoldiella1
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
 Nuculida - Nuculidae
Nuculoma haesendoncki1, Nucula (Nucula) nucleus1
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
Nucula (Nucula) nucleus1 Linnaeus 1758 nut clam
 Thraciida - Thraciidae
Thracia (Thracia) aff. inflata1
 Hiatellida - Hiatellidae
"Hiatella (Hiatella) arctica" = Hiatella arctica1, Panopea (Panopea) meynardi1
"Hiatella (Hiatella) arctica" = Hiatella arctica1 Linnaeus 1767 clam
H. (H.) arctica arctica
 Pholadida - Teredinidae
Teredinidae indet.1
Teredinidae indet.1 Rafinesque 1815 clam
 Pholadida - Corbulidae
Corbula (Varicorbula) gibba1
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Pholadida - Myidae
Sphenia pusilla1
Sphenia pusilla1 Sorgenfrel 1958 softshell clam
 Cardiida - Leptonidae
Lepton (? Lepton) transversarium1
 Cardiida - Arcticidae
Arctica islandica1
Arctica islandica1 Linnaeus 1767 clam
A. islandica islandica
 Cardiida - Ungulinidae
Diplodonta (Diplodonta) rotundata1
 Cardiida - Glossidae
Glossus (Glossus) lunulatus1
 Cardiida - Kelliellidae
Spaniodontella nitida1
 Cardiida - Veneridae
Venus (Ventricoloidea) multilamella1, Callista chione1, "Gouldia minima" = Gouldia (Gouldia) minima1
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
Callista chione1 Linnaeus 1758 venus clam
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
 Cardiida - Mactridae
Spisula elliptica1
Spisula elliptica1 Brown 1827 clam
 Cardiida - Solecurtidae
Solecurtus basteroti1, Azorinus (Azorinus) chamasolen1
Azorinus (Azorinus) chamasolen1 clam
A. chamasolen miocaenicus
 Cardiida - Tellinidae
Angulus (Laceolina) benedeni1
Angulus (Laceolina) benedeni1 Nyst and Westendorp 1839 tellin clam
A. benedeni fallax
 Cardiida - Semelidae
sensu lato Abra lehmanni1, Abra (? Abra) sorgenfreii1
 Cardiida - Cardiidae
"Habecardium subturgidum" = Laevicardium (Habecardium)1, Habecardium dingdense1, Parvicardium straeleni1, Papillicardium papillosum1
"Habecardium subturgidum" = Laevicardium (Habecardium)1 cockle
H. subturgidum subturgidum
Habecardium dingdense1 Lehmann 1885 cockle
Papillicardium papillosum1 Poli 1781 cockle
 Lucinida - Lucinidae
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1 Nyst 1861 clam
C. droueti droueti
 Carditida - Carditidae
Cyclocardia sp.1
Cyclocardia sp.1 Conrad 1867 clam
 Carditida - Astartidae
Astarte (Astarte) radiata1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata 29 specimens, Astarte radiata fa. convexior 31 specimens
 Pectinida - Pectinoidae
Korobkovia woodi1, Pseudamussium lilli1, Aequipecten seniensis1
Pseudamussium lilli1 Pusch 1837 scallop
 Pectinida - Anomiidae
Pododesmus (Monia) anitae1, "Pododesmus (Heteranomia) squamula" = Heteranomia squamula1
Pododesmus (Monia) anitae1 Janse and Janssen 1983 scallop
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1 Linnaeus 1758 scallop
 Ostreida - Pinnidae
Atrina pectinata1
Atrina pectinata1 Linnaeus 1758 oyster
 Arcida - Arcidae
Scapharca (Scapharca) diluvii1
 Arcida - Limopsidae
Limopsis (Limopsis) aurita1, "Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
Rhynchonellata
 Rhynchonellida - Cryptoporidae
Cryptopora sp.1
Cryptopora sp.1 Jeffreys 1869
 Terebratulida - Terebratulidae
Terebratula sp.1
Terebratula sp.1 Müller 1776
Gymnolaemata
 Cheilostomata - Lunulitidae
Lunulites voigti1, Lunulites burdigalensis1
 Cheilostomata - Calloporidae
Calloporidae indet.1
Calloporidae indet.1 Norman 1903
 Cheilostomata - Membraniporidae
Discoporella umbellata1, Reussirella haidingeri1
Discoporella umbellata1 Defrance 1823
Anthozoa
 Scleractinia - Flabellidae
Flabellum sp., Flabellum cristatum, Flabellum tuberculatum
Flabellum sp. Lesson 1831 stony coral
Flabellum cristatum Milne-Edwards and Haime 1851 stony coral
Flabellum tuberculatum Keferstein 1859 stony coral
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007