Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)
• coordinate based on nearby landmark
• local area-level geographic resolution
When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)
• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.
• group of beds-level stratigraphic resolution
Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone
Size classes: macrofossils, mesofossils
Preservation: adpression
Collection methods: Repository: Beijing Museum of Natural History. This collection contains specimens labeled with five-digit numbers starting with 79... or 80...
Primary reference: Y. C. Hong. 1984. Fossil insects in the diatoms of Shanwang. Bulletin of the Tianjin Institute of Geology and Mineral Resources 8:1-15 [D. Smith/E. Leckey/M. Clapham]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 128550: authorized by Matthew Clapham, entered by Matthew Clapham on 11.06.2012, edited by Erin Leckey
Creative Commons license: CC BY (attribution)
Taxonomic list
Insecta | |
Tipula insolita n. sp.5
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Sciara brevipetiola n. sp.5
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Clothonopsis miocenica n. gen. n. sp.4, Plecia solaris n. sp.5, Plecia ludongensis n. sp.1, Bibio ventricosus n. sp.5, Bibio deformatus n. sp.5, Bibio expansus5, Bibio brunneus n. sp.5, Bibio ruptineurus n. sp.5, "Fungivora bullata n. sp." = Bibio bullatus5, "Bibio miocenica n. sp." = Bibio miocenicus1, Bibio shanwangensis1, Bibio cratopodus5
Bibio ventricosus n. sp.5 Zhang 1989 march fly 79033 (Bibio shanwangensis in Hong, 1985 - pl 7, figs 1, 2, 3)
Bibio deformatus n. sp.5 Zhang 1989 march fly 79057 (Bibio shanwangensis in Hong, 1985 - pl 4, fig 1)
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Apanechura asecta n. sp.5
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Cupes longus n. sp.4
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? Carabus ovalus n. sp.2, Calosoma cf. maderae5, "Shanwangicarabus furvus n. sp." = Chlaenius furvus2, "Shanwangicarabus paucinus n. sp." = Chlaenius furvus2, Scarites mancus n. sp.5
"Shanwangicarabus furvus n. sp." = Chlaenius furvus2 Hong 1984 ground beetle 80125, 80106, 80126, 480183
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Cerambycidae "indet. 1"5, Alifasciola daedalea n. sp.5, Mesocacia pulla n. sp.5, Sinocalosoma longicornuta n. sp.2, Mesosa soteria n. sp.5
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"Sinocurculia granulata n. sp." = Atactogaster granulata, "Sinocurculia shanwangensis n. sp." = Atactogaster shanwangensis2, Hylobius medianus n. sp.5, Hylobius aff. niitakensis5, Hylobius plenus5
"Sinocurculia shanwangensis n. sp." = Atactogaster shanwangensis2 Hong 1985 snout beetle 80128, 78001, 80107, 80134, 80173, 80174
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Longifistulia parva n. gen. n. sp.3
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"Engonius alviolatus" = Gerstaeckerus alviolatus5
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? Epicauta rufithoraca n. sp.5
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Sinocistela siphla n. gen. n. sp.5
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Scarabaeoidea "indet. 1"5 Latreille 1802 scarab beetle 79045 (Magniscarabaeus furvus in Hong & Wang - pl 17, fig 2)
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Geotrupidae indet.5 Latreille 1802 earth-boring dung beetle 79107 (Magniscarabaeus brunneus in Hong, 1985 - pl 20, fig 3)
"Obitiscarabaeus jiaoyanshanense n. gen. n. sp." = Geotrupes jiaoyanshanense Hong 1984 earth-boring dung beetle 79114/5
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Aliscarabaeus granulatus n. sp.
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Holotrichia spatha n. sp.5, Anomala sp.5, "Magniscarabaeus brunneus n. gen. n. sp." = Anomala palaeobrunnea5, Anomala lochmocola n. sp.5, Anomala synemosyna n. sp.5, "Magniscarabaeus furvus" = Anomala furva2
"Magniscarabaeus brunneus n. gen. n. sp." = Anomala palaeobrunnea5 Krell 2000 scarab beetle 79105; seven including 80101, 80120, 80172, 80188, 80184, 80194, mentioned in Hong & Wang 1987
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Lathrobium homorum n. sp.5
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"Procarabus longipes n. sp." = Cistelites longipes5
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"Procimbex shandongense n. sp." = Abia shandongensis1, Abia paurocephala n. sp.5, "Procimbex shanwangensis n. sp." = Abia shanwangensis
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Luia lacaris n. gen. n. sp.5
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Apis miocenica n. sp.
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"Sinostigma spinalata n. sp." = Pemphredon spinalatum
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"Brachyopteris shanwangensis n. sp." = Tiphia shanwangensis
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"Pseudoperla shanwangensis n. sp." = Eumenes shanwangensis
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Elis pyrula n. sp.5
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Paraphaenogaster shanwangensis n. sp., "Heteromyrmex atopogaster n. gen. n. sp." = Zhangidris atopogaster5, "Shanwangella palaeoptera n. gen. n. sp." = Liometopum palaeopterum5, Liometopum potamophilum n. sp.5, ? Tapinoma baculum n. sp.5, "Liometopum jiaoyanshanense n. sp." = Dolichoderus jiaoyanshanensis5, Formica linquensis n. sp.5, "Camponotus microthoracus n. sp." = Camponotites microthoracus5, "Rabidia xiejiaheensis n. sp." = Camponotites xiejiaheensis, "Camponotus longus n. sp." = Camponotites longus5, "Camponotus shanwangense" = Camponotites shanwangensis, "Camponotus curviansatus n. sp." = Camponotites curviansatus5, ? Lasius truncatus n. sp.5, Lasius validus n. sp.5, Lasius vetulus5, Camponotus mitis5
"Heteromyrmex atopogaster n. gen. n. sp." = Zhangidris atopogaster5 Zhang 1989 ant 79054 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 3)
"Shanwangella palaeoptera n. gen. n. sp." = Liometopum palaeopterum5 Zhang 1989 ant 79051 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 2)
Liometopum potamophilum n. sp.5 Zhang 1989 ant 79036 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 6)
"Liometopum jiaoyanshanense n. sp." = Dolichoderus jiaoyanshanensis5 Hong 1985 ant 79087; 80196 in Hong & Wang 1985
Formica linquensis n. sp.5 Zhang 1989 ant 79058 (Paraphaenogaster shanwangense in Hong, 1985 - pl 23, fig 4)
"Camponotus shanwangense" = Camponotites shanwangensis Hong 1984 ant 79031; 79043, 79076 (Liometopum jiaoyanshanense in Hong, 1985); ten specimens including 80179 in Hong & Wang 1985
"Camponotus curviansatus n. sp." = Camponotites curviansatus5 Zhang 1989 ant 79042 (Camponotus shanwangensis in Hong, 1985 - pl 27, fig 1)
Lasius validus n. sp.5 Zhang 1989 ant 79121 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 4)
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"Megaxyela xiejiaheensis n. sp." = Xyelecia xiejiaheensis
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"Cinara reconditivenia n. sp." = Cinara (Cinara) reconditivenia5
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"Coriopsis miocenica n. sp." = Coreiopsis miocenica3, "Coriopsis parva n. sp." = Coreiopsis parva3, Coreiopsis punctata n. gen. n. sp., Shanwangicoris longa n. gen. n. sp.3, Cloresmus shanwangi n. sp.5, Gonocerus shandongensis n. sp.5
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Tarsonecta mecopoda n. gen. n. sp.5
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Coroides xiejiaheense n. gen. n. sp.
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Manocerus stagnans n. gen. n. sp.5
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"Oncotympana lapidescens n. sp." = Tanyocicada lapidescens5
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"Oxycephala xiejiaheensis" = Limois shanwangensis5, "Fulgoropsis fusca n. gen. n. sp." = Limois shanwangensis, Aphaena lithoecia n. sp.5
"Oxycephala xiejiaheensis" = Limois shanwangensis5 Hong 1979 fulgorid planthopper 79026, 79079, 79080
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Jagoria sinica n. sp.5
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Arachnida | |
"Retina robusta" = Heteropoda robusta6
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"Eocryphoeca parva" = Tetragnatha parva6
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