PBDB Collection

Boryslawia (Pleistocene of Ukraine)

Where: Ukraine (49.3° N, 23.4° E: paleocoordinates 49.3° N, 23.4° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Late/Upper Pleistocene (0.1 - 0.0 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: terrestrial; unlithified, carbonaceous siltstone

• Bituminous (oil-impregnated) silts with ~35% organic matter

Size classes: macrofossils, mesofossils

Preservation: original chitin

Reposited in the NHMW

• Repository: Lomnicki's collection is at Lviv Museum (some specimens in d'Orchymont's collection in Brussels) according to Angus (1973). A few specimens in Geologisch-Paläontologischen Abteilung of NHMW (listed by Ponomarenko & Schultz, 1988).

Primary reference: A. M. Lomnicki. 1894. Pleistoceńskie owady z Borysławia. Muzeum Imienia Dzieduszyckich we Lwowie 4:1-116 [M. Clapham/M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 153476: authorized by Matthew Clapham, entered by Matthew Clapham on 21.12.2013

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Diptera - Muscidae	Muscidae indet. Muscidae indet. Latreille 1802 house fly
Diptera - Dolichopodidae	Chrysotus sp. Chrysotus sp. Meigen 1824 long-legged fly
Hemiptera - Corixidae	"Corisa boryslavica n. sp." = Corixa boryslavica, "Corisa glacialis n. sp." = Corixa glacialis, "Corisa (Glaenocorisa) horvathi n. sp." = Sigara (Arctocorisa) horvathi, "Corisa ozokeritica n. sp." = Sigara (Callicorixa) producta "Corisa boryslavica n. sp." = Corixa boryslavica Lomnicki 1894 water boatman "Corisa glacialis n. sp." = Corixa glacialis Lomnicki 1894 water boatman "Corisa (Glaenocorisa) horvathi n. sp." = Sigara (Arctocorisa) horvathi Lomnicki 1894 water boatman "Corisa ozokeritica n. sp." = Sigara (Callicorixa) producta Reuter 1880 water boatman NHMW 1907/21
Lepidoptera -	Noctuites sp.⁴ Noctuites sp.⁴ Heer 1849 moth
Coleoptera - Carabidae	"Diachila arctica" = Diacheila arctica, Blethisa pleistocenica n. sp., Bembidium (Campa) subcontaminatum n. sp., Bembidium (Campa) boryslavicum n. sp., Bembidium (Peryphus) tibiale, "Bembidium (Testidium) bipunctatum" = Bembidion (Testedium) bipunctatum, Patrobus gasiorowskii n. sp., Harpalus pleistocenicus n. sp., Amara (Amara) subaenea n. sp., Amara (Cyrtonotus) boryslavica n. sp., Pterostichus (Steropus) aethiops, Pterostichus (Cryobius) blanduloides n. sp., Pterostichus (Pseudomaseus) anthracinus, Carabus kollari, Carabus (Melancarabus) dzieduszyckii n. sp., Carabus praearvensis n. sp., Carabus maeandroides n. sp., Carabus (Megadontus) praeviolaceus n. sp., "Platynus (Europhilus) micans" = Agonum micans "Diachila arctica" = Diacheila arctica Gyllenhal 1810 ground beetle Blethisa pleistocenica n. sp. Lomnicki 1894 ground beetle Bembidium (Campa) subcontaminatum n. sp. Lomnicki 1894 ground beetle Bembidium (Campa) boryslavicum n. sp. Lomnicki 1894 ground beetle Bembidium (Peryphus) tibiale Duftschmid 1812 ground beetle "Bembidium (Testidium) bipunctatum" = Bembidion (Testedium) bipunctatum Linnaeus 1761 ground beetle Patrobus gasiorowskii n. sp. Lomnicki 1894 ground beetle Harpalus pleistocenicus n. sp. Lomnicki 1894 ground beetle Amara (Amara) subaenea n. sp. Lomnicki 1894 ground beetle Amara (Cyrtonotus) boryslavica n. sp. Lomnicki 1894 ground beetle Pterostichus (Steropus) aethiops Panzer 1797 ground beetle Pterostichus (Cryobius) blanduloides n. sp. Lomnicki 1894 ground beetle Pterostichus (Pseudomaseus) anthracinus Illiger 1798 ground beetle Carabus kollari Palliardi 1825 ground beetle var. comptus Carabus (Melancarabus) dzieduszyckii n. sp. Lomnicki 1894 ground beetle Carabus praearvensis n. sp. Lomnicki 1894 ground beetle Carabus maeandroides n. sp. Lomnicki 1894 ground beetle Carabus (Megadontus) praeviolaceus n. sp. Lomnicki 1894 ground beetle "Platynus (Europhilus) micans" = Agonum micans Nicolai 1822 ground beetle
Coleoptera - Gyrinidae	"Gyrinus praemarinus n. sp." = Gyrinus marinus, Gyrinus praeopacus n. sp. "Gyrinus praemarinus n. sp." = Gyrinus marinus Gyllenhal 1808 whirligig beetle NHMW 1907/21 Gyrinus praeopacus n. sp. Lomnicki 1894 whirligig beetle NHMW 1907/21
Coleoptera - Dytiscidae	Hydroporus praenivalis n. sp., Hydroporus pleistocenicus n. sp., Hydroporus praenigrita n. sp., Hydroporus lapponum, Hydroporus praedorsalis n. sp., Hydroporus subarcticus n. sp., Hydroporus clessini n. sp., Hydroporus sandbergeri n. sp., "Hydroporus (Oreodytes) borealis" = Oreodytes borealis, "Coelambus niedzwiedzkii n. sp." = Hygrotus niedzwiedzkii, "Coelambus picipoides n. sp." = Hygrotus picipoides, "Coelambus ganglbaueri n. sp." = Hygrotus ganglbaueri, "Coelambus dzieduszyckii n. sp." = Hygrotus dzieduszyckii, "Coelambus ozokeriticus n. sp." = Hygrotus ozokeriticus, "Coelambus latefasciatus n. sp." = Hygrotus latefasciatus, Ilybius boryslavicus n. sp., Agabus parvulus n. sp., Agabus reitteri n. sp., Agabus congeneroides n. sp., Agabus niedzwiedzkii n. sp., Agabus corticeus n. sp., Dytiscus lapponicus, Acilius praesulcatus n. sp., "Rantus praesuturellus n. sp." = Rhantus praesuturellus, Cymatopterus striatus, Cymatopterus dolabratus Hydroporus praenivalis n. sp. Lomnicki 1894 predaceous diving beetle Hydroporus pleistocenicus n. sp. Lomnicki 1894 predaceous diving beetle Hydroporus praenigrita n. sp. Lomnicki 1894 predaceous diving beetle NHMW 1907/21 Hydroporus lapponum Gyllenhal 1808 predaceous diving beetle Hydroporus praedorsalis n. sp. Lomnicki 1894 predaceous diving beetle Hydroporus subarcticus n. sp. Lomnicki 1894 predaceous diving beetle NHMW 1907/21 Hydroporus clessini n. sp. Lomnicki 1894 predaceous diving beetle Hydroporus sandbergeri n. sp. Lomnicki 1894 predaceous diving beetle NHMW 1907/21 "Hydroporus (Oreodytes) borealis" = Oreodytes borealis Gyllenhal 1826 predaceous diving beetle "Coelambus niedzwiedzkii n. sp." = Hygrotus niedzwiedzkii Lomnicki 1894 predaceous diving beetle "Coelambus picipoides n. sp." = Hygrotus picipoides Lomnicki 1894 predaceous diving beetle "Coelambus ganglbaueri n. sp." = Hygrotus ganglbaueri Lomnicki 1894 predaceous diving beetle "Coelambus dzieduszyckii n. sp." = Hygrotus dzieduszyckii Lomnicki 1894 predaceous diving beetle "Coelambus ozokeriticus n. sp." = Hygrotus ozokeriticus Lomnicki 1894 predaceous diving beetle One at NHMW 1907/21 "Coelambus latefasciatus n. sp." = Hygrotus latefasciatus Lomnicki 1894 predaceous diving beetle Ilybius boryslavicus n. sp. Lomnicki 1894 predaceous diving beetle Agabus parvulus n. sp. Lomnicki 1894 predaceous diving beetle Agabus reitteri n. sp. Lomnicki 1894 predaceous diving beetle Agabus congeneroides n. sp. Lomnicki 1894 predaceous diving beetle NHMW 1907/21 Agabus niedzwiedzkii n. sp. Lomnicki 1894 predaceous diving beetle Agabus corticeus n. sp. Lomnicki 1894 predaceous diving beetle NHMW 1907/21 Dytiscus lapponicus Gyllenhal 1808 predaceous diving beetle at least 4 (from plates) and 1 var. brachypterus Acilius praesulcatus n. sp. Lomnicki 1894 predaceous diving beetle "Rantus praesuturellus n. sp." = Rhantus praesuturellus Lomnicki 1894 predaceous diving beetle One at NHMW 1907/21; at least one other Cymatopterus striatus Linnaeus 1758 predaceous diving beetle Cymatopterus dolabratus Paykull 1798 predaceous diving beetle
Coleoptera - Helophoridae	"Helophorus (Gephelophorus) lomnicki n. sp." = Helophorus (Gephelophorus) sibiricus², "Helophorus (Trichelophorus) pleistocenicus n. sp." = Helophorus (Gephelophorus) sibiricus, "Helophorus (Meghelophorus) aquaticus" = Helophorus (Meghelophorus) aquaticus¹, "Helophorus (Praehelephorus) dzieduszyckii n. sp." = Helophorus (Meghelophorus) aquaticus, "Helophorus (Rhopalhelophorus) praenanus n. sp." = Helophorus praenanus, "Helophorus (Rhopalhelophorus) polonicus n. sp." = Helophorus praenanus, "Helophorus (Rhopalhelophorus) kuwerti n. sp." = Helophorus oblongus "Helophorus (Gephelophorus) lomnicki n. sp." = Helophorus (Gephelophorus) sibiricus² Motschulsky 1860 water scavenger beetle Part of Lomnicki's H. pleistocenicus "Helophorus (Trichelophorus) pleistocenicus n. sp." = Helophorus (Gephelophorus) sibiricus Motschulsky 1860 water scavenger beetle Paratype NHMW 1907/21; 107 specimens mentioned by Angus 1973 "Helophorus (Meghelophorus) aquaticus" = Helophorus (Meghelophorus) aquaticus¹ Linnaeus 1758 water scavenger beetle Lectotype of H. pleistocenicus var. obsoletus (in d'Orchymont coll.); others belong to Lomnicki's H. pleistocenicus or H. pleistocenus var. obsoletus material "Helophorus (Praehelephorus) dzieduszyckii n. sp." = Helophorus (Meghelophorus) aquaticus Linnaeus 1758 water scavenger beetle 8 in Lviv collection, 1 in d'Orchymon collection "Helophorus (Rhopalhelophorus) praenanus n. sp." = Helophorus praenanus Lomnicki 1894 water scavenger beetle Paratype NHMW 1907/21; 23 specimens in Lviv; 1 in d'Orchymony coll. "Helophorus (Rhopalhelophorus) polonicus n. sp." = Helophorus praenanus Lomnicki 1894 water scavenger beetle "Helophorus (Rhopalhelophorus) kuwerti n. sp." = Helophorus oblongus Le Conte 1850 water scavenger beetle 7 paratypes in NHMW 1907/21; 7 specimens including lectotype in Lviv; 1 paratype in d'Orchymont coll.
Coleoptera - Hydrophilidae	Sphaeridium scarabaeoides, Laccobius flachii n. sp., Hydrobius fuscipes, "Philhydrus pleistocenicus n. sp." = Enochrus pleistocenicus Sphaeridium scarabaeoides Linnaeus 1758 water scavenger beetle Laccobius flachii n. sp. Lomnicki 1894 water scavenger beetle Hydrobius fuscipes Linnaeus 1758 water scavenger beetle "Philhydrus pleistocenicus n. sp." = Enochrus pleistocenicus Lomnicki 1894 water scavenger beetle
Coleoptera - Scarabaeidae	Aphodius rhinocerontis n. sp., Aphodius granarius, Aphodius boryslavicus n. sp., Aphodius rufipes, Aphodius subater n. sp., Aphodius ruthenus n. sp. Aphodius rhinocerontis n. sp. Lomnicki 1894 scarab beetle Aphodius granarius Linnaeus 1767 scarab beetle Aphodius boryslavicus n. sp. Lomnicki 1894 scarab beetle Aphodius rufipes Linnaeus 1758 night-flying dung beetle Aphodius subater n. sp. Lomnicki 1894 scarab beetle Aphodius ruthenus n. sp. Lomnicki 1894 scarab beetle
Coleoptera - Staphylinidae	Silpha (Blitophaga) vetusta n. sp., Silpha (Blitophaga) reitteri n. sp., Gymnusa antiqua n. sp. Silpha (Blitophaga) vetusta n. sp. Lomnicki 1894 rove beetle Silpha (Blitophaga) reitteri n. sp. Lomnicki 1894 rove beetle Gymnusa antiqua n. sp. Lomnicki 1894 rove beetle
Coleoptera - Dryopidae	Parnus prolifericornis Parnus prolifericornis Fabricius 1792 long toed water beetle
Coleoptera - Curculionidae	"Hypera praecomata n. sp." = Eremochorus concinnus, Hypera glacialis n. sp., Donus nidensis³, Sitona ovipennis³, "Otiorrhynchus blanduloides n. sp." = Otiorhynchus blanduloides, "Otiorrhynchus uhligii n. sp." = Otiorhynchus obtusus, Otiorhynchus rugosus³, "Lepyrus frigidus n. sp." = Lepyrus nordenskjoeldi, Coniocleonus ferrugineus³ "Hypera praecomata n. sp." = Eremochorus concinnus Boheman 1842 snout beetle Hypera glacialis n. sp. Lomnicki 1894 snout beetle NHMW 1907/21 Donus nidensis³ Mazur and Petryszak 1981 snout beetle part of Lomnicki's Hypera glacialis Sitona ovipennis³ Hochhuth 1851 snout beetle ssp. borealis (part of Lomnicki's Hypera glacialis) "Otiorrhynchus blanduloides n. sp." = Otiorhynchus blanduloides Lomnicki 1894 snout beetle "Otiorrhynchus uhligii n. sp." = Otiorhynchus obtusus Boheman 1842 snout beetle Otiorhynchus rugosus³ Hummel 1827 snout beetle "Lepyrus frigidus n. sp." = Lepyrus nordenskjoeldi Faust 1885 snout beetle Coniocleonus ferrugineus³ Fahraeus 1842 snout beetle "Lepyrus diluvianus" label of Lomnicki (unpublished)
Coleoptera - Chrysomelidae	Psylliodes polonica n. sp., Donacia jaroslavii n. sp., Chrysomela lichenis, Chrysomela hilberi n. sp. Psylliodes polonica n. sp. Lomnicki 1894 flea beetle Donacia jaroslavii n. sp. Lomnicki 1894 leaf beetle Chrysomela lichenis Richter 1820 leaf beetle Chrysomela hilberi n. sp. Lomnicki 1894 leaf beetle
Orthoptera - Acrididae	Stenobothrus sp. Stenobothrus sp. Fischer 1853 short-horned grasshopper

¹Angus 1973; ²d'Orchymont 1927; ³Kuska 1992; ⁴Sohn et al. 2012