PPP# 359 - Escudo de Veraguas [Escudo de Veraguas Fm] (Pliocene of Panama)

Also known as NMB Loc. 17833

Where: Bocas, Panama (9.1° N, 81.6° W: paleocoordinates 9.0° N, 81.1° W)

• coordinate stated in text

• outcrop-level geographic resolution

When: Escudo de Veraguas Formation (Bocas del Toro Group), Piacenzian (3.6 - 2.6 Ma)

• STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), not specified to Early or Late Pliocene; direct date extrapolated from Cheetham et al. (1999); age range also provided by Bybell (1999).

• bed-level stratigraphic resolution

Environment/lithology: deep-water; poorly lithified, burrowed, bioturbated, fine-grained, argillaceous sandstone and poorly lithified, burrowed, bioturbated, argillaceous siltstone

• ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
• GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, pervasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.

Size class: macrofossils

Reposited in the BMNH, NMB, USNM

Collection methods: bulk, sieve,

• COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. COLLECTORS: unknown (PPP staff), 6/4/88. REPOSITORIES:collections presumably held at USNM, BMNH and Naturhistorisches Museum Basel (NMB),

Primary reference: L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]more details

Purpose of describing collection: paleoecologic analysis

PaleoDB collection 79988: authorized by Austin Hendy, entered by Kate Bulinski on 31.03.2008, edited by Austin Hendy

Creative Commons license: CC BY (attribution)

Taxonomic list

• NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
unclassified
  -
Scaphopoda
 Gadilida - Gadilidae
Cadulus sp. Philippi 1844 tusk shell
 Dentaliida - Dentaliidae
Dentalium sp. Linnaeus 1758 tusk shell
sensu lato
Gastropoda
 Heterostropha - Murchisonellidae
Anisocycla (Saccoina) sp. Dall and Bartsch 1904 snail
 Heterostropha - Pyramidellidae
Eulimella (Eulimella) sp. Forbes and MacAndrew 1846 snail
? Eulimella (Eulimella) sp. Forbes and MacAndrew 1846 snail
Eulimella (Ebalina) sp. Thiele 1929 snail
Besla sp. Dall and Bartsch 1904 snail
Pyramidellinae indet. Gray 1840 snail
 Heterostropha - Ringiculidae
 Vanikoroidea - Eulimidae
Eulimidae indet. Philippi 1853 snail
 Sorbeoconcha - Triphoridae
Triphoridae indet. Gray 1847 snail
 Sorbeoconcha - Cassidae
Sconsia sp. Gray 1847 snail
 Sorbeoconcha - Strombidae
Strombus sp. Linnaeus 1758 conch
 Neogastropoda - Marginellidae
Volvarina sp. Hinds 1844 margin shell
Dentimargo sp. Cossmann 1899 margin shell
 Neogastropoda - Muricidae
Haustellum sp. Schumacher 1817 murex snail
 Neogastropoda - Cystiscidae
? Persicula sp. Schumacher 1817 snail
 Neogastropoda - Olividae
Olivella (Macgintiella) sp. Olsson 1956 olive snail
 Neogastropoda - Nassariidae
Nassarius sp. Duméril 1806 snail
sensu lato
 Neogastropoda - Buccinidae
Solenosteira sp. Dall 1890 true whelk
Antillophos (Antillophos) sp. Woodring 1928 true whelk
 Neogastropoda - Columbellidae
"Costoanachis sp." = Anachis (Costoanachis), Mitrella sp.
"Costoanachis sp." = Anachis (Costoanachis) Sacco 1890 snail
Mitrella sp. Risso 1826 snail
 Neogastropoda - Drilliidae
Agladrillia (Agladrillia) sp. Woodring 1928 snail
 Neogastropoda - Borsoniidae
? Microdrillia sp. Casey 1903 snail
 Neogastropoda - Turridae
Polystira sp. Woodring 1928 turrid
Vaughanites sp. Woodring 1928 turrid
 Neogastropoda - Raphitomidae
Daphnella sp. Hinds 1844 snail
 Neogastropoda - Clathurellidae
Nannodiella sp. Dall 1919 snail
 Neogastropoda - Conidae
Conus sp. Linnaeus 1758 cone shell
 Sorbeoconcha - Naticidae
Natica sp. Scopoli 1777 moon snail
Polinices sp. Monfort 1810 moon snail
 Sorbeoconcha - Tornidae
Teinostoma sp. Adams and Adams 1853 snail
 Sorbeoconcha - Atlantidae
Atlanta sp. Lesueur 1817 snail
 Sorbeoconcha - Rissoidae
Rissoidae indet. Gray 1847 snail
Alvania sp. Risso 1826 snail
 Trochoidea - Trochidae
Microgaza sp. Dall 1881 top snail
 Opisthobranchia - Haminoeidae
"Atys (Aliculastrum) sp." = Aliculastrum
"Atys (Aliculastrum) sp." = Aliculastrum Pilsbry 1896 snail
 Opisthobranchia - Retusidae
Sulcoretusa sp. Burch 1945 snail
Volvulella (Volvulella) sp. Newton 1891 snail
 Opisthobranchia - Cylichnidae
Cylichnella sp. Gabb 1873 snail
Acteocina sp. Gray 1847 barrel-bubble
 Thecosomata - Limacinidae
Limacina sp. Bosc 1817 snail
? Limacina sp. Bosc 1817 snail
 Thecosomata - Cavoliniidae
Cavolinia sp. Avilgaard 1791 snail
Creseis sp. Rang 1828 snail
Styliola sp. Gray 1850 snail
Clio sp. Linnaeus 1767 snail
Bivalvia
 Nuculanida - Yoldiidae
Yoldia sp. Möller 1842 clam
 Nuculanida - Nuculanidae
Jupiteria sp. Bellardi 1875 pointed nut clam
Saccella sp. Woodring 1925 pointed nut clam
 Pectinida - Propeamussiidae
Cyclopecten sp. Verrill 1897 mud scallop
 Arcida - Arcidae
"? Hawaiarca sp." = Barbatia (Hawaiarca) Dall et al. 1938 clam
Anadara (Rasia) sp. Gray 1857 ark
 Lucinida - Lucinidae
Parvilucina sp. Dall 1901 clam
sensu lato
 Poromyida - Verticordiidae
Trigonulina sp. d'Orbigny 1946 clam
 Poromyida - Cuspidariidae
Halonympha sp. Dall and Smith 1886 clam
 Poromyida - Poromyidae
Poromya (Poromya) sp. Forbes 1844 clam
 Pholadida - Corbulidae
Caryocorbula sp. Gardner 1926 clam
 Cardiida - Tellinidae
Tellina (Phyllodina) sp. Dall 1900 tellin clam
Angulus sp. Megerle 1811 tellin clam
 Cardiida - Semelidae
Abra sp. Lamarck 1818 abra
 Carditida - Crassatellidae
Crassinella sp. Guppy 1874 clam
Anthozoa
 Scleractinia - Caryophylliidae
Asterosmilia profunda Duncan 1864 stony coral