PBDB Collection

Bore Lullingen 1, depth 21m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.5° N, 6.2° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Dingdener stage

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, fine-grained, silty sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1992

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 61895: authorized by Martin Aberhan, entered by Sabine Nürnberg on 07.07.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Foraminifera
Rotalioidea - Elphidiidae	Elphidium inflatum Elphidium inflatum
Nodosariacea - Glandulinidae	Glandulina ovula Glandulina ovula
Globothalamea
Rotaliida - Nonionidae	Nonion boueanum Nonion boueanum
Rotaliida - Cibicididae	Cibicides lobatulus, Cibicides pseudoungerianus, Cibicides dutemplei Cibicides lobatulus incl. 1 specimen C. lob. var. ornatus Cibicides pseudoungerianus Cushman 1922 Cibicides dutemplei dOrbigny 1846
Foraminifera
- Spiroplectamminidae	Spiroplectammina carinata Spiroplectammina carinata d'Orbigny 1846
Nodosariata
Vaginulinida - Vaginulinidae	Lenticulina (Lenticulina) calcar, Lenticulina (Planularia) grundensis, Lenticulina (Planularia) kubinyi, Lenticulina (Lenticulina) inornata, Lenticulina (Robulus) grandis, Lenticulina (Robulus) cultrata, Lenticulina (Robulus) depauperata Lenticulina (Lenticulina) calcar Lenticulina (Planularia) grundensis Lenticulina (Planularia) kubinyi Lenticulina (Lenticulina) inornata d'Orbigny 1846 Lenticulina (Robulus) grandis Lenticulina (Robulus) cultrata Lenticulina (Robulus) depauperata L. (L.) depauperata for. costata
Nodosariida - Nodosariidae	Lenticulina (Robulus) clypeiformis, Vaginulina obtusicosta, Dentalina intermittens, Dentalina bifurcata, Dentalina grossicosta, Dentalina pauperata, Dentalina konincki, Nodosaria bacillum, Nodosaria raphanus, Frondicularia annularis Lenticulina (Robulus) clypeiformis d'Orbigny 1846 Vaginulina obtusicosta Dentalina intermittens Dentalina bifurcata Dentalina grossicosta Dentalina pauperata d'Orbigny 1846 Dentalina konincki Reuss 1861 Nodosaria bacillum Nodosaria raphanus Brady 1884 Frondicularia annularis
Polymorphinida - Polymorphinidae	Globulina gibba, Pyrulina fusiformis, Guttulina problema, Pseudopolymorphina subnodosa Globulina gibba d'Orbigny 1832 Pyrulina fusiformis Guttulina problema d'Orbigny 1826 Pseudopolymorphina subnodosa P. subn. for. rugosa
Foraminifera
Lituolida - Lituolidae	Textularia gramen, Bigenerina nodosaria Textularia gramen dOrbigny 1846 Bigenerina nodosaria
Polychaeta
-	Polychaeta indet.¹ Polychaeta indet.¹ Grube 1850 Sedentarida indet.
Sabellida - Serpulidae	Josephella sp.¹, Protula cf. canavarii¹, Mercierella sp.¹ Josephella sp.¹ Caullery and Mesnil 1896 Protula cf. canavarii¹ Mercierella sp.¹ Fauvel 1923
Scaphopoda
Gadilida - Gadilidae	Gadila dingdenensis¹ Gadila dingdenensis¹ tusk shell
Dentaliida - Dentaliidae	Antalis dumasi¹, Lentigodentalium pseudomutabile¹ Antalis dumasi¹ tusk shell Antalis dumasi subsp. Lentigodentalium pseudomutabile¹ tusk shell
Cephalopoda
Spirulida - Spirulirostridae	Spirulirostra hoernesi¹ Spirulirostra hoernesi¹
Gastropoda
Cocculinoidea - Cocculinidae	Cocculina (Cocculina) miocaenica² Cocculina (Cocculina) miocaenica² snail
Epitonioidea - Epitoniidae	sensu lato Turriscala straeleni², Epitonium (Spiniscala) frondiculum², Acirsa (Hemiacirsa) aff. duvergieri², sensu lato Cirsotrema ertborni², Cirsotrema (Cirsotrema) crassicostatum² sensu lato Turriscala straeleni² wentletrap T. straeleni for. germanica Epitonium (Spiniscala) frondiculum² Wood 1842 wentletrap Acirsa (Hemiacirsa) aff. duvergieri² wentletrap sensu lato Cirsotrema ertborni² wentletrap Cirsotrema (Cirsotrema) crassicostatum² Deshayes 1839 wentletrap
Vanikoroidea - Eulimidae	Niso (Niso) terebellum², Balcis (Balcis) alba² Niso (Niso) terebellum² snail Balcis (Balcis) alba² snail incl. 5 specimens B. alba for. lactea
Neotaenioglossa - Calyptraeidae	Calyptraea (Calyptraea) chinensis² Calyptraea (Calyptraea) chinensis² Linnaeus 1758 slipper shell
Cerithioidea - Cerithiidae	Bittium (Bittium) tenuispina², "Bittium (Bittium) spina" = Bittium spina² Bittium (Bittium) tenuispina² cerith snail "Bittium (Bittium) spina" = Bittium spina² Partsch 1856 cerith snail
Cerithioidea - Turritellidae	Turritella subangulata² Turritella subangulata² turret shell
Pseudomelanoidea - Pseudomelaniidae	sensu lato Inquisitor borealis³ sensu lato Inquisitor borealis³ snail
Sorbeoconcha - Naticidae	Naticidae indet.², "Natica neglecta" = Cochlis neglecta², "Natica sensu lato tigrina" = Cochlis tigrina², Euspira staringi², Euspira helicina², Natica koeneni², Tectonatica miopusilla² Naticidae indet.² Guilding 1834 moon snail Naticidae juv. et indet. "Natica neglecta" = Cochlis neglecta² Mayer 1858 moon snail "Natica sensu lato tigrina" = Cochlis tigrina² Röding 1798 moon snail Euspira staringi² moon snail Euspira helicina² Brocchi 1814 moon snail E. aggr. helicina Natica koeneni² moon snail Tectonatica miopusilla² moon snail
Sorbeoconcha - Triforidae	Cerithiella (Cerithiella) genei² Cerithiella (Cerithiella) genei² snail
Sorbeoconcha - Triphoridae	Norephora (Norephora) fritschi² Norephora (Norephora) fritschi² snail
Sorbeoconcha - Cerithiopsidae	Cerithiopsis (Dizoniopsis) "sp. 3"² Cerithiopsis (Dizoniopsis) "sp. 3"² Hanley 1849 snail
Sorbeoconcha - Rissoidae	sensu lato Alvania holsatica², Alvania (Alvania) partschi², Alvania (Alvania) basisulcata² sensu lato Alvania holsatica² snail Alvania (Alvania) partschi² snail Alvania (Alvania) basisulcata² snail
Sorbeoconcha - Cassidae	"Semicassis (Echinophoria) bicoronata" = Echinophoria² "Semicassis (Echinophoria) bicoronata" = Echinophoria² snail
Sorbeoconcha - Ranellidae	"Charonia (Sassia) tarbelliana" = Sassia² "Charonia (Sassia) tarbelliana" = Sassia² triton shell
Sorbeoconcha - Triviidae	Erato (Erato) germanica² Erato (Erato) germanica² Schilder 1929 snail
Neogastropoda - Costellariidae	Orthosurcula steinvorthi³ Orthosurcula steinvorthi³ ribbed mitre
Neogastropoda - Ancillariidae	"Ancilla (Baryspira) obsoleta" = Amalda obsoleta² "Ancilla (Baryspira) obsoleta" = Amalda obsoleta² Brocchi 1814 snail
Neogastropoda - Olividae	Oliva (Strephona) dufresnei² Oliva (Strephona) dufresnei² Basterot 1825 olive snail
Neogastropoda - Cancellariidae	Babylonella fusiformis², "Trigonostoma (Ventrilia) pouwi" = Ventrilia², Cancellaria (Merica) contorta² Babylonella fusiformis² snail "Trigonostoma (Ventrilia) pouwi" = Ventrilia² snail Cancellaria (Merica) contorta² snail C. contorta gelriana
Neogastropoda - Mitridae	Mitra (Tiara) substriatula³ Mitra (Tiara) substriatula³ d'Orbigny 1852 mitre shell
Neogastropoda - Fasciolariidae	Streptodictyon sexcostatus², ? Exilifusus attenuatus² Streptodictyon sexcostatus² snail ? Exilifusus attenuatus² snail
Neogastropoda - Nassariidae	Nassarius bocholtensis², Nassarius tenuistriatus² Nassarius bocholtensis² snail Nassarius tenuistriatus² snail
Neogastropoda - Columbellidae	Mitrella (Macrurella) nassoides² Mitrella (Macrurella) nassoides² snail
Neogastropoda - Volutidae	Scaphella (Scaphella) bolli² Scaphella (Scaphella) bolli² volute
Neogastropoda - Muricidae	"Murex (Haustellum) inornatus" = Haustellum², Lyrotyphis (Eotyphis) fistulosus², Typhis (Typhis) ? pungens² "Murex (Haustellum) inornatus" = Haustellum² murex snail Lyrotyphis (Eotyphis) fistulosus² murex snail Typhis (Typhis) ? pungens² Solander in Brander and Brander 1766 murex snail
Neogastropoda - Terebridae	Strioterebrum hoernesi³, Strioterebrum ronaldi³, Terebra (Myurellina) acuminata³ Strioterebrum hoernesi³ auger snail Strioterebrum ronaldi³ auger snail Terebra (Myurellina) acuminata³ auger snail T. acuminata neglecta
Neogastropoda - Conidae	"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ "Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ cone shell
Neogastropoda - Mangeliidae	sensu lato Mangelia andersoni³, Sorgenfreispira plicatelloides³, Sorgenfreispira "sp. 1"³, Sorgenfreispira tenella³ sensu lato Mangelia andersoni³ Wienrich and Janssen 2007 snail Sorgenfreispira plicatelloides³ snail Sorgenfreispira "sp. 1"³ Moroni 1979 snail Sorgenfreispira tenella³ Conrad 1848 snail
Neogastropoda - Raphitomidae	Teretia anceps³, Mioawateria andersoni³, Metuonella grippi³ Teretia anceps³ snail Mioawateria andersoni³ snail Metuonella grippi³ snail
Neogastropoda - Turridae	Boreodrillia hosiusi³, Nitidiclavus maitrejus³, sensu lato Unedogemmula boreoturricula³, sensu lato Unedogemmula hanseata³, sensu lato Unedogemmula stoffelsii³, Gemmula (Gemmula) sp.³, Gemmula (Gemmula) vermicularis³, Gemmula (Gemmula) zimmermanni³, Fusiturris aquensis³, Fusiturris duchastelii³ Boreodrillia hosiusi³ turrid Nitidiclavus maitrejus³ turrid sensu lato Unedogemmula boreoturricula³ turrid sensu lato Unedogemmula hanseata³ turrid sensu lato Unedogemmula stoffelsii³ turrid Gemmula (Gemmula) sp.³ Weinkauff 1875 turrid Gemmula (Gemmula) vermicularis³ turrid Gemmula (Gemmula) zimmermanni³ turrid Fusiturris aquensis³ turrid Fusiturris duchastelii³ turrid F. duchastelii flexiplicata
Neogastropoda - Drilliidae	Splendrillia selenkae³ Splendrillia selenkae³ snail
Neogastropoda - Clathurellidae	Pleurotomoides luisae³ Pleurotomoides luisae³ snail
Neogastropoda - Borsoniidae	Asthenotoma pannoides³, Bathytoma jugleri³ Asthenotoma pannoides³ snail Bathytoma jugleri³ snail
Sorbeoconcha - Aporrhaidae	Aporrhais (Aporrhais) alata² Aporrhais (Aporrhais) alata² snail
Sorbeoconcha - Xenophoridae	Xenophora deshayesi² Xenophora deshayesi² Michelotti 1847 carrier shell
Sorbeoconcha - Vitrinellidae	Circulus praecedens², Circulus hennei² Circulus praecedens² snail 2 specimens C. praecedens for. praecedens and 1 specimen C. praecedens for. gliberti Circulus hennei² Glibert 1952 snail
Entomotaeniata - Cimidae	Actaeopyramis elatus³ Actaeopyramis elatus³ snail
Heterostropha - Pyramidellidae	Chrysallida (Pyrgulina) pygmaea³, Chrysallida (Parthenina) "sp. 1"³, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³, Turbonilla (Turbonilla) gastaldi³, Turbonilla (Turbonilla) koeneniana³, Kleinella (Kleinella) sp.³, Pyramidella (Voluspa) elata³, Pyramidella (Voluspa) plicosa³, Eulimella (Eulimella) scillae³, Odostomia (Megastomia) cf. conoidea³, Odostomia (Megastomia) cf. pallidaeformis³, Odostomia (Megastomia) tuexeni³, Syrnola hoernesi³, Syrnola neumayri³ Chrysallida (Pyrgulina) pygmaea³ Grateloup 1838 snail Ch. pygmaea f. belgica Chrysallida (Parthenina) "sp. 1"³ Bucquoy et al. 1883 snail "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³ snail sensu JANSSEN "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³ snail sensu JANSSEN Turbonilla (Turbonilla) gastaldi³ snail sensu JANSSEN Turbonilla (Turbonilla) koeneniana³ snail sensu JANSSEN Kleinella (Kleinella) sp.³ Adams 1860 snail Pyramidella (Voluspa) elata³ snail Pyramidella (Voluspa) plicosa³ snail Eulimella (Eulimella) scillae³ snail Odostomia (Megastomia) cf. conoidea³ Brocchi 1814 snail Odostomia (Megastomia) cf. pallidaeformis³ snail Odostomia (Megastomia) tuexeni³ snail Syrnola hoernesi³ snail Syrnola neumayri³ snail
Heterostropha - Acteonidae	Acteon (Acteon) semistriatus³, Acteon (Acteon) "sp. 2"³ Acteon (Acteon) semistriatus³ snail spelled as 'Actaeon' Acteon (Acteon) "sp. 2"³ Montfort 1810 snail spelled as 'Actaeon'
Heterostropha - Ringiculidae	Ringicula (Ringiculina) buccinea³ Ringicula (Ringiculina) buccinea³ Sowerby 1823 snail
Trochoidea - Trochidae	Calliostoma (Ampullotrochus) ? laureatum², Calliostoma (Ampullotrochus) serratocostatum² Calliostoma (Ampullotrochus) ? laureatum² top snail Calliostoma (Ampullotrochus) serratocostatum² top snail C. serratocostatum mülleri
Lepetellida - Lepetellidae	Lepetella compressiuscula² Lepetella compressiuscula² snail
Opisthobranchia - Retusidae	Retusa (Cylichnina) elongata³, Retusa (Cylichnina) subangistoma³, Volvulella (Volvulella) acuminata³ Retusa (Cylichnina) elongata³ snail Retusa (Cylichnina) subangistoma³ snail Volvulella (Volvulella) acuminata³ snail
Opisthobranchia - Cylichnidae	Bullichna paucistriata³, Cylichna (Cylichna) pseudoconvoluta³, Scaphander grateloupi³ Bullichna paucistriata³ snail Cylichna (Cylichna) pseudoconvoluta³ snail Scaphander grateloupi³ snail
Opisthobranchia -	Roxania utriculus³ Roxania utriculus³ snail
Bivalvia
Nuculanida - Nuculanidae	"Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ "Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ Nyst and Westendorp 1839 pointed nut clam
Nuculanida - Yoldiidae	Yoldia (Yoldia) glaberrima¹, "Portlandia (Yoldiella) pygmaea" = Yoldiella¹ Yoldia (Yoldia) glaberrima¹ von Münster 1837 clam "Portlandia (Yoldiella) pygmaea" = Yoldiella¹ clam
Nuculida - Nuculidae	Nuculoma haesendoncki¹ Nuculoma haesendoncki¹ nut clam N. haesendoncki hanseata
Poromyida - Verticordiidae	Verticordia (Verticordia) punctata¹ Verticordia (Verticordia) punctata¹ clam
Hiatellida - Hiatellidae	Panopea (Panopea) meynardi¹ Panopea (Panopea) meynardi¹ clam
Solenida - Pharidae	Ensis hausmanni¹ Ensis hausmanni¹ clam
Pholadida - Teredinidae	Teredinidae indet.¹ Teredinidae indet.¹ Rafinesque 1815 clam
Pholadida - Corbulidae	Corbula (Varicorbula) gibba¹ Corbula (Varicorbula) gibba¹ Oliva 1792 clam C. (V.) gibba gibba
Cardiida - Glossidae	Glossus (Glossus) lunulatus¹ Glossus (Glossus) lunulatus¹ clam
Cardiida - Kelliellidae	Spaniodontella nitida¹ Spaniodontella nitida¹ clam
Cardiida - Veneridae	Venus (Ventricoloidea) multilamella¹, "Gouldia minima" = Gouldia (Gouldia) minima¹ Venus (Ventricoloidea) multilamella¹ Lamarck 1818 venus clam V. (V.) multilamella multilamella "Gouldia minima" = Gouldia (Gouldia) minima¹ Montagu 1803 venus clam
Cardiida - Mactridae	Lutraria frangula¹ Lutraria frangula¹ clam
Cardiida - Erycinidae	"Hemilepton mionitidum" = Erycina (Hemilepton)¹ "Hemilepton mionitidum" = Erycina (Hemilepton)¹ clam
Cardiida - Solecurtidae	Solecurtus basteroti¹ Solecurtus basteroti¹ clam
Cardiida - Tellinidae	Angulus (Moerella) aff. donacillus¹ Angulus (Moerella) aff. donacillus¹ tellin clam
Cardiida - Semelidae	sensu lato Abra lehmanni¹, Abra (? Abra) sorgenfreii¹ sensu lato Abra lehmanni¹ abra Abra (? Abra) sorgenfreii¹ abra
Cardiida - Cardiidae	"Habecardium subturgidum" = Laevicardium (Habecardium)¹, Parvicardium straeleni¹ "Habecardium subturgidum" = Laevicardium (Habecardium)¹ cockle H. subturgidum subturgidum Parvicardium straeleni¹ cockle
Lucinida - Lucinidae	"Lucinoma borealis" = Lucinoma (Lucinoma) borealis¹, "Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti¹ "Lucinoma borealis" = Lucinoma (Lucinoma) borealis¹ Linnaeus 1767 clam "Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti¹ Nyst 1861 clam C. droueti droueti
Carditida - Carditidae	Cyclocardia sp.¹ Cyclocardia sp.¹ Conrad 1867 clam
Carditida - Astartidae	Astarte (Astarte) radiata¹ Astarte (Astarte) radiata¹ Nyst and Westendorp 1839 clam Astarte radiata fa. radiata 13 specimens, Astarte radiata fa. convexior 52 specimens
Pectinida - Pectinoidae	Korobkovia woodi¹, Pseudamussium lilli¹, Aequipecten angelonii¹ Korobkovia woodi¹ scallop Pseudamussium lilli¹ Pusch 1837 scallop Aequipecten angelonii¹ scallop
Ostreida - Pinnidae	Atrina pectinata¹ Atrina pectinata¹ Linnaeus 1758 oyster
Arcida - Arcidae	Scapharca (Scapharca) diluvii¹ Scapharca (Scapharca) diluvii¹ Lamarck 1805 clam
Arcida - Limopsidae	Limopsis (Limopsis) aurita¹, "Limopsis (Pectunculina) anomala" = Limopsis¹, Limopsis (Pectunculina) lamellata¹ Limopsis (Limopsis) aurita¹ Brocchi 1814 clam "Limopsis (Pectunculina) anomala" = Limopsis¹ clam Limopsis (Pectunculina) lamellata¹ Lehmann 1885 clam
Gymnolaemata
Cheilostomata - Lunulitidae	Lunulites voigti¹, Lunulites burdigalensis¹ Lunulites voigti¹ Lunulites burdigalensis¹
Cheilostomata - Membraniporidae	Reussirella haidingeri¹ Reussirella haidingeri¹
Anthozoa
Scleractinia - Micrabaciidae	Stephanophyllia imperialis Stephanophyllia imperialis Michelin 1841 stony coral
Scleractinia - Flabellidae	Flabellum sp., Flabellum pompeckji, Flabellum tuberculatum Flabellum sp. Lesson 1831 stony coral Flabellum pompeckji Krejci 1926 stony coral Flabellum tuberculatum Keferstein 1859 stony coral

¹Wienrich 1999; ²Wienrich 2001; ³Wienrich and Janssen 2007