Krinke pit, Werder [Werder Fm] (Miocene of Germany)

Where: Germany (53.0° N, 9.1° E: paleocoordinates 53.1° N, 7.2° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Werder Formation, Burdigalian (20.4 - 16.0 Ma)

• STRATIGRAPHIC RELATIONS: From the Werder Fm, which regionally overlie the Arnum (Burdigalian) and is overlain by the Winerswijk-Miste Fm (Langhian). AGE: Burdigalian, on the basis of macrofossil biostratigraphy. STRATIGRAPHIC POSITION: Collection is a composite for the formation at this section.

• formation-level stratigraphic resolution

Environment/lithology: coastal; unlithified gravel

• ENVIRONMENT: Marine, siliciclastic, with some reduced salinity species. Note estuarine though.
• SPECIFIC LITHOLOGY: Gravel. LITHIFICATION: Unlithified, on the basis of figured specimens.

Size classes: macrofossils, mesofossils

Collection methods: quarrying,

• COLLECTORS: Not stated in text, but presumably the quarry operators, and authors. REPOSITORY: Not stated in English portion of text.

Primary reference: H. Moths, F. Albrecht, and G. Stein. 2010. Die molluskenfauna (Hermmorium, Untermiozän) aus der Kiesgrube kirnke bei Werder (Nordwest-Niedersachsen). Palaeofocus 3:1-155 [A. Hendy/A. Hendy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111599: authorized by Austin Hendy, entered by Austin Hendy on 26.06.2011, edited by Mark Uhen

Creative Commons license: CC BY (attribution)

Taxonomic list

• COVERAGE: Exhaustive for molluscs. NOMENCLATURE: Authoritative publication, with relatively modern nomenclature, and species-resolution identifications.
unclassified
  -
Bivalvia
 Mytilida - Crenellidae
Crenella sp. Brown 1827 clam
 Mytilida - Mytilidae
Modiolula phaseolina Philippi 1844 mussel
 Ostreida - Pteriidae
Pteriidae indet. Gray 1847 pearl oyster
 Ostreida - Pinnidae
Pinnidae indet. Leach 1819 oyster
 Ostreida - Ostreidae
Cubitostrea digitalina Eichwald 1830 oyster
 Pectinida - Limidae
Limatula sp. Wood 1839 file clam
Limea (Limea) sp. Bronn 1831 file clam
 Pectinida - Anomiidae
Anomiinae indet. Rafinesque 1815 scallop
 Pectinida - Pectinoidae
Patinopecten brummeli Nyst 1864 scallop
Palliolinae indet. Korbobkov in Eberzin 1960 scallop
 Arcida - Glycymerididae
"Glycymeris (Chevronia) obovata" = Glycymeris (Glycymeris) obovata Lamarck 1819 clam
SUBSPECIES: Glycymeris (Chevronia) obovata baldii
 Arcida - Arcidae
Arca sp. Linnaeus 1758 ark
Barbatia sp. Gray 1842 clam
 Arcida - Limopsidae
 Carditida - Condylocardiidae
 Carditida - Carditidae
Cardita sp. Bruguière 1792 clam
 Carditida - Astartidae
Astarte (Astarte) radiata Nyst and Westendorp 1839 clam
Goodallia (Goodallia) waeli Glibert 1945 clam
Digitaria beyschlagi Kautsky 1925 clam
Digitaria burdigalensis Cossmann and Peyrot 1912 clam
 Poromyida - Cuspidariidae
 Thraciida - Thraciidae
? Thraciidae indet. Stoliczka 1870 clam
 Thraciida - Periplomatidae
Periploma ariei Gürs 1996 clam
 Solenida - Pharidae
 Hiatellida - Hiatellidae
Panopea (Panopea) menardi Deshayes 1829 clam
"Hiatella (Hiatella) arctica" = Hiatella arctica Linnaeus 1767 clam
Cyrtodaria angusta Nyst and Westendorp 1839 clam
 Pholadida - Corbulidae
Lentidium (Ajanssenium) turonense Cossmann and Peyrot 1914 clam
Caryocorbula basteroti Hörnes 1870 clam
 Pholadida - Myidae
Myidae indet. Lamarck 1809 softshell clam
Sphenia pusilla Sorgenfrel 1958 softshell clam
 Pholadida - Teredinidae
Teredininae indet. Rafinesque 1815 clam
 Pholadida - Pholadidae
Pholadinae "indet. 1" Lamarck 1809 clam
Pholadinae "indet. 2" Lamarck 1809 clam
 Cardiida - Donacidae
 Cardiida - Semelidae
 Cardiida - Solecurtidae
 Cardiida - Tellinidae
Angulus (Peronaea) benedeni Nyst and Westendorp 1839 tellin clam
SUBSPECIES: Angulus (Peronaea) benedeni fallax
 Cardiida - Cardiidae
Acanthocardia hanseata Kautsky 1925 cockle
Bucardium sp. Gray 1853 cockle
Parvicardium sp. Monterosato 1884 cockle
 Cardiida - Veneridae
"Callista (Costacallista) xesta" = Costacallista venus clam
Gouldia (Gouldia) minima Montagu 1803 venus clam
Timoclea (Timoclea) patera Soregnfrei 1958 venus clam
Irus irus Linnaeus 1767 venus clam
Venus (Ventricoloidea) multilamella Lamarck 1818 venus clam
 Cardiida - Ungulinidae
 Cardiida - Kelliellidae
 Cardiida - Glossidae
 Cardiida - Chamidae
Chama gryphoides Linnaeus 1758 jewel box
 Cardiida - Mactridae
Spisula ? obtruncata Wood 1850 clam
 Cardiida - Trapezidae
 Cardiida -
 Cardiida - Montacutidae
 Cardiida - Erycinidae
Erycina striatissima Cerulli-Irelli 1908 clam
 Cardiida - Lasaeidae
Scacchia (Scacchia) degrangei Cossmann and Peyrot 1911 clam
 Cardiida - Leptonidae
 Cardiida - Galeommatidae
 Lucinida - Lucinidae
Lucinoma (Lucinoma) borealis Linnaeus 1767 clam
SUBSPECIES: Lucinoma (Lucinoma) borealis praecedens
"Loripes (Microloripes) dentatus" = Lucina (Microloripes) clam
"Ctena (Ctena) decussata" = Codakia (Ctena) decussata Costa 1829 clam
 Nuculida - Nuculidae
Nucula (Nucula) cromata Sorgenfrel 1958 nut clam
Leionucula sp. Quenstedt 1930 nut clam
 Nuculanida - Nuculanidae
"Lembulus emarginatus" = Leda (Lembulus)
"Lembulus emarginatus" = Leda (Lembulus) pointed nut clam
 Nuculanida - Yoldiidae
Yoldia (Yoldia) glaberrima von Münster 1837 clam
Yoldiella philippiana clam
SUBSPECIES: Yoldiella philippiana wesselinghi
Gastropoda
 Heterostropha - Ringiculidae
 Heterostropha - Acteonidae
Acteon (Acteon) arnumensis Sorgenfrei 1958 snail
Acteon (Acteon) "sp. 1" Montfort 1810 snail
Acteon (Acteon) "sp. 2" Montfort 1810 snail
Acteon (Acteon) "sp. 3" Montfort 1810 snail
 Heterostropha - Pyramidellidae
Pyramidella (Volupsa) sp. Lamarck 1799 snail
Chrysallida sp. Carpenter 1856 snail
Odostomia (Brachystomia) sp. Fleming 1817 snail
Odostomia (Odostomia) sp. Fleming 1817 snail
"Turbonilla (Pyrgolampros) "sp. 1"" = Pyrgolampros Sacco 1892 snail
"Turbonilla (Pyrgolampros) "sp. 2"" = Pyrgolampros Sacco 1892 snail
Evalea sp. Adams 1860 snail
Kleinella (Kleinella) sp. Adams 1860 snail
"Turbonilla (Strioturbonilla) borealis" = Strioturbonilla snail
"Turbonilla (Strioturbonilla) "sp. 1"" = Strioturbonilla Sacco 1892 snail
"Turbonilla (Strioturbonilla) "sp. 2"" = Strioturbonilla Sacco 1892 snail
Syrnola "sp. 1" Adams 1860 snail
Syrnola "sp. 2" Adams 1860 snail
 Heterostropha - Tjaernoiidae
 Heterostropha - Architectonicidae
 Heterostropha - Mathildidae
  -
Clathromangelia sp. Monterosato 1884 snail
 Entomotaeniata - Cimidae
 Opisthobranchia - Umbraculidae
Spiricella unguiculus Rang and Des Moulins 1828 snail
 Thecosomata - Cavoliniidae
 Thecosomata - Limacinidae
 Opisthobranchia - Cylichnidae
 Opisthobranchia - Retusidae
"Cylichnina elongata" = Retusa (Cylichnina) snail
"Cylichnina subangistoma" = Retusa (Cylichnina) snail
 Opisthobranchia - Philinidae
"Philine (Megistostoma) complanata" = Megistostoma
"Philine (Megistostoma) complanata" = Megistostoma snail
 Opisthobranchia -
 Cocculinoidea - Cocculinidae
 Epitonioidea - Epitoniidae
Cirsotrema (Cirsotrema) sallomacense wentletrap
SUBSPECIES: Cirsotrema (Cirsotrema) sallomacense nordica
Cirsotrema (Discoscala) sp. Mörch 1852 wentletrap
"Epitonium (Spiniscala) sp." = Scala (Spiniscala) De Boury 1909 wentletrap
Epitonium (Hyaloscala) sp. De Boury 1890 wentletrap
"Nodiscala scacchii" = Opalia (Nodiscala) wentletrap
"Amaea (Scalina) subreticulata" = Scalina wentletrap
 Neotaenioglossa - Calyptraeidae
Calyptraea (Calyptraea) chinensis Linnaeus 1758 slipper shell
Crepidula (Janacus) crepidula Linnaeus 1758 slipper shell
 Sorbeoconcha - Littorinidae
"Littorina (Melarhaphe) sp." = Melarhaphe
"Littorina (Melarhaphe) sp." = Melarhaphe Menke 1828 periwinkle
 Sorbeoconcha - Vermetidae
 Sorbeoconcha - Rissoidae
Rissoina (Zebinella) sp. Mörch 1876 snail
"Alvania (Taramellia) partschi" = Manzonia (Alvinia) snail
"Alvania (Taramellia) sp." = Manzonia (Alvinia) Monterosato 1884 snail
"Zebina (Stossichia) multicingulata" = Stossichia multicingulata Boettger 1887 snail
Alvania (Alvania) sp. Risso 1826 snail
 Sorbeoconcha - Naticidae
"Sigaretotrema striatella" = Eunaticina moon snail
Euspira helicina Brocchi 1814 moon snail
SUBSPECIES: Euspira helicina protracta
Neverita josephinia Risso 1826 moon snail
SUBSPECIES: Neverita josephinia olla
Cochlis hanseata moon snail
Cochlis sp. Röding 1798 moon snail
Cochlis neglecta Mayer 1858 moon snail
Tanea dillwyni Payraudeau 1826 moon snail
SUBSPECIES: Tanea dillwyni koeneni
 Sorbeoconcha - Elachisinidae
 Sorbeoconcha - Iravadiidae
Ceratia proxima Forbes and Hanley 1850 snail
 Sorbeoconcha - Caecidae
Caecum sp. Fleming 1813 snail
 Sorbeoconcha - Vitrinellidae
 Sorbeoconcha - Tornidae
 Sorbeoconcha - Eocypraeidae
 Sorbeoconcha - Ovulidae
Simnia sp. Risso 1826 snail
 Sorbeoconcha - Cypraeidae
Cypraeinae indet. Rafinesque 1815 cowry
 Sorbeoconcha - Triviidae
Erato sp. Risso 1826 snail
 Sorbeoconcha - Ranellidae
Sassia tarbelliana triton shell
Sassia sp. Bellardi 1872 triton shell
 Sorbeoconcha - Cassidae
 Sorbeoconcha - Personidae
Distorsio cancellina Lamarck 1803 snail
 Neogastropoda - Costellariidae
"Vexillum (Uromitra) sp." = Uromitra, Orthosurcula steinvorthi
"Vexillum (Uromitra) sp." = Uromitra Bellardi 1887 ribbed mitre
 Neogastropoda - Olividae
Oliva (Strephona) dufresnei Basterot 1825 olive snail
 Neogastropoda - Ancillariidae
Amalda obsoleta Brocchi 1814 snail
 Neogastropoda - Fasciolariidae
"Streptolathyrus rothi" = Streptochetus (Streptolathyrus) snail
 Neogastropoda - Columbellidae
 Neogastropoda - Melongenidae
"Melongena cornuta" = Galeodes cornutus
"Melongena cornuta" = Galeodes cornutus Agassiz 1843 snail
 Neogastropoda - Nassariidae
Nassarius "sp. 1" Duméril 1806 snail
Nassarius "sp. 2" Duméril 1806 snail
Nassarius "sp. 3" Duméril 1806 snail
Nassarius coronatus Bruguière 1789 snail
 Neogastropoda - Buccinidae
Euthria sp. Gray 1850 true whelk
"Kelletia (Borekelletia) hosiusi" = Siphonalia (Kelletia) true whelk
 Neogastropoda - Drilliidae
 Neogastropoda - Turridae
Gemmula (Gemmula) denticula turrid
SUBSPECIES: Gemmula (Gemmula) denticula borealis
Gemmula (Gemmula) sp. Weinkauff 1875 turrid
Acamptogenotia sp. Rovereto 1899 turrid
Fusiturris duchastelli turrid
SUBSPECIES: Fusiturris duchastelli flexiplicata
 Neogastropoda - Clavatulidae
 Neogastropoda - Clathurellidae
 Neogastropoda - Terebridae
 Neogastropoda - Conidae
"Conus (Chelyconus) aquensis" = Conus (Chelyoconus), "Conus (Conolithus) sp." = Conus (Conolithes)
"Conus (Chelyconus) aquensis" = Conus (Chelyoconus) cone shell
"Conus (Conolithus) sp." = Conus (Conolithes) Herrmannsen 1847 cone shell
 Neogastropoda - Mangeliidae
? Mangelia guerichi Kautsky 1925 snail
? Mangelia miorugulosa Kautsky 1925 snail
Sorgenfreispira "sp. 1" Moroni 1979 snail
Sorgenfreispira tenella Conrad 1848 snail
Sorgenfreispira calais Kautsky 1925 snail
? Bela sp. Gray 1847 snail
Benthomangelia sp. Thiele 1925 snail
Benthomangelia grippi Anderson 1964 snail
 Neogastropoda - Raphitomidae
 Neogastropoda - Borsoniidae
"Genota ramosa" = Genota valeriae Hoernes and Auinger 1891 snail
 Neogastropoda - Cancellariidae
Calcarata calcarata Brochhi 1814 snail
"Trigonostoma (Ventrilia) geslini" = Ventrilia snail
SUBSPECIES: Trigonostoma (Ventrilia) geslini josephinae
"Trigonostoma (Ventrilia) pouwi" = Ventrilia snail
Cancellaria (Bivetiella) cancellata Linnaeus 1767 snail
SUBSPECIES: Cancellaria (Bivetiella) cancellata praecedens
Sveltia sp. Jousseaume 1887 snail
Contortia sp. Sacco 1894 snail
 Neogastropoda -
 Neogastropoda - Mitridae
Mitrinae indet. Swainson 1831 mitre shell
Mitra (Tiara) substriatula d'Orbigny 1852 mitre shell
 Neogastropoda - Volutidae
 Neogastropoda - Muricidae
Chicoreus sp. Monfort 1810 murex snail
Chicoreus (Chicoreus) aquitanicus Grateloup 1833 murex snail
Murex (Haustellum) beyrichi Koenen 1889 murex snail
Pterynotus sp. Swainson 1833 mustached bat
? Ocinebrina sp. Jousseaume 1880 murex snail
Typhis (Typhis) pungens Solander in Brander and Brander 1766 murex snail
 Neogastropoda - Harpidae
"Morum (Oniscidia) sp." = Oniscidia
"Morum (Oniscidia) sp." = Oniscidia Mörch 1852 snail
 Sorbeoconcha - Xenophoridae
Xenophora sp. Fischer von Waldheim 1807 carrier shell
 Sorbeoconcha - Aporrhaidae
Aporrhais dingdenensis Marquet et al. 2002 snail
 Sorbeoconcha - Ficidae
Ficus condita Brongniart 1823 fig shell
 Sorbeoconcha - Cerithiopsidae
? Cerithiopsidae indet. Adams and Adams 1853 snail
Cerithiopsis sp. Hanley 1849 snail
? Seila sp. Adams 1861 snail
 Sorbeoconcha - Newtoniellidae
 Sorbeoconcha - Triphoridae
 Sorbeoconcha - Campanilidae
"Cerithium (Thericium) sp." = Thericium
"Cerithium (Thericium) sp." = Thericium Monterosato 1809 snail
 Pseudomelanoidea - Pseudomelaniidae
 Vanikoroidea - Eulimidae
Niso sp. Risso 1826 snail
Eulima (Eulima) sp. Risso 1826 snail
 Cerithioidea - Batillariidae
Granulolabium plicatum Bruguiére 1792 snail
 Cerithioidea - Potamididae
Terebralia bidentata Defrance 1832 horn snail
 Cerithioidea - Scaliolidae
Sandbergeria wolffi Kautsky 1925 snail
 Cerithioidea - Cerithiidae
Bittium tenuispina cerith snail
Bittium spina Partsch 1856 cerith snail
 Cerithioidea - Turritellidae
Turritellinae indet. Lovén 1847 turret shell
Haustator eryna d'Orbigny 1852 turret shell
 Lepetellida - Fissurellidae
Emarginula fissura Linnaeus 1758 snail
Diodora subcostaria d'Orbigny 1852 snail
 Trochoidea - Solariellidae
 Trochoidea - Trochidae
Calliostoma (Ampullotrochus) sp. Monterosato 1890 top snail
 Trochoidea - Skeneidae
? Solariorbis sp. Conrad 1865 snail
Cephalopoda
 Nautilida - Aturiidae
Aturia sp. Bronn 1838 nautiloid
Polyplacophora
 Neoloricata - Leptochitonidae
? Lepidopleurus sp. Gray 1847 chiton
Scaphopoda
 Dentaliida - Dentaliidae
Fissidentalium sp. Fischer 1885 tusk shell
Antalis sp. Adams and Adams 1854 tusk shell
 Gadilida - Gadilidae
Gadila benoisti tusk shell
SUBSPECIES: Gadila benoisti houtalenensis
Chondrichthyes
 Lamniformes - Cetorhinidae
Keasius rhenanus n. sp. Reinecke and Radwanski 2015 basking shark