Oeningen (Zurich collection) (Miocene of Germany)

Where: Baden-Württemberg, Germany (47.7° N, 8.9° E: paleocoordinates 47.9° N, 7.8° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: MN 7 mammal zone, Upper Oehningen beds Member (Upper Freshwater-Molasse Formation), Sarmatian (12.7 - 11.6 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: crater lake; lithified limestone

• "meromictic maar lake of approximately 1000 m in diameter and perhaps up to 100 m in depth" (Lutz, 1997). "Deposited in the Northern Alpine Foreland basin" (Uhl et al., 2006).

Size classes: macrofossils, mesofossils

Preservation: mold/impression, adpression

Collection methods: Collection of Zurich University

Primary reference: O. Heer. 1849. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Zweiter Theil: Henschrecken, Florfliegen, Aderflügler, Schmetterlinge und Fliegen 1-264 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 117252: authorized by Matthew Clapham, entered by Matthew Clapham on 18.09.2011, edited by Jered Karr and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

Arachnida
 Araneae -
Schellenbergia rotundata n. gen. n. sp.10, Clubiona eseri n. sp.10
Schellenbergia rotundata n. gen. n. sp.10 Heer 1864 spider
Clubiona eseri n. sp.10 Heer 1864 spider
 Araneae - Thomisidae
Thomisus lividus n. sp.10, Thomisus sulzeri n. sp.10, "Thomisus oeningensis n. sp." = Xysticus oeningensis10
Thomisus lividus n. sp.10 Heer 1864 crab spider
Thomisus sulzeri n. sp.10 Heer 1864 crab spider
"Thomisus oeningensis n. sp." = Xysticus oeningensis10 Heer 1864 crab spider
 Araneae - Dictynidae
? Argyroneta longipes n. sp.10 Heer 1864 spider
 Araneae - Araneidae
"Epeira molassica n. sp." = Araneus molassicus10
"Epeira molassica n. sp." = Araneus molassicus10 Heer 1864 orb-weaving spider
 Araneae - Theridiidae
Theridion maculipes n. sp.10 Heer 1864 cobweb spider
Theridion globulus n. sp.10 Heer 1864 cobweb spider
Theridion annulipes n. sp.10 Heer 1864 cobweb spider
 Araneae - Gnaphosidae
"Macaria tenella n. sp." = Micaria tenella10
"Macaria tenella n. sp." = Micaria tenella10 Heer 1864 spider
Malacostraca
 Isopoda - Armadillidiidae
"Armadillo molassicus n. sp." = Armadillidium molassicum10
"Armadillo molassicus n. sp." = Armadillidium molassicum10 Heer 1864 isopod
Insecta
 Diptera - Chironomidae
Chironomus gaudini n. sp.10 Heer 1864 non biting midge
 Diptera - Oestridae
"Dipterites obovatus n. sp." = Adipterites obovatus10
"Dipterites obovatus n. sp." = Adipterites obovatus10 Heer 1864 bot fly
 Diptera - Syrphidae
Syrphus schellenbergi n. sp.10 Heer 1864 hover fly
Syrphus bremii n. sp.10 Heer 1864 hover fly
 Diptera - Tabanidae
"? Hexatoma oeningensis n. sp." = Heptatoma oeningensis10
"? Hexatoma oeningensis n. sp." = Heptatoma oeningensis10 Heer 1864 horse fly
 Diptera - Bibionidae
Bibio elongatus16 Heer 1849 march fly
ETHZ S.N. 189 Pl. I 863 (labeled by Heer)
Dilophus pumilio n. sp.16 Skartveit and Pika 2014 march fly
ETHZ Pl. I 886
Bibio brevis16 Heer 1849 march fly
Some in lot Pl. I 856; ETHZ Pl.I 865, Pl. I 887
Bibio fusiformis16 Heer 1849 march fly
ETHZ Pl. I 858 (x2), 861 (x6)
Bibio pulchellus16 Heer 1849 march fly
ETHZ Pl. I 854
Bibio moestus16 Heer 1849 march fly
ETHZ Pl. I 855, 857(x2)
Bibio angustatus16 Heer 1849 march fly
ETHZ Pl. I 851
Bibio crassinervis n. sp.16 Skartveit and Pika 2014 march fly
ETHZ Pl. I 851, Pl.I 851(2), Pl. I 862, Pl. I 875, Pl. I 888
"Protomyia speciosa n. sp." = Penthetria amoena10 Heer 1849 march fly
ETHZ Pl. I 873; also Pl. I 872, 876 as P. amoena in Skartveit & Pika, 2014
Penthetria brevicollis16 Heer 1849 march fly
ETHZ Pl. I 870
Plecia hilaris16 Heer 1849 love bug
ETHZ Pl. I 850
Plecia macilenta n. sp.16 Skartveit and Pika 2014 love bug
ETHZ Pl. I 853, 853 female
"Protomyia jucunda" = Plecia jucunda Heer 1849 love bug
ETHZ Pl. I 871, 853(x2), 856, 861, 867, 869 (some labeled by Brongniart fide Skartveit & Pika 2014)
 Diptera - Sciaridae
Sciara deleta n. sp.10
Sciara deleta n. sp.10 Heer 1864 dark-winged fungus gnat
 Diptera - Cecidomyiidae
Cecidomyia bremii n. sp.10 Heer 1864 gall midge
 Diptera - Mycetophilidae
Mycetophila pusillima n. sp.10 Heer 1864 fungus gnat
Mycetophila orci n. sp.10 Heer 1864 fungus gnat
 Coleoptera - Dytiscidae
"Cybister nicoleti" = Megadytes nicoleti9, Cybister agassizi n. sp.9, Cybister atavus n. sp.9, "Hydroporus benzeli n. sp." = Nebrioporus benzeli9, Dytiscus lavateri9, "Hydaticus areolatus n. sp." = Graphoderus heeri9, Colymbetes aemulus n. sp.9
"Cybister nicoleti" = Megadytes nicoleti9 Heer 1862 predaceous diving beetle
from Kesselstein
Cybister agassizi n. sp.9 Heer 1862 predaceous diving beetle
from Kesselstein
Cybister atavus n. sp.9 Heer 1862 predaceous diving beetle
from Kesselstein
"Hydroporus benzeli n. sp." = Nebrioporus benzeli9 Heer 1862 predaceous diving beetle
Dytiscus lavateri9 Heer 1847 predaceous diving beetle
from Kesselstein
"Hydaticus areolatus n. sp." = Graphoderus heeri9 Nilsson 2001 predaceous diving beetle
Colymbetes aemulus n. sp.9 Heer 1862 predaceous diving beetle
from Kesselstein
 Coleoptera - Carabidae
Nebria pluto n. sp.9 Heer 1862 ground beetle
1 from lower quarry, 2 from Kesselstein
Amara primigenia n. sp.9 Heer 1862 ground beetle
from Kesselstein
Amara pinguicula n. sp.9 Heer 1862 ground beetle
from Insektenschicht (lower quarry)
Amara princeps n. sp.9 Heer 1862 ground beetle
1 from Insektenschicht (lower quarry), 1 from Kesselstein
Harpalus constrictus n. sp.9 Heer 1862 ground beetle
from Kesselstein
Harpalus tardigradus n. sp.9 Heer 1862 ground beetle
from Insektenschicht (lower quarry)
Harpalus stierlini n. sp.9 Heer 1862 ground beetle
Harpalus stygius n. sp.9 Heer 1862 ground beetle
from Insektenschicht (lower quarry)
Dichirotrichus lividus9 Heer 1862 ground beetle
from Kesselstein
Badister grandis n. sp.9 Heer 1862 ground beetle
from Kesselstein
Badister fragilis n. sp.9 Heer 1862 ground beetle
from Kesselstein (2 var. paulo major)
Badister macrocephalus n. sp.9 Heer 1862 ground beetle
"multiple examples" in Kesselstein, including var. major
Sinis brevicollis n. gen. n. sp.9 Heer 1862 ground beetle
1 male, 1 female from lower quarry
Pterostichus minutulus n. sp.9 Heer 1862 ground beetle
from Insektenschicht (lower quarry)
Pterostichus vetustus n. sp.9 Heer 1862 ground beetle
from Insektenschicht (lower quarry)
Carabites laesicollis n. sp.9 Heer 1862 ground beetle
Calosoma nauckianum n. sp.8 Heer 1860 ground beetle
Calosoma escrobiculatum n. sp.8 Heer 1860 ground beetle
"Calosoma deplanatum n. sp." = Campalita deplanatum8 Heer 1860 ground beetle
Calosoma escheri n. sp.8 Heer 1860 ground beetle
Calosoma catenulatum n. sp.8 Heer 1860 ground beetle
Kesselstein
Glenopterus laevigatus n. gen. n. sp.6 Heer 1847 ground beetle
Trechinites oblongus n. sp.9 Heer 1862 ground beetle
from lower quarry
Trechinites clairvillii n. sp.9 Heer 1862 ground beetle
from lower quarry
 Coleoptera - Gyrinidae
"Dineutus longiventris n. sp." = Dineutus (Cyclinus) longiventris9, Miodineutes oeningensis n. gen. n. sp.5, Miodineutes heeri n. sp.5, "Dineutus insignis n. sp." = Miodineutes insignis9
"Dineutus longiventris n. sp." = Dineutus (Cyclinus) longiventris9 Heer 1862 whirligig beetle
from Kesselstein
Miodineutes oeningensis n. gen. n. sp.5 Hatch 1927 whirligig beetle
Dineutus insignis specimen in pl 2, fig 28 of Heer
Miodineutes heeri n. sp.5 Hatch 1927 whirligig beetle
Dineutus longiventris Specimen in pl 2, fig 25 of Heer
"Dineutus insignis n. sp." = Miodineutes insignis9 Heer 1862 whirligig beetle
from Kesselstein
 Coleoptera - Hydrophilidae
Hydrobius couloni n. sp.9, Hydrophilopsis elongata n. gen. n. sp.9, "Hydrous braunii" = Hydrobiomorpha braunii9, "Hydrous escheri n. sp." = Hydrobiomorpha braunii9, Hydrobiomorpha heeri n. sp.4, "Hydrophilus spectabilis" = Hydrophilus spectabilis6, "Hydrophilus giganteus n. sp." = Hydrophilus spectabilis9, "Hydrous rehmanni" = Hydrochara noachica9, Hydrous ovalis n. sp.9, Escheria bella n. sp.9
Hydrobius couloni n. sp.9 Heer 1862 water scavenger beetle
from Kesselstein
Hydrophilopsis elongata n. gen. n. sp.9 Heer 1862 water scavenger beetle
from Insektenschicht (lower quarry)
"Hydrous braunii" = Hydrobiomorpha braunii9 Heer 1847 water scavenger beetle
from Kesselstein and Insektenschicht (lower quarry)
"Hydrous escheri n. sp." = Hydrobiomorpha braunii9 Heer 1847 water scavenger beetle
from Insektenschicht (lower quarry); ETHZ NO-Z51.3-39-4:4-Pl.I148
Hydrobiomorpha heeri n. sp.4 Fikáček and Schmied 2013 water scavenger beetle
ETHZ NO-Z51.3-39-4:4-Pl. I 149b (label reads Hydrophilus ovalis)
"Hydrophilus spectabilis" = Hydrophilus spectabilis6 Heer 1847 water scavenger beetle
"Hydrophilus giganteus n. sp." = Hydrophilus spectabilis9 Heer 1847 water scavenger beetle
from Kesselstein; ETHZ NO-Z51.3-39-4:4-Pl.I143b
"Hydrous rehmanni" = Hydrochara noachica9 Heer 1847 water scavenger beetle
Hydrous ovalis n. sp.9 Heer 1862 water scavenger beetle
from Kesselstein
Escheria bella n. sp.9 Heer 1862 water scavenger beetle
 Coleoptera - Staphylinidae
Protactus minor n. sp.9 Heer 1862 rove beetle
from Insektenschicht (lower quarry)
Oxytelus proaevus n. sp.9 Heer 1862 rove beetle
from Insektenschicht (lower quarry)
Bledius speciosus n. sp.9 Heer 1862 rove beetle
from Insektenschicht (lower quarry)
Silpha tricostata9 Heer 1847 rove beetle
from Kesselstein
Scaphidium deletum9 Heer 1847 rove beetle
Scaphisoma gracile n. sp.9 Heer 1862 rove beetle
2 from Kesselstein, 1 from from Insektenschicht (lower quarry)
? Oxyporus seuberti n. sp.9 Heer 1862 rove beetle
Staphylinus atavus n. sp.9 Heer 1862 rove beetle
from Insektenschicht (lower quarry)
Lathrobium oeningense n. sp.9 Heer 1862 rove beetle
from Insektenschicht (lower quarry)
 Coleoptera - Histeridae
Hister mastodontis n. sp.9 Heer 1862 clown beetle
from Kesselstein
Hister maculigerus n. sp.9 Heer 1862 clown beetle
from Kesselstein
Hister morosus n. sp.9 Heer 1862 clown beetle
from Insektenschicht (lower quarry)
Hister coprolithorum n. sp.9 Heer 1862 clown beetle
numerous specimens from Kesselstein
Hister marmoratus n. sp.9 Heer 1862 clown beetle
from Kesselstein
Hister antiquus n. sp.9 Heer 1862 clown beetle
from Kesselstein
Hister aemulus n. sp.9 Heer 1862 clown beetle
several from Kesselstein
Hister vetustus n. sp.9 Heer 1862 clown beetle
several from Kesselstein
 Coleoptera - Glaphyridae
Glaphyrus antiquus n. sp.9 Heer 1862 bumble bee scarab beetle
from Insektenschicht (lower quarry)
 Coleoptera - Hybosoridae
Hybosorus lividus n. sp.9 Heer 1862 scavenger scarab beetle
 Coleoptera -
"Onthophagus urus" = Onthophagus urusheeri9, Gymnopleurus deperditus n. sp.9
"Onthophagus urus" = Onthophagus urusheeri9 Krell 2000 scarab beetle
"several" specimens
Gymnopleurus deperditus n. sp.9 Heer 1862 scarab beetle
from Kesselstein
 Coleoptera - Scarabaeidae
Gymnopleurus sisyphus n. sp.6, Gymnopleurus rotundatus n. sp.9, Copris druidum n. sp.9, Onthophagus bisontinus n. sp.9, Onthophagus prodromus n. sp.9, Onthophagus crassus n. sp.9, Oniticellus amplicollis n. sp.9, Anomala fugax n. sp.9, "Scarabaeus proserpinae n. sp." = Pentodon proserpinae9, Aphodius brevipennis n. sp.9, Lepitrix germanica n. sp.9, Serica minutula n. sp.9, Valgus oeningensis n. sp.9, ? Trichius rotundatus n. sp.9, Trichius lugubris n. sp.9, Trichius unifasciatus n. sp.9, "Trichius aedilis n. sp." = Gnorimus aedilis9
Gymnopleurus sisyphus n. sp.6 Heer 1847 scarab beetle
Gymnopleurus rotundatus n. sp.9 Heer 1862 scarab beetle
from Kesselstein
Copris druidum n. sp.9 Heer 1862 scarab beetle
from Kesselstein
Onthophagus bisontinus n. sp.9 Heer 1862 scarab beetle
from Insektenschicht (lower quarry)
Onthophagus prodromus n. sp.9 Heer 1862 scarab beetle
several from Kesselstein
Onthophagus crassus n. sp.9 Heer 1862 scarab beetle
from Kesselstein
Oniticellus amplicollis n. sp.9 Heer 1862 scarab beetle
from Kesselstein
Anomala fugax n. sp.9 Heer 1862 scarab beetle
from Kesselstein
"Scarabaeus proserpinae n. sp." = Pentodon proserpinae9 Heer 1862 rhinoceras beetle
from Kesselstien
Aphodius brevipennis n. sp.9 Heer 1862 scarab beetle
from Kesselstein
Lepitrix germanica n. sp.9 Heer 1862 scarab beetle
Serica minutula n. sp.9 Heer 1862 scarab beetle
from Kesselstein
Valgus oeningensis n. sp.9 Heer 1862 flower chafer
from Kesselstein
? Trichius rotundatus n. sp.9 Heer 1862 flower chafer
from Kesselstein
Trichius lugubris n. sp.9 Heer 1862 flower chafer
Trichius unifasciatus n. sp.9 Heer 1862 flower chafer
from Kesselstein
"Trichius aedilis n. sp." = Gnorimus aedilis9 Heer 1862 flower chafer
from Kesselstein
 Coleoptera -
"Copris subterranea n. sp." = Copris subterraneus9
"Copris subterranea n. sp." = Copris subterraneus9 Heer 1862 scarab beetle
from Kesselstein
 Coleoptera - Geotrupidae
Geotrupes germari n. sp.9 Heer 1862 earth-boring dung beetle
from Kesselstein
 Coleoptera - Byrrhidae
Byrrhus oeningensis9 Heer 1847 pill beetle
several from Kesselstein
 Coleoptera - Lampyridae
Lampyris orciluca n. sp.10 Heer 1864 beetle
 Coleoptera - Elateridae
"Elater (Alaus) spectabilis n. sp." = Alaus spectabilis10
"Elater (Alaus) spectabilis n. sp." = Alaus spectabilis10 Heer 1864 click beetle
 Coleoptera - Cantharidae
"Telephorus macilentus n. sp." = Cantharis (Cantharis) macilenta10
"Telephorus macilentus n. sp." = Cantharis (Cantharis) macilenta10 Heer 1864 soldier beetle
 Coleoptera - Buprestidae
Protogenia escheri n. gen. n. sp.6, Anthaxia pallida n. sp.9, ? Anthaxia doris n. sp.9, Buprestites agriloides n. sp.9, "Ancylocheira alemanica n. sp." = Buprestis alemanica14, "Ancylocheira concinna n. sp." = Buprestis concinna14, "Ancylochira tincta n. sp." = Buprestis tincta9, Acmaeodera antholitha n. sp.9, Acmaeodera brevicollis n. sp.9, Capnodis antiqua9, Capnodis spectabilis n. sp.9, Capnodis puncticollis9, Chalcophora pulchella n. sp.9, Chalcophora laevigata9
Protogenia escheri n. gen. n. sp.6 Heer 1847 jewel beetle
Anthaxia pallida n. sp.9 Heer 1862 jewel beetle
from Kesselstein
? Anthaxia doris n. sp.9 Heer 1862 jewel beetle
from Kesselstein
Buprestites agriloides n. sp.9 Heer 1862 jewel beetle
"Ancylocheira alemanica n. sp." = Buprestis alemanica14 Heer 1879 jewel beetle
"Ancylocheira concinna n. sp." = Buprestis concinna14 Heer 1879 jewel beetle
"Ancylochira tincta n. sp." = Buprestis tincta9 Heer 1862 jewel beetle
from Kesselstein,includes var. b. elytris majoribus
Acmaeodera antholitha n. sp.9 Heer 1862 jewel beetle
from Kesselstein
Acmaeodera brevicollis n. sp.9 Heer 1862 jewel beetle
from Kesselstein
Capnodis antiqua9 Heer 1847 jewel beetle
"several" specimens
Capnodis spectabilis n. sp.9 Heer 1862 jewel beetle
from Kesselstein
Capnodis puncticollis9 Heer 1847 jewel beetle
Chalcophora pulchella n. sp.9 Heer 1862 jewel beetle
from Kesselstein
Chalcophora laevigata9 Heer 1862 jewel beetle
 Coleoptera - Meloidae
Lytta aesculapii6 Heer 1847 blister beetle
Zonites vetusta n. sp.10 Heer 1864 blister beetle
 Coleoptera - Tenebrionidae
Tagenopsis brevicornis n. gen. n. sp.10
Tagenopsis brevicornis n. gen. n. sp.10 Heer 1864 darkling beetle
 Coleoptera - Nitidulidae
Nitidula pallida n. sp.9 Heer 1862 sap beetle
Nitidula ancora n. sp.9 Heer 1862 sap beetle
from Kesselstein
Nitidula maculigera n. sp.9 Heer 1862 sap beetle
 Coleoptera - Trogossitidae
Trogosita longicollis n. sp.9 Heer 1862 bark gnawing beetle
from Kesselstein
Trogosita bella n. sp.9 Heer 1862 bark gnawing beetle
Trogosita sculpturata n. sp.9 Heer 1862 bark gnawing beetle
from Kesselstein
Trogosita amissa n. sp.9 Heer 1862 bark gnawing beetle
from Kesselstein and Insektenschicht (lower quarry)
Trogosita koellikeri9 Heer 1847 bark gnawing beetle
Trogosita assimilis n. sp.9 Heer 1862 bark gnawing beetle
from Kesselstein
"Gymnochila obesa n. sp." = Gymnocheilis obesa9 Heer 1862 bark gnawing beetle
from Kesselstein
 Coleoptera - Lymexylidae
Hylecoetus cylindricus n. sp.10 Heer 1864 ship-timber beetle
 Coleoptera - Curculionidae
Sitona atavina n. sp.10 Heer 1864 snout beetle
Naupactus crassirostris n. sp.10 Heer 1864 snout beetle
Brachycerus nanus n. sp.10 Heer 1864 snout beetle
Lixus oeningensis n. sp.10 Heer 1864 snout beetle
Cleonus speciosus n. sp.10 Heer 1864 snout beetle
 Coleoptera - Brentidae
Miocenocylas heeri n. gen. n. sp.15, "Rhynchites dionysus n. sp." = Omphalapion dionysus10, Antliarhinites gracilis n. gen. n. sp.10, Apion antiquum n. sp.10
Miocenocylas heeri n. gen. n. sp.15 Legalov 2018 weevil
"Rhynchites dionysus n. sp." = Omphalapion dionysus10 Heer 1864 weevil
Antliarhinites gracilis n. gen. n. sp.10 Heer 1864 weevil
Apion antiquum n. sp.10 Heer 1864 weevil
 Coleoptera - Attelabidae
"Attelabus durus n. sp." = Phialodes durus10
"Attelabus durus n. sp." = Phialodes durus10 Heer 1864 leaf rolling weevil
 Coleoptera - Chrysomelidae
Gonioctena japeti n. sp.6, Galeruca buchi n. sp.10, Galeruca gemmifera n. sp.14, "Lema vetusta n. sp." = Crioceris vetusta10
Gonioctena japeti n. sp.6 Heer 1847 leaf beetle
Galeruca buchi n. sp.10 Heer 1864 leaf beetle
Galeruca gemmifera n. sp.14 Heer 1879 leaf beetle
"Lema vetusta n. sp." = Crioceris vetusta10 Heer 1864 leaf beetle
 Coleoptera - Cerambycidae
Lamia antiqua n. sp.14, Acanthoderus sepultus n. sp.10, Acanthoderus lepidus n. sp.10, "Prionus spectabilis n. sp." = Ergates spectabilis10, "Prionus polyphemus n. sp." = Ergates polyphemus10, Clytus pulcher n. sp.10, Callidium procerum n. sp.10, Callidium escheri n. sp.10
Lamia antiqua n. sp.14 Heer 1879 longhorned beetle
Acanthoderus sepultus n. sp.10 Heer 1864 longhorned beetle
Acanthoderus lepidus n. sp.10 Heer 1864 longhorned beetle
"Prionus spectabilis n. sp." = Ergates spectabilis10 Heer 1864 longhorned beetle
"Prionus polyphemus n. sp." = Ergates polyphemus10 Heer 1864 longhorned beetle
Clytus pulcher n. sp.10 Heer 1864 longhorned beetle
Callidium procerum n. sp.10 Heer 1864 longhorned beetle
Callidium escheri n. sp.10 Heer 1864 longhorned beetle
 Coleoptera - Coccinellidae
Coccinella amabilis n. sp.10, Coccinella colorata n. sp.10, "Coccinella decempustulata n. sp." = Coccinella heeri14, "Coccinella spectabilis n. sp." = Coccinella ponomarenkoi10
Coccinella amabilis n. sp.10 Heer 1864 ladybird beetle
Coccinella colorata n. sp.10 Heer 1864 ladybird beetle
"Coccinella decempustulata n. sp." = Coccinella heeri14 Ukrainsky 2010 ladybird beetle
Written "Coccinella 10-pustulata"
"Coccinella spectabilis n. sp." = Coccinella ponomarenkoi10 Ukrainsky 2010 ladybird beetle
 Coleoptera -
Cistelites spectabilis n. sp.13 Heer 1869 beetle
Hydrobius godeti n. sp.9 Heer 1862 beetle
from Libellenschicht (upper quarry)
Hydrophilus stenopterus n. sp.9 Heer 1862 beetle
from Kesselstein
Helophorus magnus n. sp.9 Heer 1862 beetle
from Kesselstein
 Lepidoptera -
"Bombycites" oeningensis n. sp. Heer 1849 moth
pupa
Bombycites buchii n. sp.10 Heer 1864 moth
 Hymenoptera -
Pteromalites oeningensis n. gen. n. sp.10
Pteromalites oeningensis n. gen. n. sp.10 Heer 1864 chalcid wasp
 Hymenoptera - Ichneumonidae
Ichneumon infernalis n. sp.10 Heer 1864 ichneumon wasp
Ichneumonites bellus n. sp.12 Heer 1867 ichneumon wasp
 Hymenoptera - Vespidae
Polistes primitiva n. sp.10 Heer 1864 wasp
Polistes kirbyanus n. sp.3 Cockerell 1915 wasp
ETH Pl. I, 681; Osmia kirbyana in Heer's label (unpublished)
 Hymenoptera - Scoliidae
Scolia saussureana n. sp.10 Heer 1864 wasp
 Hymenoptera - Megachilidae
Megachilidae indet.17, "Apis adamitica n. sp." = Lithurge adamitica10, Osmia nigra n. sp.17
Megachilidae indet.17 Latreille 1802 mason bee
ETH Pl. I, 687; Pl. I, 688(689); Pl. I, 718
"Apis adamitica n. sp." = Lithurge adamitica10 Heer 1864 mason bee
ETH V.S. 4 (Heer's holotype); SN.200 (apparently different from Heer specimens) in Zeuner & Manning, 1976
Osmia nigra n. sp.17 Zeuner and Manning 1976 mason bee
ETH Pl. I, 719; Pl. I, 685
 Hymenoptera -
Andrena primaeva n. sp.2 Cockerell 1909 bee
SN.202
 Hymenoptera - Apidae
Bombus abavus n. sp.17, Anthophorites longaeva17, Anthophorites mellona n. gen. n. sp., Anthophorites veterana17, Xylocopa senilis17, "Bombus atavus n. sp." = Xylocopa atavus12, "Bombus jurinei n. sp." = Xylocopa jurinei12, Bombus grandaevus12
Bombus abavus n. sp.17 Heer 1867 bumble bee
Holotype (lost), ETH Pl. I, 808 (examined by Cockerell), SN.201
Anthophorites longaeva17 Heer 1867 bee
ETH Pl. I, 683, 712, 809 (not Heer's types)
Anthophorites mellona n. gen. n. sp. Heer 1849 bee
Anthophorites veterana17 Heer 1849 bee
ETH Pl. I, 684
Xylocopa senilis17 Heer 1849 large carpenter bee
ETH 21a(b), doutbfully 21c
"Bombus atavus n. sp." = Xylocopa atavus12 Heer 1867 large carpenter bee
"Bombus jurinei n. sp." = Xylocopa jurinei12 Heer 1864 large carpenter bee
ETH Pl. I, 682a (type in Heer, 1865), 682b (specimen in heer, 1867)
Bombus grandaevus12 Heer 1849 bee
 Hymenoptera - Sphecidae
Ammophila annosa n. sp.10 Heer 1864 digger wasp
Ammophila inferna n. sp.10 Heer 1864 digger wasp
 Hymenoptera - Formicidae
"Ponera veneraria n. sp." = Poneropsis veneraria10, "Ponera globosa n. sp." = Poneropsis globosa, Poneropsis stygia n. sp.12, Imhoffia pallida n. sp.12, "Formica heraclea" = Camponotus heracleus, "Formica lignitum" = Camponotus lignitus, Formica orbata
"Ponera veneraria n. sp." = Poneropsis veneraria10 Heer 1864 ant
"Ponera globosa n. sp." = Poneropsis globosa Heer 1849 ant
Poneropsis stygia n. sp.12 Heer 1867 ant
Imhoffia pallida n. sp.12 Heer 1864 ant
"Formica heraclea" = Camponotus heracleus Heer 1849 ant
"Formica lignitum" = Camponotus lignitus Germar 1837 ant
Formica orbata Heer 1849 ant
 Orthoptera - Tettigoniidae
"Decticus speciosus n. sp." = Platycleis speciosa10
"Decticus speciosus n. sp." = Platycleis speciosa10 Heer 1864 katydid
 Orthoptera - Gryllotalpidae
Gryllotalpa stricta n. sp.10 Heer 1864 mole cricket
 Orthoptera - Gryllidae
Gryllus troglodytes n. sp.10 Heer 1864 true crickets
 Orthoptera - Pyrgomorphidae
"Oedipoda fischeri n. sp." = Miopyrgomorpha fischeri10
"Oedipoda fischeri n. sp." = Miopyrgomorpha fischeri10 Heer 1864 gaudy grasshopper
 Orthoptera - Acrididae
Oedipoda germari n. sp.10 Heer 1864 short-horned grasshopper
 Orthoptera - Tetrigidae
"Tetrix gracilis n. sp." = Tettigidea gracilis10
"Tetrix gracilis n. sp." = Tettigidea gracilis10 Heer 1864 pygmy grasshopper
 Dermaptera - Forficulidae
Forficula minuta n. sp.10, Forficula recta n. sp.10
Forficula minuta n. sp.10 Heer 1864 common earwig
Forficula recta n. sp.10 Heer 1864 common earwig
 Blattodea - Blattidae
"Blattidium coloratum n. sp." = Blatta colorata11
"Blattidium coloratum n. sp." = Blatta colorata11 Heer 1864 cockroach
from Kesselstein
 Blattodea - Rhinotermitidae
"Termes hartungi n. sp." = Reticulitermes hartungi10
"Termes hartungi n. sp." = Reticulitermes hartungi10 Heer 1864 termite
 Blattodea - Termitidae
Termes buchii n. sp.10
Termes buchii n. sp.10 Heer 1864 termite
 Hemiptera - Nabidae
Prostemma oeningensis n. sp.7 Heer 1853 true bug
lower quarry
 Hemiptera - Tingidae
"Monanthia wollastoni n. sp." = Dictyla wollastoni10
"Monanthia wollastoni n. sp." = Dictyla wollastoni10 Heer 1864 lace bug
 Hemiptera - Reduviidae
Harpactor maculipes n. sp.7 Heer 1853 assassin bug
lower quarry
Harpactor obsoletus n. sp.7 Heer 1853 assassin bug
from the Kesselstein
 Hemiptera - Gerridae
Limnobates prodromus n. sp.10 Heer 1864 water strider
 Hemiptera - Alydidae
"Nabis gracillima n. sp." = Licremytra gracillima10, Alydus pulchellus n. sp.7
"Nabis gracillima n. sp." = Licremytra gracillima10 Heer 1864 broad-headed bug
Alydus pulchellus n. sp.7 Heer 1853 broad-headed bug
lower quarry
 Hemiptera - Rhyparochromidae
Pachymerus obsoletus n. sp.7 Heer 1853 seed bug
lower quarry
 Hemiptera - Lygaeidae
Cephalocoris pilosus n. gen. n. sp.7
Cephalocoris pilosus n. gen. n. sp.7 Heer 1853 milkweed bug
lower quarry
 Hemiptera - Heterogastridae
Heterogaster tristis n. sp.10 Heer 1864 true bug
 Hemiptera - Scutelleridae
Pachycoris protogaeus n. sp.7 Heer 1853 metallic shield bug
lower quarry
Pachycoris burmeisteri n. sp.10 Heer 1864 metallic shield bug
Pachycoris escheri7 Heer 1853 metallic shield bug
 Hemiptera - Cydnidae
Cydnus oeningensis7 Heer 1853 burrowing bug
lower quarry
Cydnopsis pygmaea n. sp.7 Heer 1853 burrowing bug
upper quarry
Cydnopsis deleta n. sp.7 Heer 1853 burrowing bug
lower quarry
Cydnopsis exilis n. sp.7 Heer 1853 burrowing bug
 Hemiptera - Pentatomidae
Halys bruckmanni7 Heer 1853 stink bug
from the Kesselstein
Halys spectabilis n. sp.10 Heer 1864 stink bug
Eurydema arcuata n. sp.7 Heer 1853 stink bug
from the Kesselstein
Eurydema impudica7 Heer 1853 stink bug
Pentatoma pictum n. sp.10 Heer 1864 stink bug
 Hemiptera - Corixidae
"Corisa fasciolata" = Corixa fasciolata7
"Corisa fasciolata" = Corixa fasciolata7 Heer 1853 water boatman
from the Libellen-schicht
 Hemiptera - Belostomatidae
"Belostoma speciosum n. sp." = Belostomates speciosum10
"Belostoma speciosum n. sp." = Belostomates speciosum10 Heer 1864 giant water bug
 Hemiptera - Cercopidae
Cercopis herrichi n. sp.10, Cercopis hageni n. sp.10, Cercopis germari n. sp.10
Cercopis herrichi n. sp.10 Heer 1864 froghopper
Cercopis hageni n. sp.10 Heer 1864 froghopper
Cercopis germari n. sp.10 Heer 1864 froghopper
 Thysanoptera -
Thrips annosa n. sp.10, Thrips oeningensis n. sp.10
Thrips annosa n. sp.10 Heer 1864 thrips
Thrips oeningensis n. sp.10 Heer 1864 thrips
 Ephemeroptera -
Ephemera oeningensis n. sp.10 Heer 1864 mayfly
 Odonata -
"Libellula" thetis Heer 1849 true dragonfly
larva
 Odonata - Gomphidae
Lindenia heeri n. sp.1 Boderau et al. 2024 clubtail dragonfly
ETH S.N. 228 (labeled "Aeschna tyche")
 Odonata - Sieblosiidae
"Agrion iris n. sp." = Stenolestes iris14
"Agrion iris n. sp." = Stenolestes iris14 Heer 1879 damselfly
 Odonata -
"Agrion (Sterope) parthenope n. sp." = Steropoides parthenope
"Agrion (Sterope) parthenope n. sp." = Steropoides parthenope Heer 1849 damselfly