Menat (Piton collection) (Paleocene of France)

Where: Auvergne, France (46.1° N, 2.9° E: paleocoordinates 40.3° N, 0.7° E)

• coordinate stated in text

• outcrop-level geographic resolution

When: Menat Formation, Selandian (61.6 - 59.2 Ma)

• The estimated age of the Menat site varies. It was previously dated by pollen analysis, mammal stratigraphy, and an unpublished radiometric K/Ar analysis as late Paleocene (see Kedves & Russell 1982). However, several lines of more recent evidence indicate an older, middle Paleocene (Selandian) age, of about 60-61 Ma. Paleomagnetic studies reveal that the eruption took place during a reversed interval (Kedves & Russell 1982). Fossils of the mammal Plesiadapis from Menat enable the locality to be connected to the biostratigraphic framework established in the Western Interior USA that relies extensively on this genus (Gingerich 1976; Secord et al. 2006). The abundance of the genus, its rapid evolution, and its Holarctic dispersal history allow species of Plesiadapis to play an important role in the zonation of Paleocene continental sediments and in correlation of faunas on both sides of the North Atlantic (e.g., Gingerich 1976). Two remarkable skeletons of P. insignis, one of them nearly complete, have been found in lake deposits at Menat, and this species closely compares to P. praecursor / P. anceps in North America (pers. comm. P. Gingerich August 2008). This suggests an early Selandian age, equivalent to the early Tiffanian Ti-1/Ti-2 land mammal biozonation sensu Secord et al. (2006) (see also Gingerich 1976; Russell 1967, 1975). In North America, the Ti1 and Ti2 biozones occur within magnetic polarity Chron 26r (Secord et al. 2006), consistent with the reversed polarity of the Menat sediments.

Environment/lithology: crater lake; diatomite

• Volcanic Paleolake (maar) deposit
• spongo-diatomitic

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by Piton; reposited in the MNHN

Primary reference: L. Piton. 1940. Paléontologie du Gisement Éocéne de Menat (Puy-de-Dôme) (Flore et Faune) 1-303 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 113685: authorized by Matthew Clapham, entered by Jered Karr on 04.08.2011, edited by Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Diptera - Limoniidae
"Limnobia theobaldi n. sp." = Limonia theobaldi
"Limnobia theobaldi n. sp." = Limonia theobaldi Piton 1940 crane fly
Coll. Piton 325
 Diptera - Tabanidae
Chrysops seguyi n. sp. Piton 1940 deer fly
Coll. Piton 541
 Diptera - Bibionidae
Bibio sp. Geoffroy 1762 march fly
 Odonata - Lestidae
"Lestes" zalesskyi n. sp.4 Piton 1940 spread-winged damselfly
Coll. Piton 190
 Odonata - Aeshnidae
Aeshna sp.3 Fabricius 1775 hawker dragonfly
Coll. Piton 193
 Hymenoptera - Tenthredinidae
Tenthredo primordialis n. sp. Piton 1940 wood wasp
Coll. Piton 268
 Hymenoptera -
Manevalia pachyliformis n. gen. n. sp.
Manevalia pachyliformis n. gen. n. sp. Piton 1940 wood wasp
Coll. Piton 306, 307
 Hymenoptera - Siricidae
"Eosirex ligniticus n. gen. n. sp." = Urocerus ligniticus
"Eosirex ligniticus n. gen. n. sp." = Urocerus ligniticus Piton 1940 horntail
Coll. Piton 743( MNHN IPM-R07004, IPM-B47308)
 Hymenoptera - Ichneumonidae
"? Phaenolobus arvernus n. sp." = Hallocinetus arvernus
"? Phaenolobus arvernus n. sp." = Hallocinetus arvernus Piton 1940 ichneumon wasp
Coll. Piton 663 (MNHN.F.A57301)
 Hymenoptera - Crabronidae
Gorytes statzi n. sp. Piton 1940 apoid wasp
Coll. Piton 314
 Hymenoptera - Megachilidae
Probombus hirsutus n. gen. n. sp.
Probombus hirsutus n. gen. n. sp. Piton 1940 mason bee
Coll. Piton 628
 Hymenoptera - Thynnidae
Myzine madeleinae n. sp. Piton 1940 wasp
Coll. Piton 705
 Hymenoptera - Vespidae
Eumenes projaponica n. sp. Piton 1940 potter wasp
Coll. Piton 532
Ancistrocerus eocenicus n. sp. Piton 1940 potter wasp
Coll. Piton 936
Ancistrocerus berlandi n. sp. Piton 1940 potter wasp
Coll. Piton 340
Odynerus manevali n. sp. Piton 1940 potter wasp
Coll. Piton 417, 418, 419
"Rygchium andrei n. sp." = Rhynchium andrei Piton 1940 potter wasp
Coll. Piton 851
Polistes vergnei n. sp. Piton 1940 wasp
Coll. Piton 409
 Hymenoptera - Myrmosidae
Myrmosa brunnea n. sp. Piton 1940 wasp
Coll. Piton 185
 Hymenoptera -
Prohepialus incertus n. gen. n. sp.
Prohepialus incertus n. gen. n. sp. Piton 1940 winged insect
Coll. Piton 426
 Coleoptera -
Euchroma arvernica n. sp. Piton 1940 beetle
Coll. Piton 27
Megopis lineolatus n. sp. Piton 1940 beetle
Coll. Piton 177
 Coleoptera - Elateridae
"Adelocera jungi n. sp." = Lacon jungi, Semiotus menatensis n. sp.
"Adelocera jungi n. sp." = Lacon jungi Piton 1940 click beetle
Coll. Piton 76
Semiotus menatensis n. sp. Piton 1940 click beetle
Coll. Piton 943
 Coleoptera - Buprestidae
Ancylochira pinguis n. sp., Ancylochira striatopunctata n. sp., Buprestites lineatocollis n. sp., Psiloptera vermiculata n. sp., "Ancylochira octomaculata n. sp." = Buprestis octomaculata, "Eurythyrea viridis n. sp." = Buprestis viridis, Ancylochira metallica n. sp., Eurythyrea micropunctata n. sp., "Anthaxia biimpressa n. sp." = Anthaxia eocenica, Anthaxia ventricosa n. sp., Stigmodera unicolor n. sp., Polybothris primordialis n. sp., "Lampra gautieri" = Lamprodila gautieri, "Psiloptera (Lampetis) aurostriata" = Lamprodila gautieri, Iridotaenia hovassei n. sp., Ancylochira violaceocyanescens n. sp., Chrysochroa punctata n. sp.
Ancylochira pinguis n. sp. Piton 1940 jewel beetle
Coll. Piton 412
Ancylochira striatopunctata n. sp. Piton 1940 jewel beetle
Coll. Piton 625
Buprestites lineatocollis n. sp. Piton 1940 jewel beetle
Coll. Piton 112
Psiloptera vermiculata n. sp. Piton 1940 jewel beetle
Coll. Piton 381
"Ancylochira octomaculata n. sp." = Buprestis octomaculata Piton 1940 jewel beetle
Coll. Piton 85, 86
"Eurythyrea viridis n. sp." = Buprestis viridis Piton 1940 jewel beetle
Coll. Piton 840
Ancylochira metallica n. sp. Piton 1940 jewel beetle
Coll. Piton 347, 348
Eurythyrea micropunctata n. sp. Piton 1940 jewel beetle
Coll. Piton 450
"Anthaxia biimpressa n. sp." = Anthaxia eocenica Bílý 1996 jewel beetle
Coll. Piton 926
Anthaxia ventricosa n. sp. Piton 1940 jewel beetle
Coll. Piton 270
Stigmodera unicolor n. sp. Piton 1940 jewel beetle
Coll. Piton 284, 285
Polybothris primordialis n. sp. Piton 1940 jewel beetle
Coll. Piton 606
"Lampra gautieri" = Lamprodila gautieri Bruyant 1902 jewel beetle
Coll. Piton 11
"Psiloptera (Lampetis) aurostriata" = Lamprodila gautieri Bruyant 1902 jewel beetle
Coll. Piton 400, 401
Iridotaenia hovassei n. sp. Piton 1940 jewel beetle
Coll. Piton 818
Ancylochira violaceocyanescens n. sp. Piton 1940 jewel beetle
Coll. Piton 720, 721
Chrysochroa punctata n. sp. Piton 1940 jewel beetle
Coll. Piton 814
 Coleoptera - Tenebrionidae
Ancylochira oligostriata Piton and Théobald 1937 darkling beetle
Coll. Piton 589
 Coleoptera - Attelabidae
Phytonomus punctatus n. sp. Piton 1940 leaf rolling weevil
Coll. Piton 805
 Coleoptera - Curculionidae
Hipporhinus ventricosus n. sp. Piton 1940 snout beetle
Coll. Piton 312
Lixus ligniticus n. sp. Piton 1940 snout beetle
Coll. Piton 254
Centrinus longipes n. sp. Piton 1940 snout beetle
Coll. Piton 927
"Balaninus elegans n. sp." = Menatorhis elegans Piton 1940 snout beetle
Coll. Piton 730
Rhysosternum punctatolineatum n. sp. Piton 1940 snout beetle
Coll. Piton 46
 Coleoptera - Brentidae
Apion sp. Herbst 1797 weevil
Coll. Piton 499
 Coleoptera - Cerambycidae
"Palaeoncoderes piacentinii n. sp." = Palaeoncoderes piacentini Piton 1940 longhorned beetle
Coll. Piton 525
Palaeoncoderes eocenicus Piton and Théobald 1937 longhorned beetle
Coll. Piton 12, 57
Monohammus orientalis n. sp. Piton 1940 longhorned beetle
Coll. Piton 771
Prolamioides brunneus n. sp. Piton 1940 longhorned beetle
Coll. Piton 162
Prionus sinuatus n. sp. Piton 1940 longhorned beetle
Coll. Piton 260
Chlorida magnifica n. sp. Piton 1940 longhorned beetle
Coll. Piton 610, 611
 Coleoptera - Chrysomelidae
Cryptocephalus minusculus n. sp., "Clythrina eocenica n. gen. n. sp." = Clytrina eocenica, "Haltica dryophyllorum n. sp." = Altica dryophyllorum
Cryptocephalus minusculus n. sp. Piton 1940 leaf beetle
Coll. Piton 980
"Clythrina eocenica n. gen. n. sp." = Clytrina eocenica Piton 1940 leaf beetle
Coll. Piton 843
"Haltica dryophyllorum n. sp." = Altica dryophyllorum Piton 1940 flea beetle
Coll. Piton 447
 Coleoptera - Scarabaeidae
Aphodius charauxi n. sp. Piton 1940 scarab beetle
Coll. Piton 250
 Coleoptera - Bolboceratidae
Bolboceras inermis n. sp. Piton 1940 dor beetle
Coll. Piton 247
 Coleoptera - Staphylinidae
Oxyporus impressus n. sp. Piton 1940 rove beetle
Coll. Piton 940
 Coleoptera - Carabidae
Harpalus lheritieri n. sp. Piton 1940 ground beetle
Coll. Piton 75
Bembidium cyaneomicans n. sp. Piton 1940 ground beetle
Coll. Piton 28
Agonum antiquum n. sp. Piton 1940 ground beetle
Coll. Piton 150
 Coleoptera - Gyrinidae
"Dineutes giganteus n. sp." = Gyretes giganteus
"Dineutes giganteus n. sp." = Gyretes giganteus Piton 1940 whirligig beetle
Coll. Piton 141, 143
 Coleoptera - Cupedidae
"Zonabris immaculatus n. sp." = Cainomerga immaculata
"Zonabris immaculatus n. sp." = Cainomerga immaculata Piton 1940 reticulated beetle
Coll. Piton 650
 Coleoptera -
Acmaeodera perreaui n. sp. Piton 1940 beetle
Coll. Piton 575
 Trichoptera - Leptoceridae
Prodontocerum ligniticum n. gen. n. sp.
Prodontocerum ligniticum n. gen. n. sp. Piton 1940 long-horned caddisfly
Coll. Piton 227, 228
 Trichoptera - Phryganeidae
Phryganea nigripennis n. sp. Piton 1940 large caddisfly
Coll. Piton 218,219
 Hemiptera - Aphrophoridae
"Aphrophora maculata n. sp." = Aphrophora pitoni5
"Aphrophora maculata n. sp." = Aphrophora pitoni5 Dmitriev 2020 froghopper
Coll. Piton 50
 Hemiptera - Tettigarctidae
Meuniera haupti n. sp.5 Piton 1936 hairy cicada
Coll. Piton 49 (MNHN-IPM-R0790)
 Hemiptera - Cicadidae
"Rihana beauchampi n. sp." = Chremistica beauchampi
"Rihana beauchampi n. sp." = Chremistica beauchampi Piton 1940 cicada
Coll. Piton 904
 Hemiptera - Cicadellidae
Bythoscopus pulcher n. sp. Piton 1940 leafhopper
Coll. Piton 863
Tettigonella chazei n. sp. Piton 1940 leafhopper
Coll. Piton 837
 Hemiptera -
Lithopsis punctinervis n. sp. Piton 1940 true bug
Coll. Piton 585
 Hemiptera - Ricaniidae
Hammapteryx eocenicus n. sp. Piton 1940 planthopper
Coll. Piton 88, 105
 Hemiptera -
Lithopsis lineatus n. sp. Piton 1940 true bug
Coll. Piton 432
 Hemiptera - Tessaratomidae
Pycanum griseum n. sp. Piton 1940 giant shield bug
Coll. Piton 69
Pycanum vanvollenhoveni n. sp. Piton 1940 giant shield bug
Coll. Piton 648, 649
 Hemiptera - Plataspidae
Coptosoma eocenica n. sp. Piton 1940 true bug
Coll. Piton 335, 336
 Hemiptera - Scutelleridae
Teleocydnus auriculatus n. sp. Piton 1940 metallic shield bug
Coll. Piton 333
 Hemiptera - Pentatomidae
Deryeuma primordialis n. gen. n. sp. Piton 1940 stink bug
Coll. Piton 118, 119
Doryderes laticollis n. sp. Piton 1940 stink bug
Coll. Piton 169
Asopus puncticollis n. sp. Piton 1940 stink bug
Coll. Piton 155
 Orthoptera -
Ochrilidia lineata n. sp. Piton 1940 grasshopper
Coll. Piton 161
 Orthoptera - Acrididae
Oedipoda indet. Latreille 1829 band-winged grasshopper
Coll. Piton 226
 Orthoptera - Tettigoniidae
Prophasgonura lineatocollis n. gen. n. sp. Piton 1940 katydid
Coll. Piton 148
Conocephalus martyi n. sp. Piton 1940 katydid
Coll. Piton 225
Protempusa incerta n. gen. n. sp. Piton 1940 katydid
Coll. Piton 718, 719
 Orthoptera -
Eremobiites sp. Piton 1940 grasshopper
Coll. Piton 36
 Dicondylia -
Orthacanthacris incertus n. sp. Piton 1940 winged insect
Coll. Piton 18
 Blattodea -
Prochaeradodis enigmaticus n. gen. n. sp. Piton 1940 cockroach
Coll. Piton 230
 Blattodea - Blattidae
Protectobia primordialis n. gen. n. sp. Piton 1940 cockroach
Coll. Piton 802
Zeunera superba n. sp. Piton 1940 cockroach
Coll. Piton 115, 116
Zeunera madeleinae n. sp. Piton 1936 cockroach
Coll. Piton 52
Prostylopyga gigantea n. gen. n. sp. Piton 1940 cockroach
Coll. Piton 704
Periplaneta houlberti n. sp. Piton 1940 cockroach
Coll. Piton 243, 244
Eoplaneta desforesti Piton and Théobald 1937 cockroach
Coll. Piton 223
 Blattodea - Kalotermitidae
Neotermes grassei n. sp. Piton 1940 termite
MNHN IPM-R.54380 (Coll. Piton 807)
 Blattodea - Ectobiidae
Chopardia spinipes n. gen. n. sp. Piton 1940 cockroach
Coll. Piton 751
Ectobia arverniensis n. sp. Piton 1940 croton bug
Coll. Piton 808
Ectobia menatensis n. sp. Piton 1940 croton bug
Coll. Piton 750
 Blattodea - Blaberidae
Gyna obesa n. sp.
Gyna obesa n. sp. Piton 1940 giant cockroach
Coll. Piton 979
 Mantodea - Chaeteessidae
Arvernineura insignis n. gen. n. sp., "Archaeophlebia enigmatica n. gen. n. sp." = Louispitonia enigmatica
Arvernineura insignis n. gen. n. sp. Piton 1940 mantis
Coll. Piton 715
"Archaeophlebia enigmatica n. gen. n. sp." = Louispitonia enigmatica Piton 1940 mantis
Coll. Piton 640, 641
 Dicondylia -
Menatacridium eocenicum n. sp. Piton 1936 winged insect
Coll. Piton 51
Dictyophara scudderi n. sp. Piton 1940 winged insect
Coll. Piton 925
Reptilia
 Choristodera -
Lazarussuchus sp.1 Hecht 1992 choristodere
BDL 1819 - comprises a skull and postcranial skeleton in articulation, preserved as part and counterpart
 Crocodylia -
Menatalligator bergouniouxi n. gen. n. sp.6 Piton 1940 crocodilian
Aves
 Aves -
Aves "gen. et sp. indet. A"2 bird
MNT-11-7952
Aves "gen. et sp. indet. B"2 bird
MNHN-MEN 9
 Psittaciformes - Messelasturidae
Messelasturidae indet.2 Mayr 2005 parrot
MNHN-MEN 69
 Ralliformes - Ralloidae
"? Ralloidea indet." = Ralloidae2, cf. Songziidae indet.2
"? Ralloidea indet." = Ralloidae2 Rafinesque 1815 bird
cf. Songziidae indet.2 Hou 1990 bird
MNHN-MEN 70 and MNHM.16037