PBDB Collection

Shanwang, Linqu County, collection s (Shandong Provincial Museum) (Miocene of China)

Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)

• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone

• Two minor volcanic faulted basins, the Linqu and Shanwang Basins, are present in the study area. The former lies near the county town of Linqu, and the latter in the Shanwang National Geopark, east of this town. The faulted basins are the result of intense Mesozoic-Cenozoic tectonic extension and rifting; the development of these basins was closely related in time with the nearby NE-SW and NW-SE trending faults in the Yishu Fault Zone.

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Shandong Provincial Museum. This collection contains specimens labeled with five-digit numbers prefaced by letter s.

Primary reference: J. F. Zhang. 1989. Fossil Insects From Shanwang, Shandong, China 1-459 [D. Smith/E. Leckey/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 128552: authorized by Matthew Clapham, entered by Matthew Clapham on 11.06.2012

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Arachnida
Araneae - Araneidae	Araneus luianus n. sp.³ Araneus luianus n. sp.³ Zhang et al. 1994 orb-weaving spider s82704
Araneae - Thomisidae	Miothomisus subnudus n. sp.³, "Ectona brunnea n. sp." = Ecotona brunnea³ Miothomisus subnudus n. sp.³ Zhang et al. 1994 crab spider s83401 "Ectona brunnea n. sp." = Ecotona brunnea³ Zhang et al. 1994 crab spider s85611
Insecta
Diptera - Chironomidae	Bythomyia oryctes n. gen. n. sp.¹ Bythomyia oryctes n. gen. n. sp.¹ Zhang 1989 non biting midge s32469
Diptera - Bibionidae	Plecia villosa n. sp., Plecia spinula n. sp., Plecia aculeolata n. sp., Bibio ventricosus, Bibio expansus n. sp., "Bibio abdominalis" = Bibio brunettii, Bibio tranopterus, Bibio colossoceps n. sp., Bibio varineurus Plecia villosa n. sp. Zhang 1989 love bug s82650, s82766 Plecia spinula n. sp. Zhang 1989 love bug s82652, s82675 Plecia aculeolata n. sp. Zhang 1989 love bug s82651 Bibio ventricosus Zhang 1989 march fly s82655 Bibio expansus n. sp. Zhang 1989 march fly s82654 "Bibio abdominalis" = Bibio brunettii Hardy 1973 march fly s82659 Bibio tranopterus Zhang 1989 march fly s82653 Bibio colossoceps n. sp. Zhang 1989 march fly s82658 Bibio varineurus Zhang 1989 march fly s82718
Diptera - Asilidae	Asilus brunneitibius n. sp., Asilus angustialius n. sp. Asilus brunneitibius n. sp. Zhang 1989 robber fly s82664 Asilus angustialius n. sp. Zhang 1989 robber fly s82663
Diptera - Pallopteridae	Palloptera hypolithica n. sp. Palloptera hypolithica n. sp. Zhang 1989 fly s82665
Ephemeroptera - Heptageniidae	Heptagenia sp. Heptagenia sp. Walsh 1863 stream mayfly s82707, s82712, s82724
Odonata - Macromiidae	Epophthalmia zotheca n. sp. Epophthalmia zotheca n. sp. Zhang 1989 true dragonfly s82725/6
Odonata - Aeshnidae	"Mediaeschna lucida n. sp." = Epiaeschna lucida "Mediaeschna lucida n. sp." = Epiaeschna lucida Zhang 1989 darner dragonfly s82681
Odonata - Coenagrionidae	Agriocnemis lacteola Agriocnemis lacteola Selys 1877 wisp s82682
Hemiptera - Fulgoridae	Limois shanwangensis Limois shanwangensis Hong 1979 fulgorid planthopper s82668, s82671
Hemiptera - Rhopalidae	Stictopleurus breviantennus n. sp. Stictopleurus breviantennus n. sp. Zhang 1989 scentless plant bug s82703
Hemiptera - Pentatomidae	Stollia rubibrunneus n. sp., Carbula aff. crassiventris, Carbula pustulosa n. sp. Stollia rubibrunneus n. sp. Zhang 1989 stink bug s82716 Carbula aff. crassiventris Dallas 1849 stink bug s82676 Carbula pustulosa n. sp. Zhang 1989 stink bug s82675
Hemiptera - Urostylididae	? Urochela sp., Urochela falsirubra n. sp., Urochela pardalina n. sp. ? Urochela sp. Dallas 1851 true bug s82674 Urochela falsirubra n. sp. Zhang 1989 true bug s82677 Urochela pardalina n. sp. Zhang 1989 true bug s82678
Hemiptera - Tingidae	Agramma tertiarium n. sp. Agramma tertiarium n. sp. Zhang 1989 lace bug s82730
Hemiptera - Nepidae	Laccotrephes cf. robustus Laccotrephes cf. robustus Stal 1870 true bug s82717
Hemiptera - Corixidae	Corixa triquetra n. sp. Corixa triquetra n. sp. Zhang 1989 water boatman s82679
Hemiptera - Aphididae	"Cinara limnogena n. sp." = Cinara (Cinara) limnogena, "Cinara elegans n. sp." = Cinara (Cinara) elegans "Cinara limnogena n. sp." = Cinara (Cinara) limnogena Zhang 1989 aphid s82673 "Cinara elegans n. sp." = Cinara (Cinara) elegans Zhang 1989 aphid s82674
Dicondylia -	Oiophassus nycterus n. gen. n. sp. Oiophassus nycterus n. gen. n. sp. Zhang 1989 winged insect s82702
Orthoptera - Acrididae	Tumearolium humifuse n. gen. n. sp. Tumearolium humifuse n. gen. n. sp. Zhang 1989 slant-faced grasshopper s82697
Dermaptera - Forficulidae	Allodahlia shanwangensis Allodahlia shanwangensis Zhou 1986 common earwig s82727, s82729
Blattodea - Blattidae	Periplaneta lacera n. sp. Periplaneta lacera n. sp. Zhang 1989 cockroach s82696
Blattodea - Kalotermitidae	"Kalotermes shandongianus n. sp." = Glyptotermes shandongianus "Kalotermes shandongianus n. sp." = Glyptotermes shandongianus Zhang 1989 termite s82767
Hymenoptera -	"Protelenomus miocenicus n. gen. n. sp." = Sinoprotelenomus miocenicus "Protelenomus miocenicus n. gen. n. sp." = Sinoprotelenomus miocenicus Zhang 1989 wasp s82643
Hymenoptera - Ichneumonidae	Exetastes areolus n. sp., Corynobatopsis anfracta n. gen. n. sp., Mesoleptus stigmus n. sp., ? Ovipterygium flexuosum n. sp., Yezoceryx brachycoleos n. sp., Agrothereutes townesi n. sp. Exetastes areolus n. sp. Zhang 1989 ichneumon wasp s82645 Corynobatopsis anfracta n. gen. n. sp. Zhang 1989 ichneumon wasp s82649 Mesoleptus stigmus n. sp. Zhang 1989 ichneumon wasp s82646 ? Ovipterygium flexuosum n. sp. Zhang 1989 ichneumon wasp s82647 Yezoceryx brachycoleos n. sp. Zhang 1989 ichneumon wasp s82644 Agrothereutes townesi n. sp. Zhang 1989 ichneumon wasp s82648
Hymenoptera - Vespidae	"Vespa magnifica" = Vespa mandarinia magnifica "Vespa magnifica" = Vespa mandarinia magnifica Smith 1852 Asian giant hornet s82640
Hymenoptera - Tiphiidae	Tiphia brevala n. sp., Tiphia varia n. sp., Tiphia mediovena n. sp. Tiphia brevala n. sp. Zhang 1989 wasp s82641 Tiphia varia n. sp. Zhang 1989 wasp s82633 Tiphia mediovena n. sp. Zhang 1989 wasp s82642
Hymenoptera - Formicidae	Formica linquensis, Formica ovala, Camponotus festinus, Camponotus microthoracus, Camponotus gracilis n. sp., Camponotus ampullosus n. sp., Camponotus ambon n. sp., Camponotus shanwangensis, Alloiomma changweiensis n. sp., Elaphrodites scutulatus n. gen. n. sp., Dolichoderus evolans n. sp., Dolichoderus lacinius n. sp., Technomyrmex septentrionalis n. sp., Myopopone sinensis n. sp. Formica linquensis Zhang 1989 ant s82623 Formica ovala Zhang 1989 ant s82624 Camponotus festinus Smith 1857 ant s82602 Camponotus microthoracus Zhang 1989 ant s82620 Camponotus gracilis n. sp. Zhang 1989 ant s82606 Camponotus ampullosus n. sp. Zhang 1989 ant s82603 Camponotus ambon n. sp. Zhang 1989 ant s82621, s82622 Camponotus shanwangensis Hong 1984 ant s82601, s82604, s82605 Alloiomma changweiensis n. sp. Zhang 1989 ant s82629 Elaphrodites scutulatus n. gen. n. sp. Zhang 1989 ant s82636 Dolichoderus evolans n. sp. Zhang 1989 ant s82625 Dolichoderus lacinius n. sp. Zhang 1989 ant s82627 Technomyrmex septentrionalis n. sp. Zhang 1989 ant s82626 Myopopone sinensis n. sp. Zhang 1989 ant s82628
Hymenoptera - Megachilidae	Megachile shanwangae n. sp. Megachile shanwangae n. sp. Zhang 1989 mason bee s82638
Hymenoptera - Melittidae	"Dasypoda basaltica n. sp." = Macropis basaltica "Dasypoda basaltica n. sp." = Macropis basaltica Zhang 1989 bee s82637
Hymenoptera - Apidae	Bombus luianus n. sp.², Apis longitibia n. sp.², Xylocopa diatoma n. sp.² Bombus luianus n. sp.² Zhang 1990 bumble bee s82771 Apis longitibia n. sp.² Zhang 1990 long-legged honey bee s82773 Xylocopa diatoma n. sp.² Zhang 1990 large carpenter bee s82772
Hymenoptera - Tenthredinidae	Selandria perelegans n. sp. Selandria perelegans n. sp. Zhang 1989 wood wasp s82715
Coleoptera - Carabidae	Calosoma cf. maderae, Bembidion strenum n. sp. Calosoma cf. maderae Fabricius 1775 ground beetle s82751 Bembidion strenum n. sp. Zhang 1989 ground beetle s82706
Coleoptera - Dytiscidae	Laccophilus corystes n. sp., Laccophilus pusulatus n. sp. Laccophilus corystes n. sp. Zhang 1989 predaceous diving beetle s82711 Laccophilus pusulatus n. sp. Zhang 1989 predaceous diving beetle s82721
Coleoptera - Hydrophilidae	Helochares mydaleus n. sp., "Hydrous pergrandis n. sp." = Hydrophilus pergrandis, Enochrus lapsus n. sp. Helochares mydaleus n. sp. Zhang 1989 water scavenger beetle s82720 "Hydrous pergrandis n. sp." = Hydrophilus pergrandis Zhang 1989 water scavenger beetle s82719 Enochrus lapsus n. sp. Zhang 1989 water scavenger beetle s82714
Coleoptera -	Scelocopris enertheus n. gen. n. sp. Scelocopris enertheus n. gen. n. sp. Zhang 1989 scarab beetle s82754
Coleoptera - Scarabaeidae	Anomala sp., Anomala scia n. sp., Anomala furva, Onthophagus sp., ? Phalops sp. Anomala sp. Samouelle 1819 scarab beetle s82762, s82763 Anomala scia n. sp. Zhang 1989 scarab beetle s82710 Anomala furva Hong 1984 scarab beetle s82755 Onthophagus sp. Latreille 1802 scarab beetle s82761 ? Phalops sp. Erichson 1848 scarab beetle s82760
Coleoptera - Staphylinidae	Lathrobium gensium n. sp., Medon stenus n. sp. Lathrobium gensium n. sp. Zhang 1989 rove beetle s82698 Medon stenus n. sp. Zhang 1989 rove beetle s82699
Coleoptera - Cleridae	Cleridae indet. Cleridae indet. Latreille 1802 checkered beetle s82748
Coleoptera - Tenebrionidae	Dascillus exiguus n. sp. Dascillus exiguus n. sp. Zhang 1989 comb-clawed beetle s82731
Coleoptera - Endomychidae	"Engonius alviolatus" = Gerstaeckerus alviolatus "Engonius alviolatus" = Gerstaeckerus alviolatus Hong 1984 handsome fungus beetle s82743, s82744, 82745, s82746
Coleoptera - Coccinellidae	Scymnus (Pullus) aff. kawamurai Scymnus (Pullus) aff. kawamurai Ohta 1929 ladybird beetle s82722
Coleoptera - Attelabidae	Oedeuops ephemera n. gen. n. sp. Oedeuops ephemera n. gen. n. sp. Zhang 1989 leaf rolling weevil s82689
Coleoptera - Curculionidae	Hylobius plenus, Hylobius turfaceus n. sp. Hylobius plenus Zhang 1989 snout beetle s82690 Hylobius turfaceus n. sp. Zhang 1989 snout beetle s82691
Coleoptera - Brentidae	"Balanobius crassipedalis n. sp." = Longifistulia crassipedalis, "Balanobius edentis n. sp." = Longifistulia edentis "Balanobius crassipedalis n. sp." = Longifistulia crassipedalis Zhang 1989 weevil s82688 "Balanobius edentis n. sp." = Longifistulia edentis Zhang 1989 weevil s82683, s82684, s82685, s82687
Coleoptera - Chrysomelidae	Chrysomelidae indet., Chrysothoracus tropicus n. gen. n. sp. Chrysomelidae indet. Latreille 1802 leaf beetle s82749 Chrysothoracus tropicus n. gen. n. sp. Zhang 1989 leaf beetle s82701
Coleoptera - Cerambycidae	Delostromatium curvum n. gen. n. sp., Stromatium cf. barbatum, Trosites foveolatus n. gen. n. sp., Alifasciola sirdena n. gen. n. sp., Protaxis chinensis n. sp., Mesosa viaria n. sp. Delostromatium curvum n. gen. n. sp. Zhang 1989 longhorned beetle s82705 Stromatium cf. barbatum Fabricius 1775 longhorned beetle s82708 Trosites foveolatus n. gen. n. sp. Zhang 1989 longhorned beetle s82695 Alifasciola sirdena n. gen. n. sp. Zhang 1989 longhorned beetle s82693 Protaxis chinensis n. sp. Zhang 1989 longhorned beetle s82694 Mesosa viaria n. sp. Zhang 1989 longhorned beetle s82709
Coleoptera - Dascillidae	Dascillus sp., Dascillus flatus n. sp., Dascillus shandongianus Dascillus sp. Latreille 1796 soft bodied plant beetle s82750 Dascillus flatus n. sp. Zhang 1989 soft bodied plant beetle s82734, s82735, s82736 Dascillus shandongianus Zhang 1989 soft bodied plant beetle s82732, s82733, s82740, s82741, s82742
Coleoptera - Elateridae	Elater mitrus n. sp., Elater asmodeus n. sp. Elater mitrus n. sp. Zhang 1989 click beetle s82713 Elater asmodeus n. sp. Zhang 1989 click beetle s82747
Coleoptera - Cantharidae	"Curticantharis sp." = Themus "Curticantharis sp." = Themus Motschulsky 1858 soldier beetle s82753

¹Zhang 1989; ²Zhang 1990; ³Zhang et al. 1994