Haqel (MSNM collection) (Cretaceous of Lebanon)

Also known as Hakel, Haqil

Where: Lebanon (34.2° N, 35.8° E: paleocoordinates 13.0° N, 30.4° E)

• coordinate estimated from map

• local area-level geographic resolution

When: Metoicoceras geslinianum ammonoid zone, Late/Upper Cenomanian (99.6 - 93.5 Ma)

• The precise age of Hâqel and Hjoûla varies according to different authors. ln his comprehensive sedimentological study Hückel (1970) divided the Cenomanian Limestones into seven units and concluded that Hâqel (unite V) and Hjoûla (unite IV) were deposited during the later Lower Cenomanian. Owing to the occurrence of the ammonite Allocrioceras annulatum as a member of the Metoicoceras geslinianum Zone, Wippich & Lehmann (2004) confirmed the conclusions of Hemleben (1977) and dated Hâqel and Hjoûla as Upper Cenomanian (Fig. 2). Dalla Vecchia et al. (2002) considered Hâqel and Hjoûla as lower Upper Cenomanian, because Nammoûra (another fossil-rich locality a few kilometres in the South of Hjoûla and slightly younger than Hâqel and Hjoûla) is dated as Middle Cenomanian.

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal or restricted shallow subtidal; carbonate

Size class: macrofossils

Preservation: soft parts

Collection methods: quarrying, chemical, mechanical,

• Repository: Museo Civico di Storia Naturale di Milano (MSNM)

Primary reference: O. Rieppel and J.J. Head. 2004. New specimens of the fossil snake genus Eupodophis Rage and Escuillié, from the mid-Cretaceous of Lebanon. Memorie della Società Italiana di Scienze Naturali e Museo Civico di Storia Naturale di Milano 23(2):1-26 [J. Head/J. Head/J. Head]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 60311: authorized by Jason Head, entered by Jason Head on 09.05.2006, edited by Matthew Clapham and Richard Butler

Creative Commons license: CC BY (attribution)

Taxonomic list

Cephalopoda
 Octopoda - Trachyteuthididae
Trachyteuthis bacchiai n. sp.4 Fuchs and Larson 2011 octopus
Glyphiteuthis sp.4 Reuss 1854 octopus
Glyphiteuthis abisaadiorum in Fuchs & Weis 2009
Glyphiteuthis libanotica2 Fraas 1878 octopus
Glyphiteuthis freijii4 Fuchs and Larson 2011 octopus
Part of Glyphiteuthis libanotica in Fuchs, 2006
 Octopoda - Octopodidae
Styletoctopus annae n. gen. n. sp.3
Styletoctopus annae n. gen. n. sp.3 Fuchs et al. 2009 octopus
Xiphosura
 Xiphosurida - Limulidae
Tachypleus syriacus7 Woodward 1879 horseshoe crab
Malacostraca
 Decapoda - Oplophoridae
Odontochelion cretaceum5 Roger 1946 shrimp
 Decapoda - Stenopodidae
Phoenice pasinii n. gen. n. sp.6
Phoenice pasinii n. gen. n. sp.6 Garassino 2001 decapod
 Decapoda - Palinuridae
Palinurus sp.5 Weber 1795 spiny lobster
Linuparus sp.5 White 1847 spiny lobster
 Decapoda - Scyllaridae
Palibacus praecursor5 Dames 1886 decapod
 Decapoda - Erymidae
"Eryma cretacea" = Eryma cretaceum5
"Eryma cretacea" = Eryma cretaceum5 Roger 1946 decapod
 Decapoda - Nephropidae
Homarus hakelensis5 Fraas 1878 clawed lobster
 Decapoda - Dorippidae
Telamonocarcinus gambalatus8 Larghi 2004 crab
 Decapoda - Carpopenaeidae
Carpopenaeus septemspinatus5, "Carpopenaeus callirostris" = Carpopenaeus callirostris5, "Pseudastacus dubertreti" = Carpopenaeus callirostris5
Carpopenaeus septemspinatus5 Dames 1886 decapod
"Carpopenaeus callirostris" = Carpopenaeus callirostris5 Glaessner 1946 decapod
"Pseudastacus dubertreti" = Carpopenaeus callirostris5 Glaessner 1946 decapod
 Decapoda - Aegeridae
Acanthochirana smithwoodwardi5 Van Straelen 1940 decapod
Aeger libanensis5 Roger 1946 decapod
 Decapoda - Penaeidae
Microchela rostrata n. sp.5 Garassino 1994 decapod
Libanocaris rogeri n. sp.5 Garassino 1994 decapod
Hakelocaris vavassorii n. sp.5 Garassino 1994 decapod
Carinacaris teruzzii n. sp.5 Garassino 1994 decapod
Macropenaeus incertus5 Roger 1946 decapod
Reptilia
 Squamata - Pachyophidae
Eupodophis descouensi Rage and Escuillié 2000 snake
 Pterosauria -
Ornithocheiroidea indet.1 pterosaur
MSNM V 3881, partial left forelimb
Actinopteri
 Neopterygii - Gebrayelichthyidae
Gebrayelichthys uyenoi9 Nursall and Capasso 2004