PBDB Collection

Tuanwang, Laiyang City (Shandong Prov. Museum coll.) (Cretaceous of China)

Where: Shandong, China (36.8° N, 120.6° E: paleocoordinates 40.2° N, 122.1° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Laiyang Formation, Aptian (125.0 - 113.0 Ma)

Environment/lithology: lacustrine - large; lithified shale

• Large (several hundred km2) lake in the Laiyang basin. In addition to the abundant entomofauna, plant fossil fragments, fish and a single bird feather also have been found. Water plants were apparently rare, and at the centre of the lake I have found no plant or animal fossils. Extensional regime in response to the progressive attenuation of the thickened lithosphere and orogenic collapse.

• The Third Member of the Laiyang Formation consists of 250-705 m of calcareous sandstone, siltstone and fine-granted sandstone intercalated with thin, paper-like, insect-rich shales.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collection methods: Repository: Shandong Provincial Museum

Primary reference: J. F. Zhang. 1992. Fossil Coleoptera from Laiyang, Shandong Province. Acta Entomologica Sinica 35:331-338 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 125196: authorized by Matthew Clapham, entered by Matthew Clapham on 06.03.2012

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Hymenoptera -	Shandongodes necrodes n. sp.¹ Shandongodes necrodes n. sp.¹ Zhang 1985 winged insect L85111
Hymenoptera - Tenthredinidae	Palaeathalia laiyangensis n. gen. n. sp.¹ Palaeathalia laiyangensis n. gen. n. sp.¹ Zhang 1985 wood wasp L85121
Hymenoptera -	"Oligoneuroides huadongensis n. gen. n. sp." = Rasnitsynia huadongensis¹ "Oligoneuroides huadongensis n. gen. n. sp." = Rasnitsynia huadongensis¹ Zhang 1985 wasp L85115
Hymenoptera - Angarosphecidae	"Shandongodes lithodes n. gen. n. sp." = Angarosphex lithodes¹, "Mataeosphex venulosus n. gen. n. sp." = Angarosphex venulosus¹ "Shandongodes lithodes n. gen. n. sp." = Angarosphex lithodes¹ Zhang 1985 apoid wasp L85110 "Mataeosphex venulosus n. gen. n. sp." = Angarosphex venulosus¹ Zhang 1985 apoid wasp L85112-1
Hymenoptera - Megalyridae	Stemmogaster celata n. gen. n. sp.¹ Stemmogaster celata n. gen. n. sp.¹ Zhang 1985 wasp L85113
Trichoptera -	Tuanwangica aethoneura n. gen. n. sp.¹ Tuanwangica aethoneura n. gen. n. sp.¹ Zhang 1985 caddisfly L85106
Coleoptera - Coptoclavidae	Coptoclavisca grandioculus n. sp. Coptoclavisca grandioculus n. sp. Zhang 1992 beetle L85433
Hemiptera - Saldidae	Mesolygaeus laiyangensis⁴ Mesolygaeus laiyangensis⁴ Ping 1928 shore bug L88232, 88240, 88242
Hemiptera - Archegocimicidae	Schizopteryx lacustris⁴, Schizopteryx shandongensis⁴ Schizopteryx lacustris⁴ Zhang 1991 shore bug L88257 Schizopteryx shandongensis⁴ Hong 1984 shore bug L88245, 88246, 88247, 88248, 88249, 88253
Hemiptera - Palaeoaphididae	Caudaphis spinalis n. gen. n. sp.⁶, Caudaphis leptoneura n. sp.⁶ Caudaphis spinalis n. gen. n. sp.⁶ Zhang et al. 1989 aphid L84116 Caudaphis leptoneura n. sp.⁶ Zhang et al. 1989 aphid L84117
Hemiptera - Sinaphididae	Tartaraphis peregrina n. gen. n. sp.⁶ Tartaraphis peregrina n. gen. n. sp.⁶ Zhang et al. 1989 aphid L84124(5)
Hemiptera - Oviparosiphidae	"Paroviparosiphum opimum n. gen. n. sp." = Archeoviparosiphum opimum⁶, "Paroviparosiphum camptotropum n. sp." = Archeoviparosiphum camptotropum⁶, "Mesoviparosiphum malacum n. sp." = Archeoviparosiphum malacum⁶, "Mesoviparosiphum tuanwangense n. gen. n. sp." = Archeoviparosiphum tuanwangense⁶ "Paroviparosiphum opimum n. gen. n. sp." = Archeoviparosiphum opimum⁶ Zhang et al. 1989 aphid L84126 "Paroviparosiphum camptotropum n. sp." = Archeoviparosiphum camptotropum⁶ Zhang et al. 1989 aphid L84129 "Mesoviparosiphum malacum n. sp." = Archeoviparosiphum malacum⁶ Zhang et al. 1989 aphid L84122 "Mesoviparosiphum tuanwangense n. gen. n. sp." = Archeoviparosiphum tuanwangense⁶ Zhang et al. 1989 aphid L84121
Hemiptera - Archijassidae	Mesoccus advenus n. sp.¹, Mesoccus lutarius n. gen. n. sp.¹ Mesoccus advenus n. sp.¹ Zhang 1985 true bug L85103 Mesoccus lutarius n. gen. n. sp.¹ Zhang 1985 true bug L85101, 85104
Orthoptera - Locustopsidae	Pseudoacrida cf. costata¹ Pseudoacrida cf. costata¹ Lin 1982 grasshopper L84985(84988)
Orthoptera -	Falsirameus ravus n. gen. n. sp.¹ Falsirameus ravus n. gen. n. sp.¹ Zhang 1985 grasshopper L84991(2)
Diptera - Chaoboridae	Mesochaoborus pallens n. sp.⁵, Mesochaoborus zhangshanyingensis⁵, "Petiolatendipes shouchangensis" = Chironomaptera vesca⁵ Mesochaoborus pallens n. sp.⁵ Zhang et al. 1986 phantom midge L84113 Mesochaoborus zhangshanyingensis⁵ Hong 1984 phantom midge L84112 "Petiolatendipes shouchangensis" = Chironomaptera vesca⁵ Kalugina 1980 phantom midge L84104, L84114
Diptera - Rhagionidae	Pseudopetia exilis n. sp.² Pseudopetia exilis n. sp.² Zhang 1987 snipe fly L84135
Diptera - Platypezidae	Mesopetia tuanwangensis n. gen. n. sp.² Mesopetia tuanwangensis n. gen. n. sp.² Zhang 1987 flat-footed fly L84131
Diptera - Pediciidae	"Palaeolimnobia laiyangensis n. gen. n. sp." = Zhangobia laiyangensis⁵ "Palaeolimnobia laiyangensis n. gen. n. sp." = Zhangobia laiyangensis⁵ Zhang et al. 1986 hairy-eyed cranefly L84101(2)
Diptera - Chironomidae	Oryctochlus contiguus n. sp.³ Oryctochlus contiguus n. sp.³ Zhang 1991 non biting midge L86812, 86813

¹Zhang 1985; ²Zhang 1987; ³Zhang 1991; ⁴Zhang 1991; ⁵Zhang et al. 1986; ⁶Zhang et al. 1989