PBDB

Klein Lehmhagen pit, Grimmen (LGA collection) (Jurassic of Germany)

Where: Mecklenburg-Vorpommern, Germany (54.1° N, 13.1° E: paleocoordinates 43.7° N, 18.9° E)

• coordinate based on political unit

When: Dactylioceras tenuicostatum zone, Early/Lower Toarcian (183.0 - 182.0 Ma)

• unnamed "bituminous shale bed" equivalent to Posidonian Shale (Posidonienschiefer); Lias epsilon level = lowest Toarcian; also lower part of Harpoceras falcifer Zone

• bed-level stratigraphic resolution

Environment/lithology: offshore; lithified, carbonaceous shale

• bituminous shale ("bituminösen Schiefer")

Size classes: macrofossils, mesofossils

Preservation: adpression, concretion

Collected by Ansorge; reposited in the MfN

Primary reference: W. Zessin. 1987. Variabilität, Merkmalswandel und Phylogenie der Elcanidae im Jungpaläozoikum und Mesozoikum und die Phylogenie der Ensifera. Deutsche Entomologische Zeitschrift 34(1-3):1-76 [M. Clapham/J. Karr/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 114993: authorized by Matthew Clapham, entered by Jered Karr on 26.08.2011, edited by Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Arachnida
Araneae -	Seppo koponeni n. gen. n. sp.¹¹ Seppo koponeni n. gen. n. sp.¹¹ Selden and Dunlop 2014 spider MB.A 2966
Insecta
Eoblattida - Blattogryllidae	Griphopteron molle² Griphopteron molle² Handlirsch 1939 insect LGA 850
Hemiptera - Archijassidae	Archijassidae indet.², Mesoledra pachyneura², Archijassus heeri², Ardela grimmenensis n. gen. n. sp.² Archijassidae indet.² Becker-Migdisova 1962 true bug LGA 240, 380, 384, 701 Mesoledra pachyneura² Handlirsch 1939 true bug LGA 138, 206, 742, 1050, 1086, 1299, 927 Archijassus heeri² Geinitz 1880 true bug LGA 505, 640, 757, 1204, 1288, 1340, 1300, 108 Ardela grimmenensis n. gen. n. sp.² Ansorge 1996 true bug LGA 1348, 673, 856, 1104, 1216, 505, 350, 1325, 163, 266
Hemiptera - Hylicellidae	Liassocercopis schnicki n. gen. n. sp.² Liassocercopis schnicki n. gen. n. sp.² Ansorge 1996 true bug LGA 814
Hemiptera - Procercopidae	Procercopidae indet.², Procercopina frenzeli n. sp.² Procercopidae indet.² Handlirsch 1906 froghopper LGA 676 Procercopina frenzeli n. sp.² Ansorge 1996 froghopper LGA 565, 380
Hemiptera - Protopsyllidiidae	Aphidulum ciliatum n. sp.², Toarcopsyllidium ernsti n. gen. n. sp.² Aphidulum ciliatum n. sp.² Ansorge 1996 true bug LGA 264, 641, 931 Toarcopsyllidium ernsti n. gen. n. sp.² Ansorge 1996 true bug LGA 1185
Hemiptera - Liadopsyllidae	Liadopsylla obtusa n. sp.² Liadopsylla obtusa n. sp.² Ansorge 1996 jumping plant louse LGA 731
Hemiptera - Oviparosiphidae	Grimmenaphis magnifica n. gen. n. sp.² Grimmenaphis magnifica n. gen. n. sp.² Ansorge 1996 aphid LGA 1313, 903
Hemiptera - Archiconiopterygidae	Archiconiopteryx liasina² Archiconiopteryx liasina² Handlirsch 1906 true bug LGA 620, 666, 747
Hemiptera - Fulgoridiidae	Fulgoridium "sp 1"², Fulgoridium "sp 2"², Fulgoridium sp.² Fulgoridium "sp 1"² Handlirsch 1906 planthopper LGA 820 Fulgoridium "sp 2"² Handlirsch 1906 planthopper LGA 123 Fulgoridium sp.² Handlirsch 1906 planthopper LGA 569, 749, 1183
Hemiptera -	Fulgoridulum egens², Heteroptera indet.⁵ Fulgoridulum egens² Handlirsch 1939 planthopper LGA 70, 89, 198, 714, 1313, 876, 1206, 628, 893, 1349, 920, 899, 896, 893, 1303, 920, 896, 1313 Heteroptera indet.⁵ Latreille 1810 true bug LGA 936
Lophioneurida - Lophioneuridae	Undacypha europaea n. sp.² Undacypha europaea n. sp.² Ansorge 1996 winged insect LGA 233, 223, 292, 378, 490, 645, 746, 871, 883, 967, 983, 1274, 1333, 1342, 1339, 1364, 1365
Permopsocida - Archipsyllidae	Archipsylla primitiva² Archipsylla primitiva² Handlirsch 1906 winged insect LGA 1229
Permopsocida - Psocidiidae	Liassopsocus lanceolatus n. gen. n. sp.² Liassopsocus lanceolatus n. gen. n. sp.² Ansorge 1996 winged insect LGA 1246, 997
Hymenoptera - Karatavitidae	Grimmaratavites mirabilis n. gen. n. sp.¹⁰ Grimmaratavites mirabilis n. gen. n. sp.¹⁰ Rasnitsyn et al. 2006 wood wasp LGA 2510
Lepidoptera - Eolepidopterigidae	? Eolepidopterigidae indet.¹⁵, ? Eolepidopterigidae indet.¹⁵, ? Eolepidopterigidae indet.¹⁵ ? Eolepidopterigidae indet.¹⁵ Rasnitsyn 1983 moth LGA 2150 ? Eolepidopterigidae indet.¹⁵ Rasnitsyn 1983 moth LGA 968 ? Eolepidopterigidae indet.¹⁵ Rasnitsyn 1983 moth LGA 1500
Trichoptera - Philopotamidae	Liadotaulius maior⁴ Liadotaulius maior⁴ Handlirsch 1906 finger-net caddisfly LGA 290, 458, 626, 649, 1995, 2040, 2188, 220, 1198, 1710, 672, 713, 934
Trichoptera - Necrotauliidae	Mesotrichopteridium intermedium⁴, Necrotaulius parvulus⁴ Mesotrichopteridium intermedium⁴ Handlirsch 1906 caddisfly LGA 348, 533, 675, 789, 830, 874, 784, 1425 Necrotaulius parvulus⁴ Geinitz 1884 caddisfly
Diptera -	Asilomorpha "sp 1"², Asilomorpha "sp 2"² Asilomorpha "sp 1"² Rohdendorf 1961 fly LGA 690 Asilomorpha "sp 2"² Rohdendorf 1961 fly LGA 429
Diptera - Rhagionidae	Palaeobrachyceron willmanni n. sp.², Grimmyia baltica n. gen. n. sp.² Palaeobrachyceron willmanni n. sp.² Ansorge 1996 snipe fly LGA 269 Grimmyia baltica n. gen. n. sp.² Ansorge 1996 snipe fly LGA 950
Diptera - Heterorhyphidae	Heterorhyphus triangularis n. sp.² Heterorhyphus triangularis n. sp.² Ansorge 1996 fly LGA 380
Diptera - Protorhyphidae	Protorhyphus simplex², Archirhyphus geinitzi² Protorhyphus simplex² Geinitz 1888 fly LGA 533, 571, 744, 998, 1002, 1007, 1294 Archirhyphus geinitzi² Handlirsch 1939 fly LGA 999
Diptera - Pleciofungivoridae	Pleciofungivora liassica n. sp.² Pleciofungivora liassica n. sp.² Ansorge 1996 fly LGA 1045
Diptera - Antefungivoridae	Antefungivora germanica n. sp.², Archibio mycetophilinus² Antefungivora germanica n. sp.² Ansorge 1996 fly LGA 394 Archibio mycetophilinus² Handlirsch 1939 fly LGA 214, 840, 988
Diptera - Anisopodidae	Mesorhyphus zirzipanus n. sp.² Mesorhyphus zirzipanus n. sp.² Ansorge 1996 wood gnat LGA 579
Diptera - Chaoboridae	Rhaetomyia herrigi n. sp.² Rhaetomyia herrigi n. sp.² Ansorge 1996 phantom midge LGA 1189
Diptera - Chironomidae	Podonominae indet.², ? Podonomius tumidus n. sp.², ? Oryctochlus toarciensis n. sp.², Aenne liasina n. gen. n. sp.³ Podonominae indet.² Thienemann and Edwards 1837 non biting midge LGA 1003 ? Podonomius tumidus n. sp.² Ansorge 1996 non biting midge LGA 1034 ? Oryctochlus toarciensis n. sp.² Ansorge 1996 non biting midge LGA 835 Aenne liasina n. gen. n. sp.³ Ansorge 1999 non biting midge LGA 1474, 1486, 1894, 1956
Diptera - Psychodidae	Tanypsycha connexa n. gen. n. sp.¹, Liassopsychodina pommerana n. gen. n. sp.¹ Tanypsycha connexa n. gen. n. sp.¹ Ansorge 1994 moth fly LGA 1233 Liassopsychodina pommerana n. gen. n. sp.¹ Ansorge 1994 moth fly LGA 457, 752
Diptera - Protopleciidae	Protoplecia klafacki n. sp.², Protoplecia liasina² Protoplecia klafacki n. sp.² Ansorge 1996 fly LGA 892, 924 Protoplecia liasina² Geinitz 1884 fly LGA 1180
Diptera - Eoditomyidae	Eoditomyia primitiva n. gen. n. sp.² Eoditomyia primitiva n. gen. n. sp.² Ansorge 1996 fly LGA 1305, 836
Diptera -	Archipleciomima sp.², Archipleciomima germanica n. sp.⁸ Archipleciomima sp.² Rohdendorf 1962 fly LGA 1203 Archipleciomima germanica n. sp.⁸ Krzemiński and Ansorge 2018 fly LGA 1550, 1803, 2282
Diptera - Trichoceridae	Trichoceridae indet.⁹, Mailotrichocera mikereichi⁹, Mailotrichocera variabilis⁹, Mailotrichocera zessini n. sp.⁹ Trichoceridae indet.⁹ Kertész 1902 winter crane fly LGA 1900 Mailotrichocera mikereichi⁹ Krzemińska et al. 2009 winter crane fly LGA 79, 2005 Mailotrichocera variabilis⁹ Krzemińska et al. 2009 winter crane fly LGA 288, 681, 1100, 1163, 2309 Mailotrichocera zessini n. sp.⁹ Krzemińska et al. 2009 winter crane fly LGA 611
Diptera - Ptychopteridae	"Eoptychoptera liasina" = Eoptychoptera simplex² "Eoptychoptera liasina" = Eoptychoptera simplex² Handlirsch 1906 phantom crane fly LGA 1368
Diptera - Tanyderidae	Nannotanyderus grimmenensis n. sp.⁶, Nannotanyderus krzeminskii n. gen. n. sp.¹ Nannotanyderus grimmenensis n. sp.⁶ Ansorge and Krzemiński 2002 primitive crane fly LGA 2222 Nannotanyderus krzeminskii n. gen. n. sp.¹ Ansorge 1994 primitive crane fly LGA 1145, 544
Mecoptera - Orthophlebiidae	Orthophlebia germanica², Orthophlebia brunsvicensis², Orthophlebia cf. elongata² Orthophlebia germanica² Handlirsch 1906 scorpionfly LGA 811, 1362 Orthophlebia brunsvicensis² Bode 1905 scorpionfly LGA 790 Orthophlebia cf. elongata² Handlirsch 1939 scorpionfly LGA 737
Mecoptera - Bittacidae	Bittacidae indet.², Neorthophlebia maculipennis², Mesobittacus minutus² Bittacidae indet.² Handlirsch 1906 hangingfly LGA 500 Neorthophlebia maculipennis² Handlirsch 1906 hangingfly LGA 1369 Mesobittacus minutus² Handlirsch 1939 hangingfly LGA 133, 876, 1269, 1298
Mecoptera - Cimbrophlebiidae	Cimbrophlebiidae indet.⁵ Cimbrophlebiidae indet.⁵ Willmann 1977 hangingfly LGA 1608
Mecoptera - Pseudopolycentropodidae	Pseudopolycentropus sp.², Pseudopolycentropus triangularis² Pseudopolycentropus sp.² Handlirsch 1906 scorpionfly LGA 769, 993 Pseudopolycentropus triangularis² Handlirsch 1920 scorpionfly LGA 636, 925, 20, 738, 795, 1151
Neuroptera - Prohemerobiidae	Prohemerobius latus², "Sibithone prodroma" = Prohemerobius prodromus², Actinophlebia sp.² Prohemerobius latus² Handlirsch 1920 lacewing LGA 124 "Sibithone prodroma" = Prohemerobius prodromus² Handlirsch 1906 lacewing LGA 1001, 607, 452 Actinophlebia sp.² Handlirsch 1906 lacewing LGA 1080, 324
Neuroptera - Ithonidae	Polyosmylus excelsus n. gen. n. sp.² Polyosmylus excelsus n. gen. n. sp.² Ansorge 1996 moth lacewing LGA 1033, 540
Neuroptera - Osmylidae	"Mesosmylina falciferum n. sp." = Mesosmylina falcifera² "Mesosmylina falciferum n. sp." = Mesosmylina falcifera² Ansorge 1996 lance lacewing LGA 1295
Dicondylia -	Coleoptera indet.⁵ Coleoptera indet.⁵ Linnaeus 1758 beetle LGA 1230
Coleoptera - Trachypachidae	Liassodites obsti n. gen. n. sp.⁷ Liassodites obsti n. gen. n. sp.⁷ Kirejtshuk and Ansorge 2022 false ground beetle LGA 2850
Orthoptera - Elcanidae	Parelcana tenuis², "Elcana media" = Panorpidium media, "Elcana magna" = Panorpidium magna, "Elcana geinitzi" = Panorpidium geinitzi², "Elcana minima" = Panorpidium minima Parelcana tenuis² Handlirsch 1906 grasshopper LGA 487, 1351 "Elcana media" = Panorpidium media Handlirsch 1906 grasshopper LGA 4/1/2, 58/1/2, 63, 402/1/2 "Elcana magna" = Panorpidium magna Handlirsch 1906 grasshopper LGA 34/1/2, 62 "Elcana geinitzi" = Panorpidium geinitzi² Heer 1880 grasshopper LGA 445 "Elcana minima" = Panorpidium minima Handlirsch 1906 grasshopper LGA 61/1/2
Orthoptera - Protogryllidae	Protogryllus sp.², "Protogryllus dobbertinensis" = Protogryllus (Protogryllus) dobbertinensis² Protogryllus sp.² Handlirsch 1906 grasshopper LGA 32, 51, 358, 406, 511, 604, 766, 887, 889, 1040, 1172, 1196, 1238 "Protogryllus dobbertinensis" = Protogryllus (Protogryllus) dobbertinensis² Geinitz 1880 grasshopper LGA 94, 244, 324, 395, 813, 818, 865, 932, 1113, 1136, 1267
Orthoptera - Regiatidae	Protochaeta masculina², Protochaeta lanceolata² Protochaeta masculina² Handlirsch 1939 grasshopper LGA 113, 304, 324, 497, 597, 686, 780, 937, 940, 1114, 1280 Protochaeta lanceolata² Handlirsch 1939 grasshopper LGA 235
Orthoptera - Locustopsidae	Locustopsis " sp 1"², Locustopsis "sp 2"², Locustopsis "sp 3"², Locustopsis gyra², "Locustopsis bernstorffi" = Plesioschwinzia bernstorffi², "Locustopsis kruegeri" = Plesioschwinzia kruegeri¹³ Locustopsis " sp 1"² Handlirsch 1906 grasshopper LGA 1272 Locustopsis "sp 2"² Handlirsch 1906 grasshopper LGA 1144 Locustopsis "sp 3"² Handlirsch 1906 grasshopper LGA 1309 Locustopsis gyra² Zessin 1983 grasshopper LGA 390 "Locustopsis bernstorffi" = Plesioschwinzia bernstorffi² Geinitz 1880 grasshopper LGA 100, 296 "Locustopsis kruegeri" = Plesioschwinzia kruegeri¹³ Zessin 1983 grasshopper LGA 100
Blattodea - Blattulidae	Blattula dubia¹², Blattula langfeldti¹² Blattula dubia¹² Handlirsch 1939 cockroach LGA 1725 Blattula langfeldti¹² Geinitz 1880 cockroach LGA 86, 96, 27, 331/1(2), 380, 425, 579, 594, 888, 893, 1077, 1201, 1275, 1372, 1529, 1578, 1739, 1820, 2169, 13, 118, 154, 880, 1328, 1435, 1519, 1842, 1887, 1926, 2006, 2015, 2196, 2356, 888, 1820
Blattodea - Mesoblattinidae	Mesoblattina protypa¹² Mesoblattina protypa¹² Geinitz 1880 cockroach LGA 1320, 1942
Grylloblattodea - Bajanzhargalanidae	Nele jurassica n. gen. n. sp.² Nele jurassica n. gen. n. sp.² Ansorge 1996 ice crawler LGA 1310, 1111, 211, 977
Dicondylia - Aerophasmidae	Durnovaria cf. parallela² Durnovaria cf. parallela² Whalley 1985 winged insect LGA 1086, 1323
Plecoptera - Perlariopseidae	Perlariopseidae indet.², Dicronemoura furcata n. sp.² Perlariopseidae indet.² Sinitshenkova 1985 stonefly LGA 777 Dicronemoura furcata n. sp.² Ansorge 1996 stonefly LGA 1278
Odonata - Sphenophlebiidae	Sphenophlebia pommerana² Sphenophlebia pommerana² Ansorge 1996 dragonfly LGA 480, 510, 1197
Odonata - Asiopteridae	"Oreopteridae indet." = Asiopteridae² "Oreopteridae indet." = Asiopteridae² Pritykina 1968 dragonfly LGA 250, 712, 1358
Odonata - Liassogomphidae	Liassogomphidae "indet. 1"², Liassogomphidae "indet. 2"² Liassogomphidae "indet. 1"² Tillyard 1935 dragonfly LGA 922 Liassogomphidae "indet. 2"² Tillyard 1935 dragonfly LGA 349
Odonata - Liassophlebiidae	Grimmenopteron elegantulum n. gen. n. sp.² Grimmenopteron elegantulum n. gen. n. sp.² Ansorge 1996 damsel-dragonflies LGA 345
Odonata - Heterophlebiidae	Trigonophlebia zessini n. gen. n. sp.², Heterophlebia buckmani² Trigonophlebia zessini n. gen. n. sp.² Ansorge 1996 dragonfly LGA 739 Heterophlebia buckmani² Brodie 1845 dragonfly LGA 46
Dicondylia - Protomyrmeleontidae	Protomyrmeleon brunonis², "Zirzipanagrion quadriordinum n. gen. n. sp." = Protomyrmeleon quadriordinum¹⁴, Obotritagrion tenuiformum² Protomyrmeleon brunonis² Geinitz 1888 winged insect LGA 621, 815, 988, 1000 "Zirzipanagrion quadriordinum n. gen. n. sp." = Protomyrmeleon quadriordinum¹⁴ Zessin 1991 winged insect LGA 162/1 Obotritagrion tenuiformum² Zessin 1991 winged insect LGA 876
Dicondylia -	Ephemeroptera indet.² Ephemeroptera indet.² Hyatt and Arms 1890 mayfly LGA 1502

¹Ansorge 1994; ²Ansorge 1996; ³Ansorge 1999; ⁴Ansorge 2002; ⁵Ansorge 2003; ⁶Ansorge and Krzemiński 2002; ⁷Kirejtshuk and Ansorge 2022; ⁸Kopeć et al. 2018; ⁹Krzemińska et al. 2009; ¹⁰Rasnitsyn et al. 2006; ¹¹Selden and Dunlop 2014; ¹²Vršanský and Ansorge 2007; ¹³Zessin 1988; ¹⁴Zessin 1991; ¹⁵Zhang et al. 2018