Apomarma (Miocene of Greece)

Where: Crete, Greece (35.1° N, 24.9° E: paleocoordinates 34.9° N, 24.7° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Varvara Formation, Messinian (7.2 - 5.3 Ma)

• Lower Messinian

Environment/lithology: basinal (siliceous); lithified, calcareous mudstone

• "A slumped horizon as well as two intercalated beds of laminated intraclast grainstone (calciturbidites), the lower with imported fragments of shallow water biota (faviid corals), point to a deeper position on an unstable slope. The ant fossil described in this study was discovered within a package of laminated calcareous mudstone in the upper part of the section (Fig. 2). Associated with the insect are leaves, seeds, and blossoms of terrestrial plants, partly articulated fish remains, siliceous sponge spicules, and small planktic foraminifers. Such laminated sediments document episodes when water exchange was interrupted in deeper parts of the basin and anoxic bottom water had formed in response to salinity stratification of the water column during periods of wet climate at times of precession minima (Ten Veen and Postma 1996)." Wappler et al., 2009 "This early supra-detachment basin dates back to 12–11 Ma and became increasingly fragmented in the course of the late Miocene by dominantly N-S–striking faults reflecting arc-parallel extension associated with ongoing outward motion of the Aegean arc."
• "The studied outcrop exposes a 5-m-thick succession of laminated and homogeneous calcareous mudstones of the Varvara Formation (Fig. 2). The high carbonate content in the sediments and their stratigraphic and spatial vicinity to gypsum deposits (Fig. 2) indicate an early Messinian age." Wappler et al., 2009

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Steinmann Institute for Geology, Mineralogy, Palaeontology, University of Bonn

Primary reference: T. Wappler, G. M. Dlussky, and M. Reuter. 2009. The first fossil record of Polyrhachis (Hymenoptera: Formicidae: Formicinae) from the Upper Miocene of Crete (Greece). Paläontologische Zeitschrift 83:431-438 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 133428: authorized by Matthew Clapham, entered by Jered Karr on 13.09.2012

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hymenoptera - Formicidae
Polyrhachis annosus n. sp. Wappler et al. 2009 ant
IPB-WAPPLER-1a,b