Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)
• coordinate based on nearby landmark
• local area-level geographic resolution
When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)
• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.
• group of beds-level stratigraphic resolution
Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone
Size classes: macrofossils, mesofossils
Preservation: adpression
Collection methods: Repository: Beijing Museum of Natural History. This collection contains specimens labeled with five-digit numbers starting with 79... or 80...
Primary reference: Y. C. Hong. 1984. Fossil insects in the diatoms of Shanwang. Bulletin of the Tianjin Institute of Geology and Mineral Resources 8:1-15 [D. Smith/E. Leckey/M. Clapham]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 128550: authorized by Matthew Clapham, entered by Matthew Clapham on 11.06.2012, edited by Erin Leckey
Creative Commons license: CC BY (attribution)
Taxonomic list
Arachnida | |
"Retina robusta" = Heteropoda robusta6
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"Eocryphoeca parva" = Tetragnatha parva6
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Insecta | |
Tipula insolita n. sp.5
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Sciara brevipetiola n. sp.5
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Plecia solaris n. sp.5, Plecia ludongensis n. sp.1, Clothonopsis miocenica n. gen. n. sp.4, Bibio ventricosus n. sp.5, Bibio expansus5, "Fungivora bullata n. sp." = Bibio bullatus5, Bibio brunneus n. sp.5, Bibio deformatus n. sp.5, Bibio shanwangensis1, Bibio ruptineurus n. sp.5, "Bibio miocenica n. sp." = Bibio miocenicus1, Bibio cratopodus5
Bibio ventricosus n. sp.5 Zhang 1989 march fly 79033 (Bibio shanwangensis in Hong, 1985 - pl 7, figs 1, 2, 3)
Bibio deformatus n. sp.5 Zhang 1989 march fly 79057 (Bibio shanwangensis in Hong, 1985 - pl 4, fig 1)
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Jagoria sinica n. sp.5
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Aphaena lithoecia n. sp.5, "Fulgoropsis fusca n. gen. n. sp." = Limois shanwangensis, "Oxycephala xiejiaheensis" = Limois shanwangensis5
"Oxycephala xiejiaheensis" = Limois shanwangensis5 Hong 1979 fulgorid planthopper 79026, 79079, 79080
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Shanwangicoris longa n. gen. n. sp.3, "Coriopsis miocenica n. sp." = Coreiopsis miocenica3, "Coriopsis parva n. sp." = Coreiopsis parva3, Coreiopsis punctata n. gen. n. sp., Gonocerus shandongensis n. sp.5, Cloresmus shanwangi n. sp.5
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Manocerus stagnans n. gen. n. sp.5
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Coroides xiejiaheense n. gen. n. sp.
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Tarsonecta mecopoda n. gen. n. sp.5
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"Oncotympana lapidescens n. sp." = Tanyocicada lapidescens5
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"Cinara reconditivenia n. sp." = Cinara (Cinara) reconditivenia5
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Apanechura asecta n. sp.5
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Luia lacaris n. gen. n. sp.5
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"Pseudoperla shanwangensis n. sp." = Eumenes shanwangensis
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"Brachyopteris shanwangensis n. sp." = Tiphia shanwangensis
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Elis pyrula n. sp.5
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Lasius vetulus5, Lasius validus n. sp.5, ? Lasius truncatus n. sp.5, Formica linquensis n. sp.5, Camponotus mitis5, Camponotus microthoracus n. sp.5, Camponotus longus n. sp.5, "Camponotus shanwangense" = Camponotus shanwangensis, Camponotus curviansatus n. sp.5, "Rabidia xiejiaheensis n. sp." = Oecophylla xiejiaheensis, "Liometopum jiaoyanshanense n. sp." = Dolichoderus jiaoyanshanensis5, Shanwangella palaeoptera n. gen. n. sp.5, Liometopum potamophilum n. sp.5, ? Tapinoma baculum n. sp.5, Paraphaenogaster shanwangensis n. sp., "Heteromyrmex atopogaster n. gen. n. sp." = Zhangidris atopogaster5
Lasius validus n. sp.5 Zhang 1989 ant 79121 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 4)
Formica linquensis n. sp.5 Zhang 1989 ant 79058 (Paraphaenogaster shanwangense in Hong, 1985 - pl 23, fig 4)
"Camponotus shanwangense" = Camponotus shanwangensis Hong 1984 ant 79031; 79043, 79076 (Liometopum jiaoyanshanense in Hong, 1985); ten specimens including 80179 in Hong & Wang 1985
Camponotus curviansatus n. sp.5 Zhang 1989 ant 79042 (Camponotus shanwangensis in Hong, 1985 - pl 27, fig 1)
"Liometopum jiaoyanshanense n. sp." = Dolichoderus jiaoyanshanensis5 Hong 1985 ant 79087; 80196 in Hong & Wang 1985
Shanwangella palaeoptera n. gen. n. sp.5 Zhang 1989 ant 79051 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 2)
Liometopum potamophilum n. sp.5 Zhang 1989 ant 79036 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 6)
"Heteromyrmex atopogaster n. gen. n. sp." = Zhangidris atopogaster5 Zhang 1989 ant 79054 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 3)
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Apis miocenica n. sp.
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"Sinostigma spinalata n. sp." = Pemphredon spinalatum
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Abia paurocephala n. sp.5, "Procimbex shanwangensis n. sp." = Abia shanwangensis, "Procimbex shandongense n. sp." = Abia shandongensis1
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"Megaxyela xiejiaheensis n. sp." = Xyelecia xiejiaheensis
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? Carabus ovalus n. sp.2, Calosoma cf. maderae5, Scarites mancus n. sp.5, "Shanwangicarabus furvus n. sp." = Chlaenius furvus2, "Shanwangicarabus paucinus n. sp." = Chlaenius furvus2
"Shanwangicarabus furvus n. sp." = Chlaenius furvus2 Hong 1984 ground beetle 80125, 80106, 80126, 480183
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Scarabaeoidea "indet. 1"5 Latreille 1802 scarab beetle 79045 (Magniscarabaeus furvus in Hong & Wang - pl 17, fig 2)
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Geotrupidae indet.5 Latreille 1802 earth-boring dung beetle 79107 (Magniscarabaeus brunneus in Hong, 1985 - pl 20, fig 3)
"Obitiscarabaeus jiaoyanshanense n. gen. n. sp." = Geotrupes jiaoyanshanense Hong 1984 earth-boring dung beetle 79114/5
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Anomala sp.5, "Magniscarabaeus brunneus n. gen. n. sp." = Anomala palaeobrunnea5, Anomala synemosyna n. sp.5, Anomala lochmocola n. sp.5, "Magniscarabaeus furvus" = Anomala furva2, Holotrichia spatha n. sp.5
"Magniscarabaeus brunneus n. gen. n. sp." = Anomala palaeobrunnea5 Krell 2000 scarab beetle 79105; seven including 80101, 80120, 80172, 80188, 80184, 80194, mentioned in Hong & Wang 1987
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Lathrobium homorum n. sp.5
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Sinocistela siphla n. gen. n. sp.5
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? Epicauta rufithoraca n. sp.5
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"Engonius alviolatus" = Gerstaeckerus alviolatus5
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Hylobius plenus5, Hylobius aff. niitakensis5, Hylobius medianus n. sp.5, "Sinocurculia shanwangensis n. sp." = Atactogaster shanwangensis2, "Sinocurculia granulata n. sp." = Atactogaster granulata
"Sinocurculia shanwangensis n. sp." = Atactogaster shanwangensis2 Hong 1985 snout beetle 80128, 78001, 80107, 80134, 80173, 80174
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Longifistulia parva n. gen. n. sp.3
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Cerambycidae "indet. 1"5, Alifasciola daedalea n. sp.5, Sinocalosoma longicornuta n. sp.2, Mesosa soteria n. sp.5, Mesocacia pulla n. sp.5
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Cupes longus n. sp.4
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"Procarabus longipes n. sp." = Cistelites longipes5
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