PBDB Collection

Guanling, Lower Xiaowa Formation fossil lagerstätte (Triassic of China)

Where: Guizhou, China (25.9° N, 105.4° E: paleocoordinates 26.9° N, 87.4° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Lower Member (Xiaowa Formation), Tuvalian (232.0 - 221.5 Ma)

• 12 m thick, previously known as Wayao Member of Falang Formation or as Wayao Formation

•JA: originally entered as Carnian; the Metapolygnathus polygnatiformis and M. nodosus zones are both Tuvalian according to Lucas et al. 2012

• group of beds-level stratigraphic resolution

Environment/lithology: deep-water; lithified shale and lithified marl

• deep and quiet water, the lower unit with generally oxygenated bottom water; the upper unit shaly interval with anoxic conditions and the greatest water depth

• lower unit (5 m thick) of medium- to thick-bedded, grey bioclastic micrites with yellow-green shale interbeds; upper unit (7 m thick) of greyish to black, thin- to medium-bedded laminated mudstone with argillaceous limestone interbeds and marlstones

Size class: macrofossils

Collection methods: quarrying, mechanical,

Primary reference: C. Li and O. Rieppel. 2002. A new cyamodontid placodont from Triassic of Guizhou, China. Chinese Science Bulletin 47(5):403-407 [M. Carrano/M. Carrano]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 85166: authorized by Wolfgang Kiessling, entered by Uta Merkel on 04.12.2008, edited by Bethany Allen

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

unclassified
-	Similingula cf. lipoldi² Similingula cf. lipoldi²
Holothuroidea
-	Holothuroidea indet.² Holothuroidea indet.² de Blainville 1834 sea cucumber
Crinoidea
Encrinida - Traumatocrinidae	Traumatocrinus sp.² Traumatocrinus sp.² Woehrmann 1889 Sea lily
Roveacrinida - Somphocrinidae	Osteocrinus cf. virgatus², Osteocrinus cf. spinosus² Osteocrinus cf. virgatus² Kristan-Tollmann 1970 Sea lily Osteocrinus cf. spinosus² Kristan-Tollmann 1970 Sea lily
Conodonta
Ozarkodinida - Gondolellidae	"Metapolygnathus polygnathiformis" = Paragondolella polygnathiformis², "Metapolygnathus nodosus" = Hayashiella nodosa² "Metapolygnathus polygnathiformis" = Paragondolella polygnathiformis² Budurov and Stefanov 1965 conodont "Metapolygnathus nodosus" = Hayashiella nodosa² Hayashi 1968 conodont
Reptilia
Placodontia - Placochelyidae	Psephochelys polyosteoderma n. gen. n. sp. Psephochelys polyosteoderma n. gen. n. sp. Li and Rieppel 2002 placodont IVPP V 12442
Placodontia - Cyamodontidae	Cyamodus orientalis n. sp.³, Sinocyamodus xinpuensis² Cyamodus orientalis n. sp.³ Wang et al. 2019 placodont Zhejiang Museum of Natural History, Hangzhou, ZMNH M8820 (holotype), "an articulated and nearly complete skeleton with three-dimensionally preserved skull and lacking only the forelimbs, representing an adult individual based on overall body size" Sinocyamodus xinpuensis² Li 2000 placodont
Ichthyosauria - Shastasauridae	"Guanlingichthyosaurus liangae" = Guanlingsaurus liangae², "Ghuizhouichthyosaurus tangae" = Guizhouichthyosaurus tangae² "Guanlingichthyosaurus liangae" = Guanlingsaurus liangae² Yin et al. 2000 ichthyosaur "Ghuizhouichthyosaurus tangae" = Guizhouichthyosaurus tangae² Cao et al. 2000 ichthyosaur
Ichthyosauria - Toretocnemidae	Qianichthyosaurus zhoui² Qianichthyosaurus zhoui² Li 1999 ichthyosaur
Thalattosauria - Askeptosauridae	Anshunsaurus huangguoshuensis² Anshunsaurus huangguoshuensis² Liu 1999 diapsid
Thalattosauria -	Xinpusaurus bamaolinensis², Xinpusaurus suni² Xinpusaurus bamaolinensis² Cheng 2003 diapsid Xinpusaurus suni² Yin 2000 diapsid
Testudines -	Odontochelys semitestacea n. gen. n. sp.¹ Odontochelys semitestacea n. gen. n. sp.¹ Li et al. 2008 turtle IVPP V 13240 (paratype), V 15639 (holotype), V 15653
Craniata
Craniida - Craniidae	? Crania sp.² ? Crania sp.² Retzius 1781
Cephalopoda
Ceratitida - Clionitidae	"Clionites cf. zeilleri" = Clionitites² "Clionites cf. zeilleri" = Clionitites² ceratite
Ceratitida - Tibetitidae	Hauerites cf. himalayanus² Hauerites cf. himalayanus² ceratite
Ceratitida - Trachyceratidae	Trachyceras sinensis², Trachyceras cf. aon², Trachyceras multituberculatum², Paratrachyceras cf. hofmanni², Protrachyceras cf. douvillei², Protrachyceras longiangense², Protrachyceras costulatum², Protrachyceras deprati², Protrachyceras ladinum² Trachyceras sinensis² ceratite Trachyceras cf. aon² Munster 1834 ceratite Trachyceras multituberculatum² Hsu 1940 ceratite Paratrachyceras cf. hofmanni² Boekh 1873 ceratite Protrachyceras cf. douvillei² ceratite Protrachyceras longiangense² ceratite Protrachyceras costulatum² Mansuy 1912 ceratite Protrachyceras deprati² Mansuy 1912 ceratite Protrachyceras ladinum² Mojsisovics 1882 ceratite
Bivalvia
Ostreida - Bakevelliidae	Angustella sp.² Angustella sp.² Waagen 1907 oyster
Ostreida - Pergamidiidae	Krumbeckiella sp.² Krumbeckiella sp.² Ichikawa 1958 oyster
Ostreida - Halobiidae	Halobia subcomata², Daonella bifurcata² Halobia subcomata² Kittl 1912 oyster Daonella bifurcata² oyster

¹Li et al. 2008; ²Wang et al. 2008; ³Wang et al. 2019