Elmo, YPM 1921 collection (Permian of the United States)

Also known as Insect Hill

Where: Kansas (38.6° N, 97.2° W: paleocoordinates 3.7° N, 25.8° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Wellington Formation, Artinskian (290.1 - 279.3 Ma)

• The Elmo limestone itself about five feet thick; only the very bottom, more massive layer of this stratum contains insects. Overlies underlying terrestrial (swamp) shale with a sharp and irregular contact.

• group of beds-level stratigraphic resolution

Environment/lithology: pond; lithified, gray, argillaceous limestone

• Presence of Myalinella may suggest brackish-marine conditions, but stratigraphic position and presence of conchostracans and freshwater ostracods suggest a lacustrine/pond setting.
• Soft, argillaceous limestone of very light grey color and almost chalky texture, disposed in layers 0.5 to 4 inches thick.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by C.O. Dunbar, R.J. Tillyard in 1921; reposited in the YPM

Collection methods: quarrying,

Primary reference: C. O. Dunbar and R. J. Tillyard. 1924. Kansas Permian insects: Part 1. The geologic occurrence and the environment of the insects, with a description of a new paleodictyopterid. American Journal of Science, Series 5 7(39):171-209 [C. Labandeira/C. Labandeira/R. Day]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111219: authorized by Matthew Clapham, entered by Matthew Clapham on 21.06.2011, edited by Jered Karr and Roy Plotnick

Creative Commons license: CC BY (attribution)

Taxonomic list

Euchelicerata
 Eurypterida - Adelophthalmidae
Adelophthalmus sellardsi Dunbar 1924 eurypterid
Xiphosura
 Xiphosurida - Paleolimulidae
"Paleolimulus avitus n. gen. n. sp." = Paleolimulus signatus5
"Paleolimulus avitus n. gen. n. sp." = Paleolimulus signatus5 Beecher 1904
Insecta
 Eoblattida - Euryptilonidae
Stereopterum rotundum3 Carpenter 1950 insect
YPM 15634
Stereopterum maculosum n. sp.4 Carpenter 1966 insect
YPM 15653
 Reculida - Sylvaphlebiidae
Phenopterum elongatum3 Sellards 1909 winged insect
YPM 15637,15583,15617,15632
 Dicondylia - Camptotaxineuridae
Camptotaxineura ephialtes n. gen. n. sp.25
Camptotaxineura ephialtes n. gen. n. sp.25 Tillyard 1937 winged insect
YPM 15590
 Dicondylia - Kennedyidae
Kennedya mirabilis n. gen. n. sp.10
Kennedya mirabilis n. gen. n. sp.10 Tillyard 1925 winged insect
YPM 5005-6
 Dicondylia - Ditaxineuridae
Ditaxineura anomalostigma n. gen. n. sp.11
Ditaxineura anomalostigma n. gen. n. sp.11 Tillyard 1926 winged insect
YPM 5054
 Meganisoptera - Meganeuridae
Meganeuridae indet.10, Megatypus ingentissimus n. sp.10, Megatypus schucherti n. gen. n. sp.10, "? Typus sp." = Tupus10, "Typus permianus" = Tupus permianus10
Meganeuridae indet.10 Handlirsch 1906 griffinfly
YPM 5014-9, 5027-34; wing fragments
Megatypus ingentissimus n. sp.10 Tillyard 1925 griffinfly
YPM 5010, ?5025, ?5026
Megatypus schucherti n. gen. n. sp.10 Tillyard 1925 griffinfly
YPM 1021, 5009
"? Typus sp." = Tupus10 Sellards 1906 griffinfly
YPM 5011, 5023; abdominal fragments
"Typus permianus" = Tupus permianus10 Sellards 1906 griffinfly
YPM 5008, ?5024, ?5012, ?5013
 Ephemerida - Protereismatidae
Protereisma sp.20, Protereisma insigne n. sp.20, Protereisma elongatum20, Protereisma latum20, "Protereisma permianum" = Protereisma permianum20, "Protereisma minus" = Protereisma permianum20, "Protereisma gracile" = Protereisma permianum20, Protereisma arcuatum20, Protereisma sellardsi n. sp.20
Protereisma sp.20 Sellards 1907 mayfly
YPM 5466-5469,5473-5506
Protereisma insigne n. sp.20 Tillyard 1932 mayfly
YPM 1112
Protereisma elongatum20 Sellards 1907 mayfly
YPM 1102,1113,5432,5439,5453-4
Protereisma latum20 Sellards 1907 mayfly
YPM 1109,1115-6,5452,5460-4
"Protereisma permianum" = Protereisma permianum20 Sellards 1907 mayfly
YPM 1009,1023,1103,1105,1114,1119,1120,5427-30,5433,5436,5437,5449-50
"Protereisma minus" = Protereisma permianum20 Sellards 1907 mayfly
YPM 5451
"Protereisma gracile" = Protereisma permianum20 Sellards 1907 mayfly
YPM 1104,5431,5435,5438,5440-2,5444,5446-8
Protereisma arcuatum20 Sellards 1907 mayfly
YPM 5455-7
Protereisma sellardsi n. sp.20 Tillyard 1932 mayfly
YPM 5434
 Ephemerida - Misthodotidae
"Eudoter delicatulus n. gen. n. sp." = Misthodotes delicatulus21, Misthodotes ovalis n. sp.20, Misthodotes obtusus20, Misthodotes biguttatus n. sp.20
"Eudoter delicatulus n. gen. n. sp." = Misthodotes delicatulus21 Tillyard 1936 mayfly
YPM 1014
Misthodotes ovalis n. sp.20 Tillyard 1932 mayfly
YPM 1118
Misthodotes obtusus20 Sellards 1907 mayfly
YPM 1100,1117,5443,?5459,?5465,5470,5471
Misthodotes biguttatus n. sp.20 Tillyard 1932 mayfly
YPM 1122
 Palaeodictyoptera - Spilapteridae
Dunbaria fasciipennis n. gen. n. sp.
Dunbaria fasciipennis n. gen. n. sp. Tillyard 1924 winged insect
YPM 1001,1002,1050, 5020-2
 Palaeodictyoptera -
Kansasia pulchra n. gen. n. sp.25
Kansasia pulchra n. gen. n. sp.25 Tillyard 1937 winged insect
YPM 15589
 Palaeodictyoptera - Calvertiellidae
Calvertiella permiana n. gen. n. sp.10
Calvertiella permiana n. gen. n. sp.10 Tillyard 1925 winged insect
YPM 5007
 Megasecoptera - Protohymenidae
Protohymen permianus n. gen. n. sp.9, Permohymen schucherti n. gen. n. sp.9
Protohymen permianus n. gen. n. sp.9 Tillyard 1924 winged insect
YPM 5001/5002 (actually counterpart), Cawthron Institute 1702, YPM 5035-43, 5053
Permohymen schucherti n. gen. n. sp.9 Tillyard 1924 winged insect
YPM 5003, 5043
 Diaphanopterodea - Elmoidae
Elmoa trisecta n. gen. n. sp.25
Elmoa trisecta n. gen. n. sp.25 Tillyard 1937 winged insect
YPM 15591
 Diaphanopterodea - Martynoviidae
"Martynoviella protohymenoides n. gen. n. sp." = Martynovia protohymenoides19, Martynovia insignis n. gen. n. sp.19
"Martynoviella protohymenoides n. gen. n. sp." = Martynovia protohymenoides19 Tillyard 1932 winged insect
YPM 5424
Martynovia insignis n. gen. n. sp.19 Tillyard 1932 winged insect
YPM 5423
 Diaphanopterodea - Asthenohymenidae
Asthenohymen sp.11 Tillyard 1924 winged insect
YPM 5052
Asthenohymen affinis n. sp.11 Tillyard 1926 winged insect
YPM 5050, ?5048
Asthenohymen dunbari n. gen. n. sp.9 Tillyard 1924 winged insect
YPM 5004, Cawthron Institute 1706
Asthenohymen stigmatizans n. sp.11 Tillyard 1926 winged insect
YPM 5046
Asthenohymen pusillus n. sp.11 Tillyard 1926 winged insect
YPM 5051
Asthenohymen kansasensis n. sp.11 Tillyard 1926 winged insect
YPM 5044
Asthenohymen stenobasis n. sp.11 Tillyard 1926 winged insect
YPM 5049
Asthenohymen gracilis n. sp.11 Tillyard 1926 winged insect
YPM 5045, ?5047
 Miomoptera - Palaeomanteidae
"Delopteridae indet." = Palaeomanteidae17, Delopsocus fasciatus n. sp.17, Delopsocus latus17, Delopsocus elongatus17, "Delopterum minutum" = Delopterum minutum17, "Delopterum dunbari n. sp." = Delopterum minutum17
"Delopteridae indet." = Palaeomanteidae17 Handlirsch 1906 winged insect
YPM 5395,5396,5398
Delopsocus fasciatus n. sp.17 Tillyard 1928 winged insect
YPM 5376,5377,5404
Delopsocus latus17 Sellards 1909 winged insect
YPM 5378-5384,5401
Delopsocus elongatus17 Sellards 1909 winged insect
YPM 5386,5389
"Delopterum minutum" = Delopterum minutum17 Sellards 1909 winged insect
YPM 5385, ?5393, ?5394
"Delopterum dunbari n. sp." = Delopterum minutum17 Sellards 1909 winged insect
YPM ?5387-8,?5391,5392,?5397,?5399,?5400,5402
 Hemiptera - Archescytinidae
Archescytina permiana n. gen. n. sp.14, "Permoscytina kansasensis n. gen. n. sp." = Archescytina kansasensis14, "Permoscytina muiri n. sp." = Archescytina muiri14, Permopsylla americana n. gen. n. sp.14
Archescytina permiana n. gen. n. sp.14 Tillyard 1926 true bug
YPM 5111
"Permoscytina kansasensis n. gen. n. sp." = Archescytina kansasensis14 Tillyard 1926 true bug
YPM 5112
"Permoscytina muiri n. sp." = Archescytina muiri14 Tillyard 1926 true bug
YPM 5113
Permopsylla americana n. gen. n. sp.14 Tillyard 1926 true bug
YPM 5114
 Permopsocida - Psocidiidae
"Psocidium minimum n. sp." = Dichentomum minimum13, "Psocidium sp." = Dichentomum13, "Dichentomum tinctum n. gen. n. sp." = Dichentomum tinctum13, "Psocidium kansasense n. sp." = Dichentomum tinctum13, "Permentomum tenuiforme n. gen. n. sp." = Dichentomum tinctum13, "Psocidium abnormale n. sp." = Dichentomum tinctum13, "Pentapsocidium indistinctum n. gen. n. sp." = Dichentomum tinctum13, "Psocidium permianum n. gen. n. sp." = Dichentomum tinctum13, "Metapsocidium loxoneurum n. gen. n. sp." = Dichentomum tinctum13, "Psocidium robustum n. sp." = Dichentomum tinctum13, "Chaetopsocidium sellardsi n. gen. n. sp." = Dichentomum tinctum13
"Psocidium minimum n. sp." = Dichentomum minimum13 Tillyard 1926 winged insect
YPM 5110
"Psocidium sp." = Dichentomum13 Tillyard 1926 winged insect
YPM 5083, 5090-5097, 5103
"Dichentomum tinctum n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5071
"Psocidium kansasense n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5075-5078, ?5079-82, ?5084-89
"Permentomum tenuiforme n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5101
"Psocidium abnormale n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5073
"Pentapsocidium indistinctum n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5100
"Psocidium permianum n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5072
"Metapsocidium loxoneurum n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5099
"Psocidium robustum n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5074, 5102
"Chaetopsocidium sellardsi n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5098
 Permopsocida - Permopsocidae
"Permopsocus latipennis n. gen. n. sp." = Permopsocus latipennis13, "Ancylopsocus insolitus n. gen. n. sp." = Permopsocus latipennis13, "Permopsocus congener n. sp." = Permopsocus latipennis13, "Permopsocus enderleini n. sp." = Permopsocus latipennis13, "Progonopsocus permianus n. gen. n. sp." = Progonopsocus permianus13, "Progonopsocus pusillus n. sp." = Progonopsocus permianus13
"Permopsocus latipennis n. gen. n. sp." = Permopsocus latipennis13 Tillyard 1926 winged insect
YPM 5106
"Ancylopsocus insolitus n. gen. n. sp." = Permopsocus latipennis13 Tillyard 1926 winged insect
YPM 5109
"Permopsocus congener n. sp." = Permopsocus latipennis13 Tillyard 1926 winged insect
YPM 5107
"Permopsocus enderleini n. sp." = Permopsocus latipennis13 Tillyard 1926 winged insect
YPM 5108
"Progonopsocus permianus n. gen. n. sp." = Progonopsocus permianus13 Tillyard 1926 winged insect
YPM 5104
"Progonopsocus pusillus n. sp." = Progonopsocus permianus13 Tillyard 1926 winged insect
YPM 5105
 Orthoptera - Pruvostitidae
Paroedischia recta4 Carpenter 1966 grasshopper
YPM 15757,15638
 Orthoptera - Proparagryllacrididae
Crinoedischia buttsi n. gen. n. sp.1
Crinoedischia buttsi n. gen. n. sp.1 Béthoux and Beckemeyer 2007 grasshopper
YPM 204902
 Orthoptera - Permelcanidae
Promartynovia venicosta n. gen. n. sp.25
Promartynovia venicosta n. gen. n. sp.25 Tillyard 1937 grasshopper
YPM 15594
 Orthoptera - Permoraphidiidae
Permoraphidia americana n. gen. n. sp.19
Permoraphidia americana n. gen. n. sp.19 Tillyard 1932 grasshopper
YPM 5422
 Paoliida - Anthracoptilidae
"Homocladus ornatus n. sp." = Homocladus grandis4
"Homocladus ornatus n. sp." = Homocladus grandis4 Carpenter 1966 winged insect
YPM 15584
 Paoliida - Blattinopsidae
Blattinopsis kukalovae4 Carpenter 1966 winged insect
YPM 15582,15633
 Grylloblattodea - Probnidae
Probnis speciosa23 Sellards 1909 ice crawler
YPM 15508-15557,15560,15565-67,15599-601,15603-5,15575,15751-87
 Grylloblattodea - Chelopteridae
Chelopterum peregrinum3 Carpenter 1950 ice crawler
 Grylloblattodea - Camptoneuritidae
Demopterum gracile3 Carpenter 1950 ice crawler
YPM 15793,15562,15646
 Grylloblattodea - Lemmatophoridae
"Lemmatophora minuta" = Lisca minuta15, "Lemmatophora reducta n. sp." = Lisca minuta15, "Lemmatophora cubitalis n. sp." = Lisca minuta7, Paraprisca grandis25, "Paraprisca fragilis" = Paraprisca fragilis16, "Paraprisca stigmatica n. sp." = Paraprisca fragilis25, "Lemmatophora longipennis n. sp." = Paraprisca fragilis7, "Lecorium elongatum" = Lecorium elongatum16, "Sellardsia kansasensis n. gen. n. sp." = Lecorium elongatum16, "Sellardsia lecorioides n. sp." = Lecorium elongatum16, "Sellardsia abnormis n. sp." = Lecorium elongatum7, "Metalecorium confusum n. sp." = Lecorium elongatum7, "Metalecorium anomalum n. sp." = Lecorium elongatum7, "Sellardsia separanda n. sp." = Lecorium elongatum7, "Lemmatophora typica" = Lemmatophora typa15, Artinska sellardsi n. sp.16, "Artinska clara" = Artinska clara16, "Orta quadrifida n. sp." = Artinska clara7, "Estadia maculipennis n. sp." = Artinska clara7, "Estadia altera n. sp." = Artinska clara7, "Estadia banneria n. sp." = Artinska clara7, "Artinska pentamera n. sp." = Artinska clara7, "Estadia reducta n. sp." = Artinska clara7, "Artinska simplex n. sp." = Artinska clara7, "Estadia aucta n. sp." = Artinska clara7, "Artinska tillyardi n. sp." = Artinska clara7, "Estadia kansasia n. sp." = Artinska clara7, "Artinska nova n. sp." = Artinska clara7, "Estadia confusa n. sp." = Artinska clara7, "Artinska inversa n. sp." = Artinska clara7, "Artinska ovata" = Artinska ovata16, "Orta reducta n. sp." = Artinska ovata7
"Lemmatophora minuta" = Lisca minuta15 Sellards 1909 ice crawler
YPM 5193,5196-5198,5206-5223,5225-5246,5259-5261
"Lemmatophora reducta n. sp." = Lisca minuta15 Sellards 1909 ice crawler
YPM 5224
"Lemmatophora cubitalis n. sp." = Lisca minuta7 Sellards 1909 ice crawler
YPM 5208 (one of Lemmatophora minuta in Tillyard)
Paraprisca grandis25 Carpenter 1935 ice crawler
YPM 15576
"Paraprisca fragilis" = Paraprisca fragilis16 Sellards 1909 ice crawler
YPM 5195,5250-5258,5262,5284
"Paraprisca stigmatica n. sp." = Paraprisca fragilis25 Sellards 1909 ice crawler
YPM 15579,15596-7
"Lemmatophora longipennis n. sp." = Paraprisca fragilis7 Sellards 1909 ice crawler
YPM 5194 (one of Paraprisca fragilis in Tillyard)
"Lecorium elongatum" = Lecorium elongatum16 Sellards 1909 ice crawler
YPM 5267,5270,5272-7,5279,5281 (several specimens made types of Handlirsch spp.)
"Sellardsia kansasensis n. gen. n. sp." = Lecorium elongatum16 Sellards 1909 ice crawler
YPM 5266,5268-9,5278,5280,5282-3 (several specimens made types of Handlirsch spp.)
"Sellardsia lecorioides n. sp." = Lecorium elongatum16 Sellards 1909 ice crawler
YPM 5271
"Sellardsia abnormis n. sp." = Lecorium elongatum7 Sellards 1909 ice crawler
YPM 5268
"Metalecorium confusum n. sp." = Lecorium elongatum7 Sellards 1909 ice crawler
YPM 5272
"Metalecorium anomalum n. sp." = Lecorium elongatum7 Sellards 1909 ice crawler
YPM 5267
"Sellardsia separanda n. sp." = Lecorium elongatum7 Sellards 1909 ice crawler
YPM 5278
"Lemmatophora typica" = Lemmatophora typa15 Sellards 1909 ice crawler
YPM 5115-5192,5194,5195,5199-5205
Artinska sellardsi n. sp.16 Tillyard 1928 ice crawler
YPM 5291
"Artinska clara" = Artinska clara16 Sellards 1909 ice crawler
YPM 1003,5263-5265,5285-5290,5292-5312,5332-5363 (several specimens made types of Handlirsch spp.)
"Orta quadrifida n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5269
"Estadia maculipennis n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5264
"Estadia altera n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5298, 5303
"Estadia banneria n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5302, 5304
"Artinska pentamera n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5263
"Estadia reducta n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5296
"Artinska simplex n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5286
"Estadia aucta n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5295
"Artinska tillyardi n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5265
"Estadia kansasia n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5297, 5299
"Artinska nova n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5290
"Estadia confusa n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5306
"Artinska inversa n. sp." = Artinska clara7 Sellards 1909 ice crawler
YPM 5287
"Artinska ovata" = Artinska ovata16 Sellards 1909 ice crawler
YPM 5313-5331,5364-5375
"Orta reducta n. sp." = Artinska ovata7 Sellards 1909 ice crawler
YPM 5277
 Grylloblattodea - Liomopteridae
Semopterum venosum3 Carpenter 1950 ice crawler
Liomopterum elongatum3 Sellards 1909 ice crawler
YPM 15612, 15749, 15573
Liomopterum ornatum3 Sellards 1909 ice crawler
Liomopterum sellardsi3 Carpenter 1950 ice crawler
YPM 16180
 Protorthoptera -
? Lecopterum delicosum23 Sellards 1909 winged insect
YPM 15574
 Dicondylia - Phyloblattidae
"Pycnoblattina compacta" = Phyloblatta compacta26, "Pycnoblattina curvata" = Phyloblatta curvata26
"Pycnoblattina compacta" = Phyloblatta compacta26 Sellards 1908 winged insect
YPM 15684-90, 15694-5, 15735-9
"Pycnoblattina curvata" = Phyloblatta curvata26 Sellards 1908 winged insect
YPM 15692-3,15740-5
 Dicondylia - Spiloblattinidae
? Spiloblattinidae indet.26 Handlirsch 1906 winged insect
protnotum YPM 15698
Pycnoblattina sp.26 Sellards 1908 winged insect
hindwings YPM 15701,15746-9, pronotum YPM 15697
Permoblattina permiana22 Sellards 1908 winged insect
YPM 15677-83,15699,15702,15724-32
Permoblattina curta n. gen.22 Sellards 1908 winged insect
YPM 15663-76,15678,15700,15703-23
 Dicondylia - Nugonioneuridae
Nugonioneura problematica n. gen. n. sp.25
Nugonioneura problematica n. gen. n. sp.25 Tillyard 1937 winged insect
YPM 15561, 27536
 Cnemidolestodea - Protembiidae
"Protembia permiana n. gen. n. sp." = Protembia permiana24, "Telactinopteryx striatipennis n. gen. n. sp." = Protembia permiana23
"Protembia permiana n. gen. n. sp." = Protembia permiana24 Tillyard 1937 winged insect
YPM 15507
"Telactinopteryx striatipennis n. gen. n. sp." = Protembia permiana23 Tillyard 1937 winged insect
YPM 15558-9,15602
 Caloneurodea - Permobiellidae
Permobiella perspicua n. gen. n. sp.25
Permobiella perspicua n. gen. n. sp.25 Tillyard 1937 winged insect
YPM 15593
 Caloneurodea -
Paleuthygramma acuta2 Carpenter 1943 winged insect
YPM 15561,15655-7,15571,15572
 Protomeroptera - Protomeropidae
"Platychorista venosa n. gen. n. sp." = Platychorista venosa12, "Protomerope permiana n. gen. n. sp." = Platychorista venosa12
"Platychorista venosa n. gen. n. sp." = Platychorista venosa12 Tillyard 1926 winged insect
YPM 5067
"Protomerope permiana n. gen. n. sp." = Platychorista venosa12 Tillyard 1926 winged insect
YPM 5069, 5070
 Mecoptera - Anormochoristidae
Anormochorista oligoclada n. gen. n. sp.12
Anormochorista oligoclada n. gen. n. sp.12 Tillyard 1926 scorpionfly
YPM 5068
 Mecoptera - Permopanorpidae
Permopanorpa schucherti n. sp.12, Permopanorpa formosa n. sp.12, "Permopanorpa inaequalis n. sp." = Permopanorpa inaequalis12, "Permopanorpa sellardsi n. sp." = Permopanorpa inaequalis12, "Permopanorpa dunbari n. sp." = Permopanorpa inaequalis12, "Permopanorpa tenuis n. sp." = Permopanorpa inaequalis12, "Permopanorpa gracilis n. sp." = Permopanorpa inaequalis12, "Protopanorpa pusilla n. sp." = Lithopanorpa pusilla12
Permopanorpa schucherti n. sp.12 Tillyard 1926 scorpionfly
YPM 5061
Permopanorpa formosa n. sp.12 Tillyard 1926 scorpionfly
YPM 5057
"Permopanorpa inaequalis n. sp." = Permopanorpa inaequalis12 Tillyard 1926 scorpionfly
YPM 5058
"Permopanorpa sellardsi n. sp." = Permopanorpa inaequalis12 Tillyard 1926 scorpionfly
YPM 5063
"Permopanorpa dunbari n. sp." = Permopanorpa inaequalis12 Tillyard 1926 scorpionfly
YPM 5062
"Permopanorpa tenuis n. sp." = Permopanorpa inaequalis12 Tillyard 1926 scorpionfly
YPM 5059
"Permopanorpa gracilis n. sp." = Permopanorpa inaequalis12 Tillyard 1926 scorpionfly
YPM 5060
"Protopanorpa pusilla n. sp." = Lithopanorpa pusilla12 Tillyard 1926 scorpionfly
YPM 5066
 Mecoptera - Permochoristidae
"Protochorista tetraclada n. gen. n. sp." = Protochorista tetraclada12, "Protochorista pentaclada n. sp." = Protochorista tetraclada12, Protopanorpa permiana n. gen. n. sp.12
"Protochorista tetraclada n. gen. n. sp." = Protochorista tetraclada12 Tillyard 1926 scorpionfly
YPM 5055
"Protochorista pentaclada n. sp." = Protochorista tetraclada12 Tillyard 1926 scorpionfly
YPM 5056
Protopanorpa permiana n. gen. n. sp.12 Tillyard 1926 scorpionfly
YPM 5064, 5065
 Protoneuroptera - Permoberothidae
"Permoberotha villosa n. gen. n. sp." = Permoberotha villosa19, "Permoberotha convergens n. gen. n. sp." = Permoberotha villosa25, "Dictyobiella nervosa n. gen. n. sp." = Permoberotha villosa25
"Permoberotha villosa n. gen. n. sp." = Permoberotha villosa19 Tillyard 1932 winged insect
YPM 5426
"Permoberotha convergens n. gen. n. sp." = Permoberotha villosa25 Tillyard 1932 winged insect
YPM 15595
"Dictyobiella nervosa n. gen. n. sp." = Permoberotha villosa25 Tillyard 1932 winged insect
YPM 15592
 Dicondylia - Permembiidae
Permembia delicatula n. gen. n. sp.17
Permembia delicatula n. gen. n. sp.17 Tillyard 1928 winged insect
YPM 5403
 Dicondylia -
Paraneoptera indet.8 winged insect
YPM 5419
 Protelytroptera - Permelytridae
"? Parablattelytron sp." = Parablattelytron18, Parablattelytron rectum n. sp.18, "Acosmelytron elongatum n. gen. n. sp." = Parablattelytron elongatum18, "Acosmelytron acutum n. sp." = Parablattelytron elongatum18, Parablattelytron subincisum n. gen. n. sp.18, "? Acosmelytron sp." = Parablattelytron18, "Acosmelytron delicatum n. sp." = Parablattelytron delicatum18, Permelytron schucherti n. gen. n. sp.18, Blattelytron permianum n. gen. n. sp.18
"? Parablattelytron sp." = Parablattelytron18 Tillyard 1931 winged insect
YPM 5412
Parablattelytron rectum n. sp.18 Tillyard 1931 winged insect
YPM 5413
"Acosmelytron elongatum n. gen. n. sp." = Parablattelytron elongatum18 Tillyard 1931 winged insect
YPM 5415
"Acosmelytron acutum n. sp." = Parablattelytron elongatum18 Tillyard 1931 winged insect
YPM 5414
Parablattelytron subincisum n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 5408
"? Acosmelytron sp." = Parablattelytron18 Tillyard 1931 winged insect
YPM 5407,5410
"Acosmelytron delicatum n. sp." = Parablattelytron delicatum18 Tillyard 1931 winged insect
YPM 5416
Permelytron schucherti n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 5411
Blattelytron permianum n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 5409
 Protelytroptera - Megelytridae
Megelytron robustum n. gen. n. sp.18
Megelytron robustum n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 5406
 Protelytroptera - Protelytridae
? Protelytron sp.18 Tillyard 1931 winged insect
YPM 5421
Protelytron dunbari n. sp.18 Tillyard 1931 winged insect
YPM 5405,5417-8
Protelytron permianum n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 1019
 Protelytroptera -
Protelytropsis grandis n. gen. n. sp.18
Protelytropsis grandis n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 5420
 Dicondylia - Choristosialidae
Choristosialis enigmatica n. gen. n. sp.19
Choristosialis enigmatica n. gen. n. sp.19 Tillyard 1932 winged insect
YPM 5425
 Archaeognatha - Dasyleptidae
"Lepidodasypus sharovi n. gen. n. sp." = Dasyleptus sharovi6
"Lepidodasypus sharovi n. gen. n. sp." = Dasyleptus sharovi6 Durden 1978 jumping bristletails
YPM 28152
Bivalvia
 Myalinida - Myalinidae
"Myalina meeki n. sp." = Myalinella meeki
"Myalina meeki n. sp." = Myalinella meeki Dunbar 1924 clam