Hagen-Balve Reef Complex, Binolen outcrop - section (Devonian of Germany)

Where: Germany (51.4° N, 7.9° E: paleocoordinates 22.1° S, 7.5° E)

• coordinate stated in text

When: Binolen Member (Hagen-Balve Formation), Early/Lower Givetian (388.1 - 383.7 Ma)

• member-level stratigraphic resolution

Environment/lithology: reef, buildup or bioherm; siltstone

• biostromal initial reef succession with total thickness of 14m
• biostromal limestones; bed-by-bed survey started with the sharp onset of dark- to middle-grey, calcareous, coarse crinoid limestones and first substrate-stabilising organisms

•(e.g. sponges, tabulate corals, and stromatoporoids)

Size classes: macrofossils, mesofossils, microfossils

Primary reference: M. Löw, T. Söte, R. T. Becker, S. Stichling, A. May, Z. S. Aboussalam, and S. F. Zoppe. 2022. The initial phase of the Hönne Valley Reef at Binolen (northern Rhenish Massif, Middle Devonian). Palaeobiodiversity and Palaeoenvironments [B. Seuss/B. Seuss]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 227413: authorized by Barbara Seuss, entered by Barbara Seuss on 11.10.2022

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
unclassified
  -
Bipennatus bipennatus, Dvorkia chattertoni, Eonodosarioidea indet., Octactinellidae indet., Gerronostromaria sp., Pachytheca stellimicans, Bryozoa indet., Murchisonia sp., Binoleniceras stichlingi, Scolecodonta indet., Gastropoda indet., Podocopidde indet., Ophiuroidea indet., Vertebrata indet., Givetianella tsienii, ? Resteignella sp., Rothpletzella sp., Calcitarcha indet.
Bipennatus bipennatus
Beds 4, 18a, 18b, 23, and 29
Bryozoa indet.
indeteminate branch-like bryozoans, indeterminate fenestellids
Gastropoda indet.
juveniles
Vertebrata indet.
indeterminate tooth plate
Calcitarcha indet.
calcispheres
Chlorophyceae
 Dasycladales - Beresellaceae
Kamaena sp.
Kamaena sp. Antropov 1967
Ulvophyceae
 Bryopsidales - Codiaceae
Litanaia sp.
Litanaia sp. Maslov 1956
Foraminifera
 Ammodiscoidea - Saccamminidae
Thurammina cf. congesta, Thurammina cf. sphaerica
 Ammodiscoidea - Ammodiscidae
Tolypammina cf. continuus, Tolypammina tortuosa, Tolypammina aff. tortuosa
 Astrorhizacea - Hemisphaeramminidae
Webbinelloidea hemispherica
 Astrorhizacea - Rhizamminidae
Rhizammina cf. algaeformis
 Astrorhizacea - Psammosphaeridae
Psammosphaera fusca
Florideophyceae
 Corallinales - Corallinophycidae
Halysis moniliformis
Protozoa
  -
Angochitina pseudoavelinoi, Lagenochitina avelinoi
Anthozoa
 Favositida - Favositidae
Favosites sp., Favosites goldfussi, Pachyfavosites polymorphus
Favosites sp. Lamarck 1816 tabulate coral
Favosites goldfussi Lecompte 1939 tabulate coral
 Favositida - Pachyporidae
Thamnopora angusta, Thamnopora sp., "Trachypora circulipora" = Thamnoptychia
Thamnopora angusta tabulate coral
Thamnopora sp. Steininger 1831 tabulate coral
"Trachypora circulipora" = Thamnoptychia tabulate coral
 Favositida - Alveolitidae
Alveolites (Tetralites) sp., Alveolites (Alveolitella) sp., Alveolites (Alveolites) sp., Alveolites (Alveolites) pseudorbicularis, Scoliopora insueta, Caliapora sp., Squameoalveolites sp.
Alveolites (Tetralites) sp. Lamarck 1801 tabulate coral
Alveolites (Alveolitella) sp. Lamarck 1801 tabulate coral
Alveolites (Alveolites) sp. Lamarck 1801 tabulate coral
Alveolites (Alveolites) pseudorbicularis Iven 1980 tabulate coral
Alveolites (Alveolites) suborbicularis pseudorbicularis
Scoliopora insueta tabulate coral
Caliapora sp. Schlüter 1889 tabulate coral
Squameoalveolites sp. Mironova 1969 tabulate coral
 Favositida - Coenitidae
Platyaxum (Egosiella) clathratum, Platyaxum (Egosiella) sp., Platyaxum (Roseoporella) gradatum, Platyaxum (Roseoporella) sp.
Platyaxum (Egosiella) clathratum tabulate coral
Platyaxum (Egosiella) ex gr. clathratum
Platyaxum (Egosiella) sp. Davis 1887 tabulate coral
Platyaxum (Roseoporella) sp. Davis 1887 tabulate coral
 Auloporida - Syringoporidae
Roemerolites rhiphaeus, Roemerolites sp., Syringopora expansa, Syringopora hanshanensis
Roemerolites rhiphaeus tabulate coral
Roemerolites brevis rhiphaeus
Roemerolites sp. Dubatolov 1963 tabulate coral
Syringopora expansa tabulate coral
Syringopora hanshanensis tabulate coral
 Auloporida - Auloporidae
Aulopora (Mastopora) sp., Aulopora (Aulopora) sp.
Aulopora (Mastopora) sp. Goldfuss 1829 tabulate coral
Aulopora (Aulopora) sp. Goldfuss 1829 tabulate coral
 Auloporida - Romingeriidae
Remesia crispa
Remesia crispa tabulate coral
 Stauriida - Stauriidae
Dendrostella trigemme
 Stauriida - Ptenophyllidae
Dohmophyllum difficile
 Stauriida - Spongophyllidae
Spongophyllum pusillum
Spongophyllum pusillum horn coral
Spongophyllum immersum pusillum
 Stauriida - Disphyllidae
? Disphyllum sp.
? Disphyllum sp. Fromentel 1861 horn coral
 Cystiphyllida - Digonophyllidae
Mesophyllum (Mesophyllum) vesiculosum
 Rugosa - Charactophyllidae
Temnophyllum latum, Temnophyllum astrictum, Spinophyllum ex gr. spongiosum
Spinophyllum ex gr. spongiosum Schlüter 1889 horn coral
unclassified
  -
Porifera indet.
Porifera indet. Grant 1836
Monaxones
Dvorcia
  - Polyactinellidae
Dvorcia cf. mira
Polyactinellidae
  - Polyactinellidae
Praephobetractinia eocambrica
Heteractinida
 Octactinellida - Eiffeliidae
Gondekia sp.
Gondekia sp. Rigby 1991
 Octactinellida - Astraeospongiidae
Ensiferites langeri
Ensiferites langeri Kozur et al. 2008
Stromatoporoidea
 Stromatoporellida - Idiostromatidae
Idiostroma sp.
Idiostroma sp. Winchell 1867 stromatoporoid sponge
 Stromatoporellida - Stromatoporellidae
Clathrocoilona sp., Clathrocoilona (Clathrocoilona) curiosa, Clathrocoilona spissa, Clathrocoilona obliterata, Stromatoporella sp.
Clathrocoilona sp. Yavorsky 1931 stromatoporoid sponge
Clathrocoilona spissa Lecompte 1951 stromatoporoid sponge
Clathrocoilona (Clathrocoilona) solidula spissa
Clathrocoilona obliterata Lecompte 1951 stromatoporoid sponge
Stromatoporella sp. Nicholson 1886 stromatoporoid sponge
 Stromatoporellida - Trupetostromatidae
Hermatostroma sp.
Hermatostroma sp. Nicholson 1886 stromatoporoid sponge
 Stromatoporida - Stromatoporidae
Stromatopora huepschii
Stromatopora huepschii Bargatzky 1881 stromatoporoid sponge
 Amphiporida - Amphiporidae
Amphipora sp.
Amphipora sp. Schulz 1883 stromatoporoid sponge
 Syringostromatida - Stachyoditidae
Stachyodes caespitosa, Stachyodes (Stachyodes) sp., Stachyodes radiata
Stachyodes caespitosa stromatoporoid sponge
Stachyodes (Stachyodes) ex gr. caespitosa
Stachyodes (Stachyodes) sp. Bargatzky 1881 stromatoporoid sponge
Stachyodes radiata Lecompte 1952 stromatoporoid sponge
Stachyodes (Stachyodes) ex gr. radiata
Osteichthyes
 Onychodontiformes - Onychodontidae
? Onychodus sp.
? Onychodus sp. Newberry 1857 lobe-finned fish
 Actinopterygii -
Actinopterygii indet.
Actinopterygii indet. Cope 1887 ray-finned fish
scale
Conodonta
 Panderodontida - Panderodontidae
Panderodus cf. unicostatus, Panderodus cf. unicostatus, ? Neopanderodus sp., Neopanderodus perlineatus, Neopanderodus perlineatus
Panderodus cf. unicostatus Branson and Mehl 1933 conodont
Bed 16 and Bed 20b
Panderodus cf. unicostatus Branson and Mehl 1933 conodont
? Neopanderodus sp. Ziegler and Lindström 1971 conodont
Bed 5
Neopanderodus perlineatus Ziegler and Lindstroem 1971 conodont
Bed 6, probably a juvenile in bed 21
Neopanderodus perlineatus Ziegler and Lindstroem 1971 conodont
 Ozarkodinida - Polygnathidae
Polygnathidae indet.
Polygnathidae indet. Bassler 1925 conodont
Beds 4, 18a, 23; polygnathid ramiforms
 Prioniodontida - Icriodontidae
cf. Icriodus latecarinatus, Icriodus ? latecarinatus
 Prioniodontida - Prioniodinidae
Prioniodinidae indet., Prioniodina sp.
Prioniodinidae indet. Bassler 1925 conodont
elements occur in beds 6 and 9
Prioniodina sp. Ulrich and Bassler 1925 conodont
 Belodellida - Belodellidae
Belodella resima, Dvorakia klapperi
Belodella resima Philip 1965 conodont
Bed 16 and Bed 20b
 Protopanderodontida - Protopanderodontidae
Drepanodus sp.
Drepanodus sp. Pander 1856 conodont
Bed 4
Crinoidea
  -
Crinoidea indet.
Crinoidea indet. Miller 1821 Sea lily
columnals, 3 morphotypes
 Cyathocrinida - Cupressocrinitidae
Cupressocrinitidae indet.
Cupressocrinitidae indet. Roemer 1954 Sea lily
Cupressocrinitid columnal Morphotype A and B
  -
Glyphidocrinus sp.
Glyphidocrinus sp. Dubatolova 1971 Sea lily
Holothuroidea
 Dendrochirotida - Calclamnidae
Eocaudina cf. patella, Eocaudina ovalis
Eocaudina cf. patella Boczarowski 2001 sea cucumber
Eocaudina ovalis Matyja et al. 1973 sea cucumber
 Dendrochirotida - Cucumariidae
Staurocaudina cf. dombrowiana
Staurocaudina cf. dombrowiana Boczarowski 2001 sea cucumber
Echinoidea
  -
Echinoidea indet.
Echinoidea indet. Leske 1778 sea urchin
Gastropoda
 Euomphalina - Euomphalidae
Euomphalidae indet.
Euomphalidae indet. White 1877 snail
juveniles
Bivalvia
 Megalodontida - Megalodontidae
Eomegalodus sp.
Rhynchonellata
 Terebratulida - Stringocephalidae
Stringocephalus burtini
Stringocephalus burtini Defrance 1825
Stringocephalus ex. gr. burtini
 Atrypida - Atrypidae
Atrypidae indet.
Atrypidae indet. Gill 1871
Stenolaemata
 Cystoporata - Fistuliporidae
Fistulipora sp.
Fistulipora sp. McCoy 1850
 Cryptostomata -
Cryptostomata indet.
Cryptostomata indet. Vine 1884
rhabdomesid Cryptostomata
 Cryptostomata - Fenestellidae
? Spinofenestella sp.
? Spinofenestella sp. Termier and Termier 1971
Polychaeta
 Eunicemorpha -
Skalenoprion sp.
 Eunicida - Polychaetaspidae
Oenonites sp.
Oenonites sp. Hinde 1879
Deuteropoda
  -
Ostracoda indet.
Ostracoda indet. Latreille 1802 ostracod
Ostracoda
 Palaeocopida - Hollinellidae
? Hollinella sp.
? Hollinella sp. Coryell 1928 ostracod
 Palaeocopida - Lichviniidae
? Evlanella sp.