PBDB Collection

Gram Formation (Miocene of Denmark)

Where: Denmark (55.3° N, 9.0° E: paleocoordinates 55.5° N, 8.0° E)

• coordinate based on nearby landmark

• basin-level geographic resolution

When: Gram Formation, Tortonian (11.6 - 7.2 Ma)

• STRATIGRAPHIC RELATIONS: From the Gram Fm, which conformably overlies the Hodde Fm (Serravallian) and is unconformably overlain by Quaternary deposits. AGE: Tortonian. STRATIGRAPHIC POSITION: General list for formation.

• formation-level stratigraphic resolution

Environment/lithology: marine; poorly lithified, micaceous, silty claystone

• SPECIFIC LITHOLOGY: Silty, dark, micaceous clay. LITHIFICATION: Poorly-lithified, on the basis of figured specimens.

Size classes: macrofossils, mesofossils

Collection methods: surface (in situ),

• COLLECTORS: Unknown. REPOSITORY: Unknown.

Primary reference: F. Roth and K. Hoedemakers. 2005. The Marine Gram Formation at Gram, Denmark: Late Miocene Geology and Palaeontology. Palaeontos 7:1-189 [A. Hendy/A. Hendy]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 98312: authorized by Austin Hendy, entered by Austin Hendy on 30.09.2010

Creative Commons license: CC BY (attribution)

Taxonomic list

• COVERAGE: Exhaustive for molluscs. NOMENCLATURE: Modern nomenclature, with species-resolution identifications.

Show authors, comments, and common names

Echinoidea
Spatangoida - Brissidae	Brissopsis lyrifera² Brissopsis lyrifera² heart urchin
Chondrichthyes
Carcharhiniformes - Galeocerdidae	Galeocerdo sp.¹ Galeocerdo sp.¹ Müller and Henle 1837 tiger shark
Carcharhiniformes - Scyliorhinidae	Scyliorhinus cf. coupatezi¹ Scyliorhinus cf. coupatezi¹ Herman 1974 cat shark
Lamniformes - Odontaspididae	Odontaspis sp.¹ Odontaspis sp.¹ Agassiz 1838 sand tiger shark
Lamniformes - Otodontidae	"Carcharodon megalodon" = Otodus megalodon¹ "Carcharodon megalodon" = Otodus megalodon¹ Agassiz 1835 megalodon
Lamniformes - Cetorhinidae	Cetorhinus maximus¹ Cetorhinus maximus¹ Gunner 1765 basking shark
Lamniformes - Lamnidae	Isurus sp.¹, Isurus escheri¹, "Isurus hastalis" = Carcharodon hastalis¹ Isurus sp.¹ Rafinesque 1810 mackerel shark Isurus escheri¹ Agassiz 1843 mackerel shark "Isurus hastalis" = Carcharodon hastalis¹ Agassiz 1838 white shark
Hexanchiformes - Hexanchidae	Hexanchus primigenus¹ Hexanchus primigenus¹ sixgill shark
Rajiformes - Rajidae	Raja sp.¹ Raja sp.¹ Linnaeus 1758 skate
Squaliformes - Squalidae	Squalus cf. acanthias¹ Squalus cf. acanthias¹ Linnaeus 1758 spiny dogfish
Scaphopoda
Gadilida - Gadilidae	Polyschides weinbrechti Polyschides weinbrechti tusk shell
Dentaliida - Dentaliidae	Dentalium cf. michelotti Dentalium cf. michelotti tusk shell
Bivalvia
Nuculanida - Yoldiidae	Yoldia (Yoldia) glaberrima, Yoldiella pygmaea, Yoldiella spiegleri Yoldia (Yoldia) glaberrima von Münster 1837 clam Yoldiella pygmaea clam Yoldiella spiegleri clam
Nuculida - Nuculidae	Nucula (Lamellinucula) georgiana, Leionucula cf. laevigata Nucula (Lamellinucula) georgiana nut clam Leionucula cf. laevigata Sowerby 1818 nut clam
Arcida - Limopsidae	"Limopsis (Pectunculina) cf. anomala" = Limopsis, Limopsis (Limopsis) aurita "Limopsis (Pectunculina) cf. anomala" = Limopsis clam Limopsis (Limopsis) aurita Brocchi 1814 clam
Arcida - Glycymerididae	"Glycymeris (Chevronia) obovata" = Glycymeris (Glycymeris) obovata "Glycymeris (Chevronia) obovata" = Glycymeris (Glycymeris) obovata Lamarck 1819 clam
Pectinida - Anomiidae	"Pododesmus (Heteranomia) squamula" = Heteranomia squamula "Pododesmus (Heteranomia) squamula" = Heteranomia squamula Linnaeus 1758 scallop
Pectinida - Pectinoidae	Korobkovia woodi, Pseudamussium clavatum, Pseudamussium cf. lilli, Aequipecten seniensis Korobkovia woodi scallop Pseudamussium clavatum Poli 1795 scallop Pseudamussium cf. lilli Pusch 1837 scallop Aequipecten seniensis scallop
Ostreida - Pinnidae	Atrina sp. Atrina sp. Gray 1842 oyster
Mytilida - Mytilidae	Mytilus sp. Mytilus sp. Linnaeus 1758 mussel
Lucinida - Lucinidae	Lucinidae indet. Lucinidae indet. Fleming 1828 clam
Lucinida - Thyasiridae	Thyasira (Thyasira) flexuosa Thyasira (Thyasira) flexuosa Montagu 1803 clam
Cardiida - Lasaeidae	"Lepton transversarium" = Lepton (Lepton) transversarium "Lepton transversarium" = Lepton (Lepton) transversarium Cossmann 1896 kelly clam
Cardiida - Montacutidae	Mioerycina coarctata Mioerycina coarctata Wood 1840 clam
Cardiida - Mactridae	Spisula aff. subtruncata Spisula aff. subtruncata da Costa 1778 clam
Cardiida - Glossidae	Glossus (Glossus) olearii Glossus (Glossus) olearii clam
Cardiida - Kelliellidae	Spaniodontella nitida Spaniodontella nitida clam
Cardiida - Semelidae	Abra (Abra) cf. prismatica Abra (Abra) cf. prismatica Montagu 1808 abra
Cardiida - Cardiidae	Parvicardium straeleni Parvicardium straeleni cockle
Pholadida - Teredinidae	Teredinidae indet. Teredinidae indet. Rafinesque 1815 clam
Pholadida - Xylophagaidae	Xylophaga sp. Xylophaga sp. Turton 1822 clam
Pholadida - Corbulidae	Corbula (Varicorbula) gibba Corbula (Varicorbula) gibba Oliva 1792 clam
Hiatellida - Hiatellidae	"Hiatella (Hiatella) arctica" = Hiatella arctica "Hiatella (Hiatella) arctica" = Hiatella arctica Linnaeus 1767 clam
Poromyida - Cuspidariidae	Cuspidaria (Cuspidaria) cf. cuspidata, Cardiomya (Cardiomya) costellata Cuspidaria (Cuspidaria) cf. cuspidata clam Cardiomya (Cardiomya) costellata Deshayes 1833 clam
Thraciida - Thraciidae	Thracia (Thracia) inflata Thracia (Thracia) inflata clam
Thraciida - Periplomatidae	Periploma ariei Periploma ariei Gürs 1996 clam
Carditida - Astartidae	Goodallia (Goodallia) esbjergensis, "Carinastarte vetula" = Astarte (Carinastarte) Goodallia (Goodallia) esbjergensis clam "Carinastarte vetula" = Astarte (Carinastarte) clam
Carditida - Carditidae	Cyclocardia (Cyclocardia) orbicularis, Pteromeris corbis, Venericardia ampla, Venericardia ronchetti, Venericardia squamulosa Cyclocardia (Cyclocardia) orbicularis clam Pteromeris corbis clam Venericardia ampla clam Venericardia ronchetti clam Venericardia squamulosa clam
Gastropoda
Epitonioidea - Epitoniidae	Acirsa (Hemiacirsa) aff. duvergieri, Cirsotrema sp. Acirsa (Hemiacirsa) aff. duvergieri wentletrap Cirsotrema sp. Mörch 1852 wentletrap
Cerithioidea - Turritellidae	Turritella (Turritella) tricarinata, "Turritella (Archimediella) cochlias" = Archimediella Turritella (Turritella) tricarinata Brocchi 1814 turret shell "Turritella (Archimediella) cochlias" = Archimediella turret shell
Pseudomelanoidea - Pseudomelaniidae	Crassispira borealis Crassispira borealis snail
Sorbeoconcha - Triphoridae	Norephora (Norephora) fritschi Norephora (Norephora) fritschi snail
Sorbeoconcha - Cerithiopsidae	Krachia zelandica Krachia zelandica snail
Sorbeoconcha - Aporrhaidae	Aporrhais dingdenensis Aporrhais dingdenensis Marquet et al. 2002 snail
Sorbeoconcha - Cassidae	"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata, Galeodea bicatenata "Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata Sacco 1890 snail Galeodea bicatenata Sowerby 1815 snail
Neogastropoda - Volutidae	Scaphella bolli Scaphella bolli volute
Neogastropoda - Muricidae	"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta, Lyrotyphis (Eotyphis) sejunctus, "Trophonopsis (Pagodula) semperi" = Pagodula, Spinucella reimersi "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta Bronn 1831 murex snail Lyrotyphis (Eotyphis) sejunctus murex snail "Trophonopsis (Pagodula) semperi" = Pagodula murex snail Spinucella reimersi murex snail
Neogastropoda - Nassariidae	Amyclina sp., Nassarius cf. wienrichi, Nassarius cf. holsaticus, Nassarius cf. syltensis, Nassarius (Tritonella) ronaldjansseni, Nassarius (Uzita) prysmaticus Amyclina sp. Iredale 1918 snail Nassarius cf. wienrichi snail Nassarius cf. holsaticus snail Nassarius cf. syltensis snail Nassarius (Tritonella) ronaldjansseni snail Nassarius (Uzita) prysmaticus Brocchi 1814 snail
Neogastropoda - Fasciolariidae	"Streptolathyrus rothi" = Streptochetus (Streptolathyrus) "Streptolathyrus rothi" = Streptochetus (Streptolathyrus) snail
Neogastropoda - Buccinidae	Aquilofusus semiglaber, Aquilofusus klugorum, Aquilofusus cf. puggaardi, Colus distinctus, Liomesus ventrosus Aquilofusus semiglaber true whelk Aquilofusus klugorum true whelk Aquilofusus cf. puggaardi true whelk Colus distinctus true whelk Liomesus ventrosus true whelk
Neogastropoda - Mitridae	"Mitra grateloupi" = Mitra (Tiara) grateloupi "Mitra grateloupi" = Mitra (Tiara) grateloupi d'Orbigny 1852 mitre shell
Neogastropoda - Borsoniidae	Bathytoma (Bathytoma) cataphracta, Microdrillia serratula, Asthenotoma ornata Bathytoma (Bathytoma) cataphracta Brocchi 1814 snail Microdrillia serratula snail Asthenotoma ornata snail
Neogastropoda - Horaiclavidae	Haedropleura maitreja Haedropleura maitreja snail
Neogastropoda - Turridae	Gemmula (Gemmula) badensis, Unedogemmula annae, Acamptogenotia intorta, Fusiturris (Fusiturris) helena Gemmula (Gemmula) badensis turrid Unedogemmula annae turrid Acamptogenotia intorta turrid Fusiturris (Fusiturris) helena turrid
Neogastropoda - Raphitomidae	Teretia guersi, Pleurotomella (Pleurotomella) mioweberi, Daphnella abrahamssoni, Raphitoma spinosoreticulata, Raphitoma variccostata Teretia guersi snail Pleurotomella (Pleurotomella) mioweberi snail Daphnella abrahamssoni snail Raphitoma spinosoreticulata snail Raphitoma variccostata snail
Neogastropoda - Mangeliidae	Benthomangelia obtusangula Benthomangelia obtusangula snail
Neogastropoda - Clathurellidae	Pleurotomoides mariae Pleurotomoides mariae snail
Neogastropoda - Terebridae	Terebra forchhammeri Terebra forchhammeri auger snail
Neogastropoda - Drilliidae	Spirotropis gramensis Spirotropis gramensis Janssen 1993 snail
Neogastropoda - Conidae	"Conus (Conolithus) antediluvianus" = Conus (Conolithes), Oenopota kochi "Conus (Conolithus) antediluvianus" = Conus (Conolithes) cone shell Oenopota kochi cone shell
Neogastropoda - Costellariidae	"Vexillum (Uromitra) cimbrica" = Uromitra "Vexillum (Uromitra) cimbrica" = Uromitra ribbed mitre
Neogastropoda - Cancellariidae	Babylonella fusiformis, Narona rothi, Narona calcarata Babylonella fusiformis snail Narona rothi snail Narona calcarata snail
Sorbeoconcha - Xenophoridae	Xenophora (Stellaria) testigera Xenophora (Stellaria) testigera Bronn 1831 carrier shell
Sorbeoconcha - Ficidae	Ficus (Ficus) conditus Ficus (Ficus) conditus Brongniart 1823 fig shell
Sorbeoconcha - Naticidae	Naticarius koeneni, Euspira helicina Naticarius koeneni moon snail Euspira helicina Brocchi 1814 moon snail
Sorbeoconcha - Rissoidae	Obtusella gottscheana Obtusella gottscheana snail
Sorbeoconcha - Vitrinellidae	Circulus (Circulus) hennei Circulus (Circulus) hennei Glibert 1952 snail
Vanikoroidea - Eulimidae	Eulima (Eulima) glabra Eulima (Eulima) glabra snail
Vanikoroidea - Aclididae	Aclis minor Aclis minor aclis
Heterostropha - Ringiculidae	Ringicula (Ringiculina) buccinea Ringicula (Ringiculina) buccinea Sowerby 1823 snail
Heterostropha - Acteonidae	Acteon (Acteon) semistriatus Acteon (Acteon) semistriatus snail
Heterostropha - Tjaernoiidae	Tjaernoeia punctata Tjaernoeia punctata snail
Heterostropha - Pyramidellidae	Odostomia (Megastomia) conoidea, Odostomia (Megastomia) tuexeni, Pyramidella (Voluspa) plicosa, Eulimella (Eulimella) scillae, Eulimella (Eulimella) acicula, "Turbonilla (Strioturbonilla) rothi" = Strioturbonilla, "Turbonilla (Strioturbonilla) sp." = Strioturbonilla, Brachystomia sp., Chrysallida (Pyrgulina) interita, Chrysallida (Parthenina) semireticulata, Chrysallida (Parthenina) modesta, Syrnola (Syrnola) neumayri, Turbonilla (Turbonilla) costellata, Turbonilla (Turbonilla) facki, Kleinella (Kleinella) nordmanni Odostomia (Megastomia) conoidea Brocchi 1814 snail Odostomia (Megastomia) tuexeni snail Pyramidella (Voluspa) plicosa snail Eulimella (Eulimella) scillae snail Eulimella (Eulimella) acicula snail "Turbonilla (Strioturbonilla) rothi" = Strioturbonilla snail "Turbonilla (Strioturbonilla) sp." = Strioturbonilla Sacco 1892 snail Brachystomia sp. Monterosato 1884 snail Chrysallida (Pyrgulina) interita snail Chrysallida (Parthenina) semireticulata Sorgenfrei 1958 snail Chrysallida (Parthenina) modesta snail Syrnola (Syrnola) neumayri snail Turbonilla (Turbonilla) costellata snail Turbonilla (Turbonilla) facki snail Kleinella (Kleinella) nordmanni snail
Heterostropha - Cimidae	Actaeopyramis elatus Actaeopyramis elatus snail
Thecosomata - Limacinidae	Limacina atlanta, Limacina wilhelminae Limacina atlanta snail Limacina wilhelminae snail
Opisthobranchia - Retusidae	Retusa (Cylichnina) elongata Retusa (Cylichnina) elongata snail
Opisthobranchia - Cylichnidae	Cylichna (Cylichna) pseudoconvoluta Cylichna (Cylichna) pseudoconvoluta snail
Opisthobranchia -	Roxania (Roxania) utriculus Roxania (Roxania) utriculus snail
Heterobranchia - Diaphanidae	Toledonia flensburgensis, Diaphana moerchi Toledonia flensburgensis snail Diaphana moerchi snail
Trochoidea - Skeneidae	Skenea ariejansseni Skenea ariejansseni snail
Trochoidea - Solariellidae	Solariella (Solariella) jutensis Solariella (Solariella) jutensis snail
Lepetellida - Pseudococculinidae	Pseudococculina dittmeri, Mesopelex anderseni Pseudococculina dittmeri snail Mesopelex anderseni snail
Malacostraca
Decapoda - Munididae	Munida sp. Munida sp. Leach 1820 decapod
Decapoda - Geryonidae	Chaceon miocenicus Chaceon miocenicus Fraaije et al. 2005 crab
Decapoda - Cancridae	Tasadia carniolica Tasadia carniolica Bittner 1884 crab
Decapoda - Retroplumidae	Retropluma borealis Retropluma borealis Fraaije et al. 2005 crab
Decapoda - Nephropidae	Nephrops kvistgaardae, Homarus cf. gammarus Nephrops kvistgaardae Fraaije et al. 2005 clawed lobster Homarus cf. gammarus Linnaeus 1758 clawed lobster

¹Bendix-Almgreen 1983; ²Hansen and Hansen 2005