Guanling, Lower Xiaowa Formation fossil lagerst├Ątte (Triassic of China)

Where: Guizhou, China (25.9° N, 105.4° E: paleocoordinates 26.9° N, 87.4° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Lower Member (Xiaowa Formation), Tuvalian (232.0 - 221.5 Ma)

• 12 m thick, previously known as Wayao Member of Falang Formation or as Wayao Formation

•JA: originally entered as Carnian; the Metapolygnathus polygnatiformis and M. nodosus zones are both Tuvalian according to Lucas et al. 2012

• group of beds-level stratigraphic resolution

Environment/lithology: deep-water; lithified shale and lithified marl

• deep and quiet water, the lower unit with generally oxygenated bottom water; the upper unit shaly interval with anoxic conditions and the greatest water depth
• lower unit (5 m thick) of medium- to thick-bedded, grey bioclastic micrites with yellow-green shale interbeds; upper unit (7 m thick) of greyish to black, thin- to medium-bedded laminated mudstone with argillaceous limestone interbeds and marlstones

Size class: macrofossils

Collection methods: quarrying, mechanical,

Primary reference: C. Li and O. Rieppel. 2002. A new cyamodontid placodont from Triassic of Guizhou, China. Chinese Science Bulletin 47(5):403-407 [M. Carrano/M. Carrano]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 85166: authorized by Wolfgang Kiessling, entered by Uta Merkel on 04.12.2008, edited by Bethany Allen

Creative Commons license: CC BY (attribution)

Taxonomic list

 Ceratitida - Tibetitidae
 Ceratitida - Clionitidae
"Clionites cf. zeilleri" = Clionitites2
"Clionites cf. zeilleri" = Clionitites2 ceratite
 Ceratitida - Trachyceratidae
Protrachyceras costulatum2 Mansuy 1912 ceratite
Protrachyceras ladinum2 Mojsisovics 1882 ceratite
Protrachyceras deprati2 Mansuy 1912 ceratite
Paratrachyceras cf. hofmanni2 Boekh 1873 ceratite
Trachyceras cf. aon2 Munster 1834 ceratite
Trachyceras multituberculatum2 Hsu 1940 ceratite
 Ostreida - Halobiidae
Halobia subcomata2 Kittl 1912 oyster
 Ostreida - Bakevelliidae
Angustella sp.2 Waagen 1907 oyster
 Ostreida - Pergamidiidae
Krumbeckiella sp.2 Ichikawa 1958 oyster
 Craniida - Craniidae
? Crania sp.2 Retzius 1781
 Ozarkodinida - Gondolellidae
"Metapolygnathus nodosus" = Hayashiella nodosa2, "Metapolygnathus polygnathiformis" = Paragondolella polygnathiformis2
"Metapolygnathus nodosus" = Hayashiella nodosa2 Hayashi 1968 conodont
"Metapolygnathus polygnathiformis" = Paragondolella polygnathiformis2 Budurov and Stefanov 1965 conodont
 Testudines -
Odontochelys semitestacea n. gen. n. sp.1
Odontochelys semitestacea n. gen. n. sp.1 Li et al. 2008 turtle
IVPP V 13240 (paratype), V 15639 (holotype), V 15653
 Thalattosauria - Askeptosauridae
 Thalattosauria -
Xinpusaurus suni2 Yin 2000 diapsid
Xinpusaurus bamaolinensis2 Cheng 2003 diapsid
 Placodontia - Cyamodontidae
Sinocyamodus xinpuensis2 Li 2000 placodont
Cyamodus orientalis n. sp.3 Wang et al. 2019 placodont
Zhejiang Museum of Natural History, Hangzhou, ZMNH M8820 (holotype), "an articulated and nearly complete skeleton with three-dimensionally preserved skull and lacking only the forelimbs, representing an adult individual based on overall body size"
 Placodontia - Placochelyidae
Psephochelys polyosteoderma n. gen. n. sp. Li and Rieppel 2002 placodont
IVPP V 12442
 Ichthyosauria - Toretocnemidae
Qianichthyosaurus zhoui2 Li 1999 ichthyosaur
 Ichthyosauria - Shastasauridae
"Ghuizhouichthyosaurus tangae" = Guizhouichthyosaurus tangae2, "Guanlingichthyosaurus liangae" = Guanlingsaurus liangae2
"Ghuizhouichthyosaurus tangae" = Guizhouichthyosaurus tangae2 Cao et al. 2000 ichthyosaur
"Guanlingichthyosaurus liangae" = Guanlingsaurus liangae2 Yin et al. 2000 ichthyosaur
Holothuroidea indet.2 de Blainville 1834 sea cucumber
 Roveacrinida - Somphocrinidae
Osteocrinus cf. virgatus2 Kristan-Tollmann 1970 Sea lily
Osteocrinus cf. spinosus2 Kristan-Tollmann 1970 Sea lily
 Encrinida - Traumatocrinidae
Traumatocrinus sp.2 Woehrmann 1889 Sea lily