Gratkorn locality (Miocene of Austria)

Where: Austria (47.1° N, 15.3° E: paleocoordinates 46.7° N, 15.1° E)

• coordinate stated in text

• outcrop-level geographic resolution

When: Porosononion granosum foram zone, Sarmatian (12.7 - 11.6 Ma)

• Lower Sarmatian

•Upper Ervilia Zone (mollusc zones) / C5An.1r

Environment/lithology: lagoonal or restricted shallow subtidal; mudstone

• pelite

Size classes: macrofossils, mesofossils

Primary reference: M. Harzhauser, M. Gross, and H. Binder. 2008. Biostratigraphy of Middle Miocene (Sarmatian) wetland systems in an Eastern Alpine intramontane basin (Gratkorn Basin, Austria): the terrestrial gastropod approach. Geologica Carpathica 59(1):45-58 [W. Kiessling/M. Krause]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 175587: authorized by Wolfgang Kiessling, entered by Mihaela Krause on 17.12.2015, edited by Terri Cleary

Creative Commons license: CC BY (attribution)

Taxonomic list

Gastropoda
 Architaenioglossa - Aciculidae
Platyla callosiuscula Andreae 1904 snail
 Rissooidea - Bithyniidae
Bithynia sp. Leach 1818 snail
 Pulmonata - Vertiginidae
Negulus gracilis Gottschick and Wenz 1919 pulmonate
 Pulmonata - Clausiliidae
Pseudidyla martingrossi Harzhauser and Binder 2008 slug
 Pulmonata - Gastrocoptidae
 Pulmonata - Punctidae
Punctum parvulum Gottschick 1920 slug
 Pulmonata - Pupillidae
Pupilla iratiana Dupuy 1850 slug
Vertigo angulifera Boettger 1884 slug
 Pulmonata - Valloniidae
 Pulmonata - Endodontidae
 Pulmonata - Zonitidae
Nesovitrea boettgeriana Clessin 1877 true glass snail
 Limacoidea - Limacidae
Limax sp. Linnaeus 1758 slug
 Limacoidea - Vitrinidae
 Pulmonata - Testacellidae
Testacella schuetti Schlickum 1967 slug
 Pulmonata - Camaenidae
Pleurodonte michalkovaci Binder and Harzhauser 2008 slug
 Basommatophora - Planorbidae
Gyraulus vermicularis Stolickza 1862 pulmonate
Actinopteri
 Gobiiformes - Gobiidae
Gobiidae indet. Bonaparte 1832 goby
GPIT/OS/00800, scale
 Cypriniformes - Cyprinidae
Leuciscinae indet. Howes 1991 carp-like fish
UMJGP GR 3-79, 1 pharyngeal tooth
Osteichthyes
 Salientia - Pelobatidae
Pelobates sanchizi spadefoot toad
UMJGP 204.708, 1 frontoparietal; GPIT/AM/00728, GPIT/AM/ 00729, UMJGP 204.692, 2 right and 3 left fragments of maxilla; UMJGP 204.692, 1 right premaxilla; UMJGP 204.692, 1 sphenethmoid; GPIT/AM/00729, UMJGP 204.692, 1 right and 1 left squamosals; UMJGP 204.692, 1 scapula; UMJGP 204.692, 5 trunk verts; UMJGP 204.692, UMJGP 204.731, 2 sacral verts
 Salientia -
Latonia sp. von Meyer 1843 frog
GPIT/AM/00727, 1 left and right premaxilla, 1 left and right maxilla, 1 atlas, 2 verts, 1 urostyle
 Salientia - Ranidae
Rana sp. Linnaeus 1758 frog
GPIT/AM/00732. 1 right ilium
Pelophylax sp. Fitzinger 1843 frog
GPIT/AM/00731, 1 right ilium
 Salientia - Bufonidae
Bufo cf. viridis Laurenti 1768 European green toad
GPIT/AM/00726, 2 right ilia, 1 right and 1 left scapula, 1 right frontoparietal
 Caudata - Urodela
Chelotriton aff. paradoxus Pomel 1853 newt
UMJGP 204.730, partial skeleton with 1 frontal, 2 trunk verts, 6 ribs, and fragmentary limb bones
Triturus aff. vulgaris Linnaeus 1758 smooth newt
GPIT/AM/00724, 1 right maxilla, 1 left otoccipital, 5 trunk verts, 1 femur
Salamandra sansaniensis Lartet 1851 salamander
GPIT/AM/ 00725, 1 left and right dentaries, 11 fragmentary to complete trunk verts, 5 ribs
Reptilia
 Testudines - Panchelydridae
Chelydropsis murchisoni Bell 1836 turtle
UMJGP 204.725, 1 fragmentary left 10th peripheral
 Testudines - Pantestudinidae
Testudo cf. steinheimensis turtle
UMJPG 204.690, 1 complete anterior plastron and associated partial carapace; UMJPG 204.672, 1 fragmentary anterior plastron; UMJPG 204.108 and 203.661, 2 left hyoplastra
Testudo cf. kalksburgensis Toula 1896 turtle
UMJPG 204.113, 1 fragmentary anterior plastron and associated partial carapace
 Testudines - Pangeoemydidae
Clemmydopsis turnauensis Meyer 1847 turtle
UMJPG 204.057-1, 1 isolated neural I; UMJPG 204.726-1 and 2, 2 fragmentary pleurals
 Squamata - Lacertidae
Lacertidae indet. Bonaparte 1831 lizard
UMJGP 204.750, 1 parietal
sensu lato Lacerta sp. Linnaeus 1758 lizard
GPIT/RE/09235, fragmentary preserved 20 left and 17 right dentaries, 16 left and 18 right maxillae, 3 premaxillae, 5 jaw bones; UMJGP 204.134, 204.147, 2 left dentaries; UMJGP 204.131, 1 right dentary; UMJGP 204.132, 204.136, 2 left maxillae; UMJGP 204.135, right maxilla
sensu lato Lacerta "sp. 3" Linnaeus 1758 lizard
UMJGP 204.107, 1 left dentary
sensu lato Lacerta "sp. 2" Linnaeus 1758 lizard
GPIT/RE/09230, GPIT/RE/09231, 2 left dentaries
sensu lato Lacerta "sp. 1" Linnaeus 1758 lizard
UMJGP 204.131, 2014.133, GPIT/RE/09226–GPIT/RE/09229, GPIT/RE/ 09260, 1 left and 6 right dentaries
Miolacerta tenuis Roček 1984 lizard
UMJGP 204.106, 204.031, GPIT/RE/ 09232−/09234, GPIT/RE/09262, GPIT/RE/09263, 4 left and 4 right dentaries
Edlartetia sp. Augé and Rage 2000 lizard
UMJGP 204.752, right maxilla; GPIT/RE/ 09225, left maxilla
 Squamata - Varanidae
Varanus sp. Merrem 1820 monitor lizard
UMJGP 210.905, 1 right dentary; GPIT/RE/09236, 1 fragmentary left maxilla; UMJGP 203.732, 204.733, 203.457, GPIT/RE/09261, 5 trunk vertebrae; UMJGP 203.457, 3 caudal verts; UMJGP 203.457, 5 ribs
 Squamata - Anguidae
Ophisaurus spinari Klembara 1979 glass lizard
UMJGP 204.728, partial skeleton including 1 premaxilla, 1 quadrate, 5 trunk verts, 2 sacral verts, osteoderms; UMJGP 204.037, 204.137, 204.749, 204.750, 4 parietals; UMJGP 204.032, 1 left maxilla; UMJGP 204.130, 204.728, osteoderms; GPIT/RE/09253, 1 palatine, 1 left maxilla, 1 right dentary, 32 trunk verts; UMJGP 204.122–204.124, 204.128, 204.129, GPIT/RE/09252, GPIT/RE/09253, numerous osteoderms
 Squamata - Elapidae
Naja sp. Laurenti 1768 cobra
UMJGP 203.430 and GPIT/RE/ 09255, 2 trunk verts
 Squamata - Colubridae
Colubrinae "sp. 2" Oppel 1811 colubrid snake
GPIT/RE/09258, 2 trunk verts
Colubrinae "sp. 1" Oppel 1811 colubrid snake
UMJGP 204.041, 3 trunk verts; GPIT/RE/09257, 3 isolated trunk verts
Natricinae sp. Bonaparte 1838 colubrid snake
GPIT/RE/09256, 2 trunk verts
 Squamata - Gekkonidae
Gekkonidae indet. Gray 1825 gecko
GPIT/RE/09224, 1 left dentary
 Squamata - Scincidae
Scincidae indet. Gray 1825 skink
UMJGP 204.748, GPIT/RE/09264–GPIT/RE/09266, 3 left and 1 right dentaries; UMJGP 204.039, GPIT/RE/09267, 1 right and 1 left maxillae