Kohfidisch (Miocene of Austria)

Where: Burgenland, Austria (47.1° N, 16.3° E: paleocoordinates 46.9° N, 16.2° E)

• coordinate stated in text

When: MN 11 (8.7 - 7.8 Ma)

• Locality considered to be in MN 11 biozone

Environment/lithology: terrestrial

Size classes: macrofossils, mesofossils

Collection methods: surface (float), surface (in situ), mechanical,

Primary reference: F. Bachmayer and Z. Szyndlar. 1985. Ophidians (Reptilia: Serpentes) from the Kohfidisch fissures of Burgenland, Austria. Annalen des Naturhistorischen Museums in Wien, Serie A: Mineralogie und Petrographie, Geologie und Palaeontologie, Anthropologie Praehistorie 87:79-100 [J. Head/J. Head/L. van den Hoek Ostende]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 37789: authorized by Jason Head, entered by Jason Head on 23.03.2004, edited by Lars van den Hoek Ostende, David Nicholson, Terri Cleary and Torsten Liebrecht

Creative Commons license: CC BY (attribution)

Taxonomic list

Gastropoda
 Heterostropha - Planorbidae
"Planorbis (Anisus) confusus" = Anisus confusus8
"Planorbis (Anisus) confusus" = Anisus confusus8 Soos 1934 snail
 Heterostropha - Carychiidae
 Stylommatophora - Helicidae
Tacheocampylaea (Mesodontopsis) doderleini8 Brusina 1897 slug
mispelled in text as "Tacheocampylea"
Bivalvia
 Cardiida - Dreissenidae
Insecta
 Lepidoptera - Tineidae
Tinea sp.8 Linnaeus 1758 moth
Osteichthyes
 Teleosti -
"Teleosti indet." = Teleostei8
"Teleosti indet." = Teleostei8 Müller 1846 ray-finned fish
Amphibia
 Salientia -
Anura indet.8 Fischer von Waldheim 1813 frog
 Salientia - Discoglossidae
Bombina cf. bombina8 Linnaeus 1761 European fire-bellied toad
1 sphenethmoideum(??) (AO: Inv. Nr.: NHMW 2004z0014/0003). 1 presacral vertebra, 1 coccyx (AO: Inv. Nr.: NHMW2004z0014/0001). 1 right ilium (AO: Inv. Nr.: NHMW2004z0014/0002). 1 right (female??) humerus (BM: Inv. Nr.: NHMW2004z0014/0005). Again, translating from Latin, sorry.
 Salientia - Bufonidae
Bufo bufo8 Linnaeus 1758 toad
3 sphenethmoidea (AO: Inv. Nr.: NHMW2004 zOO 19/0005). 6 right prearticulars, 1 left prearticular, 4 atlases (AO: Inv. Nr.: NHMW2004z0019/0002). 222 presacral vertebrae, 53 sacral vertebrae, 13 coccyges. 59 right ilia. 59 left ilia. (AO: Inv. Nr.: NHMW2004z0019/0003, 0006). 8 ischia (AO: Inv. Nr.: NHMW2004z0019/0004). 28 right scapulae. 19 left scapulae. 13 coracoids (AO: Inv. Nr.: NHMW2004z0019 /0001). 57 female(?) right humeri. 33 right male(?) humeri. 58 left female(?) humeri. 34 left male(?) humeri. 1 humerus indet. 34 right radioulnare. 31 left radioulnare. 28 radioulnare indet. (BM: Inv. Nr.: NHMW2004z0019/0007).
 Salientia - Ranidae
Rana "ridibunda sp."8 Linnaeus 1758 frog
1 right fronto-parietal (AO: Inv. Nr.: NHMW 2004z0021/0003). 8 right premaxillae, 5 left premaxillae, 1 indet. premaxilla, 35 right maxillaries, 31 left maxillaries, 14 indet. maxillaries, 6 right prearticulars. 6 left prearticulars. 1 atlas. 65 presacral vertebrae (AO: Inv. Nr.: NHMW2004z0021/0001). 4 presacral vertebrae with connection to sacrals, 32 sacral vertebrae, 11 coccyges, 19 right ilia. (AO: Inv. Nr.: NHMW2004z0021/0002). 16 left ilia. (AO: Inv. Nr.: NHMW2004z0021/0004,0005). 18 ischia, 11 coracoids, 35 right female? humeri, 28 right male? humeri, 22 left female humeri, 28 left male? humeri, 8 left male? humeri, 15 right radio-ulnare, 11 left radio-ulnare, 4 indet. radio-ulnare
 Salientia - Hylidae
Hyla sp.8 Laurenti 1768 tree frog
2 right maxillae. 1 presacral vertebra (AO: Inv. Nr.: NHMW2004z0020/0001). 8 right ilia. (AO: Inv. Nr.: NHMW2004z0020/0004). 8 left ilia. (AO: Inv. Nr.: NHMW2004z0020/0002). 1 ischium (AO: Inv. Nr.: NHM- W2004z0020/0003). 4 right humeri. 2 left humeri. (BM: Inv. Nr.: NHMW2004z0020/0005).
 Salientia - Pelobatidae
Pelobates cf. cultripes8 Tschudi 1838 spadefoot toad
110 fragmentary fronto-parietals, 10 right squamosals, 3 left squamosals (AO: Inv. Nr.: NHM- W2004z0018/0001). 1 right nasal, 5 right premaxillae, 6 left premaxillae, 55 right maxillae, 38 left maxillae, 10 indet. maxillae, 1 left prearticular, 1 atlas, 19 presacral vertebrae, 3 sacral vertebrae, 13 right ilia (AO: Inv. Nr.: NHMW2004z0018/0002, 0003). 10 left ilia, 1 ischium, 1 right scapula, 10 right humerii, 3 left humerii, 1 left radioulnare (BM: Inv. Nr.: NHM- W2004z0018/0004).
 Salientia -
Latonia sp.8 von Meyer 1843 frog
56 fragmentary fronto-parietals, 2 right prootic-occipitals, 1 left prootic-occipital, 7 right premaxillae (AO: Inv. Nr.: NHMW2004z0017/0001). 8 left premaxillae, 34 right maxillae, 28 left maxillae, 45 indet. maxillae, 33 right prearticulars, 22 left prearticulars, 21 atlases, 210 presacral vertebrae, 52 sacral vertebrae, 1 costal, 59 coccyges, 47 right ilia (AO: Inv. Nr.: NHMW2004z0017/0002). 46 left ilia (AO: Inv. Nr.: NHMW2004z0017/0004). 3 ischia (AO: Inv. Nr.: NHMW2004z0017/0003). 17 right scapulae, 18 left scapulae, 24 coracoids, 17 right humerii, 27 left humerii, 22 right radioulnare, 13 left radioulnare, 2 indet. radioulnare (BM: Inv. Nr.: NHMW2004z0017/0005).
Latonia gigantea8 Lartet 1851 frog
23 right maxillae, 23 left maxillae (BM: Inv. Nr.: NHMW2004z0015/0001).
Latonia ragei8 Hossini 1993 frog
7 right maxillae, 5 left maxillae
 Caudata - Urodela
Triturus "cristatus sp."8 Rafinesque 1815 newt
170 presacral vertebrae (AO: Inv.Nr.: NHM W2004z0011/0001).
Triturus "vulgaris sp."8 Rafinesque 1815 newt
5 atlases, 46 presacral vertebrae (AO: Inv. Nr.: NHMW2004z0012/0001). 1 left humerus (of a female?). (AO: Inv. Nr.: NHMW2004z0012/0002). (BM: Inv. Nr.: NHMW2004z0012/0003). The specimen numberings in this paper do not make sense and half is in Latin, sorry.
Triturus sp.8 Rafinesque 1815 newt
65 presacral vertebrae, 19 caudal vertebrae (BM: Inv. Nr.: NHMW2004z0013/0001).
Chelotriton paradoxus8 Pomel 1853 newt
1 right squamosal, 3 atlases (AO: Inv. Nr.: NHMW2004z0010/0001). 19 presacral vertebrae (BM: Inv. Nr.: NHMW2004z0010/0002).
Mioproteus caucasicus8 Estes and Darevsky 1978 mudpuppy
1 left dentary (AO: Inv. Nr.: NHMW2004z0009/ 0001); 2 atlases, 30 presacral vertebrae (AO: Inv. Nr.: NHMW2004z0009/0002). (BM: Inv. Nr.: NHMW 2004z0009/0003).
Mammalia
 Rodentia - Hystricidae
 Rodentia - Gliridae
Vasseuromys pannonicus8 Kretzoi 1978 dormouse
Graphiurops austriacus n. sp.4 Bachmayer and Wilson 1980 dormouse
Myomimus cf. multicristatus3 dormouse
pro partim ex. Peridyromys compositus Bachmayer & Wilson, 1970
Myomimus cf. dehmi3 de Bruijn 1966 dormouse
Glis cf. minor8 Kowalski 1963 dormouse
Muscardinus austriacus4 Bachmayer and Wilson 1970 dormouse
n. ssp. austriacus
 Rodentia - Sciuridae
Spermophilinus cf. bredai2 von Meyer 1848 squirrel
 Rodentia - Eomyidae
Keramidomys sp.3 Hartenberger 1966 rodent
Leptodontomys sp.4 Shotwell 1956 rodent
 Rodentia - Castoridae
 Rodentia - Zapodidae
Protozapus intermedius n. gen. n. sp.2
Protozapus intermedius n. gen. n. sp.2 Bachmayer and Wilson 1970 jumping mouse
 Rodentia - Muridae
Progonomys woelferi n. sp.2 Bachmayer and Wilson 1970 mouse
Epimeriones austriacus4 Daxner-Höck 1972 gerbil
Parapodemus lugdunensis3 Schaub 1938 mouse
 Rodentia - Cricetidae
Neocricetodon sp.5, "Kowalskia fahlbuschi n. sp." = Neocricetodon fahlbuschi2, Prospalax petteri n. sp.2, Promimomys sp.3
Neocricetodon sp.5 Schaub 1934 mouse
"Kowalskia fahlbuschi n. sp." = Neocricetodon fahlbuschi2 Bachmayer and Wilson 1970 mouse
Prospalax petteri n. sp.2 Bachmayer and Wilson 1970 mouse
Promimomys sp.3 Kretzoi 1955 mouse
subgenus Prosomys
 Lagomorpha - Ochotonidae
Prolagus cf. oeningensis2 König 1825 pika
 Theriamorpha - Erinaceidae
"Galerix zapfei n. sp." = Schizogalerix zapfei2, "Schizogalerix zapfei" = Schizogalerix zapfei8, "Schizogalerix cf. moedlingensis" = Schizogalerix moedlingensis8, "Galerix cf. moedlingensis" = Schizogalerix moedlingensis4, ? Erinaceus sp.2, Lantanotherium sp.3
"Galerix zapfei n. sp." = Schizogalerix zapfei2 Bachmayer and Wilson 1970 hedgehog
"Schizogalerix zapfei" = Schizogalerix zapfei8 Bachmayer and Wilson 1970 hedgehog
"Schizogalerix cf. moedlingensis" = Schizogalerix moedlingensis8 Rabeder 1973 hedgehog
"Galerix cf. moedlingensis" = Schizogalerix moedlingensis4 Rabeder 1973 hedgehog
? Erinaceus sp.2 Linnaeus 1758 hedgehog
Lantanotherium sp.3 Filhol 1888 hedgehog
incorrectly written as Lanthanotherium
 Theriamorpha - Soricidae
Petenyia dubia8, ? Neomyini indet.3, "? Petenyiella repenningi n. sp." = Paenelimnoecus repenningi2, "Anourosorex kormosi n. sp." = Crusafontina kormosi2
Petenyia dubia8 Bachmayer and Wilson 1970 red-toothed shrew
(=”Blarinellia”)
? Neomyini indet.3 Matschie 1909 red-toothed shrew
"? Petenyiella repenningi n. sp." = Paenelimnoecus repenningi2 Bachmayer and Wilson 1970 red-toothed shrew
"Anourosorex kormosi n. sp." = Crusafontina kormosi2 Bachmayer and Wilson 1970 red-toothed shrew
 Theriamorpha - Talpidae
Desmanella cf. crusafonti3 Rümke 1974 mole
"Dibolia vinea" = Archaeodesmana vinea7 Storch 1978 mole
Talpa gilothi4 Storch 1978 mole
 Chiroptera - Rhinolophidae
Rhinolophus grivensis3 Depéret 1892 horseshoe bat
Rhinolophus delphinensis2 Gaillard 1899 horseshoe bat
 Chiroptera - Megadermatidae
Megaderma vireti2 Mein 1965 false vampire
 Chiroptera - Vespertilionidae
Plecotus sp.3 Geoffroy 1818 vesper bat
subgenus Paraplecotus
cf. Myotis sp.3 Kaup 1829 vesper bat
Myotis aff. boyeri3 Mein 1965 vesper bat
 Perissodactyla - Rhinocerotidae
Aceratherium sp.8 Kaup 1834 rhinoceros
 Perissodactyla - Equidae
Hipparion sp.8 de Christol 1832 hipparionine horse
 Artiodactyla - Cervidae
Cervidae indet.8 Gray 1821 deer
 Artiodactyla - Bovidae
Bovidae indet.8 Gray 1821 antelope
 Carnivora - Percrocutidae
Percrocuta eximia8 carnivoran
 Carnivora - Hyaenidae
Ictitherium sp.8 Wagner 1848 hyaena
 Proboscidea -
Proboscidea indet.8 Illiger 1811 proboscidean
 Proboscidea - Deinotheriidae
Deinotherium giganteum8 Kaup 1829 deinothere
Reptilia
 Testudines - Testudinidae
Testudinidae indet.8 Batsch 1788 turtle
 Testudines - Geoemydidae
Mauremys aff. gaudryi6 Depéret 1885 pond turtle
 Squamata -
Serpentes indet.8 Linnaeus 1758 snake
19 fragmentary skulls. 1 premaxilla. 1 quadrate. 16 indet. fragmentary maxillae. 67 fragmentary dentaries. 3 teeth. 5 atlases. 59 cervical vertebrae. 6016 presacral vertebrae. 5925 caudal vertebrae. 4 articulated caudal vertebrae. 107 costae.
 Squamata - Typhlopidae
Typhlops cf. grivensis8 blind snake
27 presacral vertebrae (AO: Inv. Nr.: NHMW 2004z0034/0001). 4 associated caudal vertebrae (=1 “os coccygis”; AO: Inv. Nr.: NHMW2004z0034/0002).
 Squamata - Elapidae
Naja romani8 Hoffstetter 1939 cobra
Holotype: basiparasphenoid (NMW 1984/98); referred specimens: 2 frontals, 4 maxillae, "several fragments" of dentaries and pterygoids (mainly catalogued as NMW 1984/104), and about 1000 vertebrae + 3 left and 2 right compound bones, 1 left palatine incl. a tiny fragment of right palatine, and 3 fragmentary dentaries (listed in Bachmayer & Szyndlar, 1987; catalogued as NMW 1986/7)
 Squamata - Crotalidae
Vipera burgenlandica n. sp.1 Bachmayer and Szyndlar 1987 pit viper
"= Daboia sp."; 1 basioccipital and 71 precaudal vertebrae (NMW 1984/106)
 Squamata - Colubridae
Elaphe kohfidischi n. sp., "Nanus planicarinatus n. gen. n. sp." = Coluber planicarinatus, "Coluber planicarinatus" = Coluber planicarinatus8, Natrix longivertebrata
Elaphe kohfidischi n. sp. Bachmayer and Szyndlar 1985 colubrid snake
Holotype: mid-trunk vertebra (NMW 1984/96); referred specimens: "several hundred" vertebrae (NMW 1984/101) + 10 cranial elements (listed in Bachmayer & Szyndlar, 1987: 3 basiparasphenoids,1 fragmentary parietal, 2 basioccipitals,1 fragmentary right dentary, 1 fragmentary left dentary, and 2 fragments of right maxilla(e), apparently all catalogued as NMW 1986/4)
"Nanus planicarinatus n. gen. n. sp." = Coluber planicarinatus Bachmayer and Szyndlar 1985 racer
Holotype: trunk vertebra (NMW 1984/97); referred specimens: 6 further trunk vertebrae (NMW 1984/102) + 1 basiparasphenoid (listed in Bachmayer & Szyndlar, 1987; catalogued as NMW 1986/5)
"Coluber planicarinatus" = Coluber planicarinatus8 Bachmayer and Szyndlar 1985 racer
Natrix longivertebrata Szyndlar 1984 water snake
15 precaudal vertebrae (NMW 1984/103) + 1 complete parietal (listed in Bachmayer & Szyndlar, 1987; catalogued as NMW NMW 1986/6)
 Squamata - Lacertidae
Edlartetia sansaniensis8 Augé and Rage 2000 squamates
9 right maxillae. 3 left maxillae. 12 right dentaries. (AO: Inv. Nr.: NHMW2004z0028/0001). 12 left dentaries. (BM: Inv. Nr.: NHMW2004z0028/0002).
Lacerta "sp. A"8 Linnaeus 1758 squamates
13 osteoderms. 17 supraocular osteoderms. 136 fragmentary frontals. 1 right pterygoid. (AO: Inv. Nr.: NHMW2004z0025/0001). 1 left pterygoid. 3 right quadrates. 5 left quadrates. 60 premaxillae (AO: Inv. Nr.: NHMW2004z0025/0002). 403 right maxillae. 322 left maxillae. 99 indet. maxillae. 208 right dentaries. 236 left dentaries. (AO: Inv. Nr.: NHMW2004z0025/0003). 1 atlas (AO: Inv. Nr.: NHMW2004z0025/0007). 2 epistrophei(?). 7 cervical vertebrae (AO: Inv. Nr.: NH- MW2004z0025/0004). 319 presacral vertebrae (AO: Inv. Nr.: NHMW2004z0025/0005). 2 sacral vertebrae. 96 anterior caudal vertebrae. 165 caudal vertebrae (AO: Inv. Nr.: NHMW2004z0025/0006). 2 right ilia. 5 left ilia. 6 right humeri. 4 left humeri.
Lacerta "sp. B"8 Linnaeus 1758 squamates
2 premaxillae. 12 right maxillae. 9 left maxillae. 16 right dentaries. 22 left dentaries. (AO: Inv. Nr.: NHMW2004z0026/0001). 8 presacral vertebrae. 2 caudal vertebrae
Lacerta cf. viridis8 Laurenti 1768 European green lizard
4 parietals (AO: Inv. Nr.: NHMW2004 z0024/0001). 1 neurocranium. 5 right quadrates. 3 left quadrates
Miolacerta tenuis8 Roček 1984 squamates
2 left maxillae, 3 right dentaries, (AO: Inv. Nr.: NHMW2004z0027/0001 ). 1 indet. dentary
 Squamata - Scincidae
Scincidae indet.8 Gray 1825 skink
7 right maxillae. 5 left maxillae. 1 right dentary. (AO: Inv. Nr.: NHMW2004z0029/0001). 2 left dentaries. 3 presacral vertebrae
 Squamata - Gekkonidae
Gekkonidae indet.8 Gray 1825 gecko
2 left dentaries. 1 caudal vertebra (AO: Inv. Nr.: NHM W2004z0023/0001).
 Squamata -
Lacertilia indet.8 Owen 1842 lizard
8 indet. maxillae, 6 indet. dentaries, 11 claws(?). (BM: Inv. Nr.: NHMW2004z0033/0001).
 Squamata - Varanidae
Varanus cf. hofmanni8 monitor lizard
2 right dentaries. (AO: Inv. Nr.: NHMW 2004 z0032/0003). 4 left dentaries. 5 teeth (AO: Inv. Nr.: NHMW2004z0032/0001, 0002). 3 cervical vertebrae (AO: Inv.Nr.: NHMW2004z0032/0005). 38 presacral vertebrae (AO: Inv.Nr.: NHMW2004 z0032/0004). 26 caudal vertebrae (AO: Inv. Nr.: NHMW 2004z0032/0006).
 Squamata - Anguidae
Anguis fragilis8, "Ophisaurus pannonicus" = Pseudopus pannonicus8, "Pseudopus pannonicus" = Pseudopus pannonicus8
Anguis fragilis8 Linneaus 1758 Slow worm
842 osteoderms. 1 right quadrate. (AO: Inv. Nr.: NHMW2004z0030/0006). 2 right maxillae. 11 left maxillae. 11 right dentaries. 14 left dentaries. (AO: Inv. Nr.: NHMW2004z0030/0001). 1 cervical vertebra (AO: Inv. Nr.: NHMW2004z0030/0002). 554 presacral vertebrae (AO: Inv. Nr.: NHMW2004z0030/0003). 5 sacral vertebrae. 253 caudal vertebrae (AO: Inv. Nr.: NHMW2004z0030/0004, 0005).
"Ophisaurus pannonicus" = Pseudopus pannonicus8 Kormos 1911 legless lizard
"Pseudopus pannonicus" = Pseudopus pannonicus8 Kormos 1911 legless lizard
11276 osteoderms (AO: Inv. Nr.: NHMW 2004z0031/0008). 4 right frontals. 6 left frontals. 34 parietals and fragmentary parietals (AO: Inv. Nr.: NHMW2004z0031/0001). 1 right palatine. (AO: Inv. Nr.: NHMW2004z0031/0002). 1 left palatine. 3 right pterygoids. 4 left pterygoids. 5 premaxillae (AO: Inv. Nr.: NHMW2004z0031/0003). 7 right maxillae. 15 left maxillae. (AO: Inv. Nr.: NHMW2004z 0031/0004). 13 indet. maxillae. 17 right dentaries. (AO: Inv. Nr.: NHMW2004z0031/0005). 14 left dentaries. 140 presacral vertebrae (AO: Inv. Nr.: NHMW2004z0031/0006). 3 sacral vertebrae. 283 caudal vertebrae (AO: Inv. Nr.: NHMW2004z0031/0007).
Actinopteri
 Gobiiformes - Gobiidae
Gobiidae indet.8 Bonaparte 1832 goby
 Siluriformes - Siluridae
Silurus sp.8 Linnaeus 1758 catfish