Cernay (Paleocene of France)
Also known as Cernay-lès-Reims; carrières Lemoine; Lemoine Quarry
Where: Champagne-Ardenne, France (49.3° N, 4.1° E: paleocoordinates 43.8° N, 1.7° E)
• coordinate based on nearby landmark
• small collection-level geographic resolution
When: Conglomérat de Cernay Formation, Thanetian (59.2 - 56.0 Ma)
• "Thanétien supérieur"
• group of beds-level stratigraphic resolution
Environment/lithology: terrestrial; lithology not reported
Size class: macrofossils
Reposited in the MNHN
• Lemoine's collections are from multiple nearby quarries
Primary reference: D. E. Russell. 1981. Un primate nouveau du Paléocène supérieur de France. Geobios 14(3):399-405 [J. Alroy/J. Alroy]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 87925: authorized by John Alroy, entered by John Alroy on 28.03.2009, edited by Philip Mannion, Jonathan Tennant, Terri Cleary, Patricia Holroyd and Mark Uhen
Creative Commons license: CC BY (attribution)
Taxonomic list
• remaining fauna not discussed
Show authors, comments, and common names
| Amphibia | |
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Neobatrachia indet.17 Reig 1958 frog MNHN CR 6752 (vertebra), 6753 (right radio-ulna) and 5391 (left humerus)
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Bufonidae indet.20 Gray 1825 toad MNHN CR 16016 and MNHN CR 16017 (two ilia); MNHN CR 16018 (pterygoid)
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Palaeobatrachus sp.20 Tschudi 1839 frog MNHN CR 6748 (ilium), 6749 (vertebra) and 6750 (left humerus)
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Palaeoproteus gallicus n. sp.5
Palaeoproteus gallicus n. sp.5 Estes et al. 1967 salamander MNHN CR 6692 - type (trunk vertebra); referred material: MNHN CR 6688-6691, 6702-6704 (atlantes), 6693, 6694, 6713-6717 (trunk vertebrae), 6695-6699, 6718-6720 (dentaries), 6700 (left premaxilla), 6701 (left maxilla) and 6721 (first ceratobranchial)
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| Aves | |
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Berruornis orbisantiqui17 Mourer-Chauviré 1994 owl L 3096 (cast of a complete right tarsometatarsus)
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"Gastornis edwardsii n. sp." = Gastornis parisiensis15
"Gastornis edwardsii n. sp." = Gastornis parisiensis15 Hébert 1855 bird MNHN CRL 2457 - lectotype (partial tibiotarsi); additional MHNH CRL referred material
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Remiornis heberti n. gen. n. sp.15
Remiornis heberti n. gen. n. sp.15 Lemoine 1881 bird MNHN CRL 2481 - lectotype (partial tarsometatarsus); additional MHNH CRL referred material
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Eupterornis remensis n. gen. n. sp.17
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| Reptilia | |
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Camptognathosaurus parisiensis6 Folie et al. 2013 worm lizard About fifteen dentaries and maxillae including MNHN CR 17420, a well-preserved right dentary, MNHN CR 17421, posterior part of a right dentary and MNHN CR 17425, anterior part of a left dentary
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Cernaycerta duchaussoisi n. gen. n. sp.1 Augé 2005 squamates MNHN CR 17403 (holotype), incomplete left dentary; MNHN CR 17402, partial posterior left dentary with a portion of the splenial
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| Mammalia | |
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Phakodon levei12 Russell et al. 1967 eutherian LM1, MNHN.F.CR-285; RM2, MNHN.F.CR-4420; RM3, MNHN.F.CR-292; LM1, MNHN.F.CR-4416; RM1, MNHN. F.CR-4414; two LM2s, MNHN.F.CR-1309, CR-4413; LM1, MNHN.F.CR-63-Pn; RM1, MNHN.F.CR-72-L; RM2, MNHN.F.CR-4483; LM3, NMB.CY-370; RM3, MNHN.F.CR- 15926
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"Paschatherium petri n. gen. n. sp." = Dipavali petri13, Walbeckodon girardi n. sp.13, Louisina mirabilis13, Louisina marci n. sp.13, Berrulestes pellouini13, Berrulestes phelizoni13, Berrulestes poirieri13, Thryptodon brailloni n. gen. n. sp.13, Gigarton sigogneauae13, Gigarton meyeri13, Gigarton louisi n. sp.13
"Paschatherium petri n. gen. n. sp." = Dipavali petri13 Russell 1964 elephant shrew MNHN.F.CR-142 - holotype (Left dentary fragment with M2 and alveoli for M1, M3); referred material: LP3, MNHN.F.CR-4390; three RP3s, MNHN.F.CR-329, CR-1137, CR-115-Bn; eight LP4s, MNHN.F.CR-633, CR-15938(Collier 837), CR-65-Ph, CR-885-Ph, CR-366-Pn, CR-67-Bn, CR-282-L, CR-55- MD; 15 RP4s, MNHN.F.CR-279, CR-1107, CR- 1176, CR-1207, CR-124-Pn, CR-359-Pn, CR-1728-Pn, CR-337-Ph, CR-338-Ph, CR-218-Bn, CR-283-L, CR-285-L, CR-286-L, CR-82-MD, CR-86-MD; five LM1s, MNHN.F.CR-64-Pn, CR-283-Bn, CR-696-Ph, CR-295-L, SLP.CR-404; nine RM1s, MNHN.F.CR-628, CR-4519, CR-15935(Collier 834), CR-86-Pn, CR-280- Bn, CR-292-L, CR-293-L, CR-294-L, CR-17481-L; five LM2s, MNHN.F.CR-217, CR-11897, CR-692-Ph, CR-297-L, CR-1418-Pn; 13 RM2s, MNHN.F.CR-218, CR-11895, CR-13775, CR-89-Ph, CR-107-Ph, CR-296-L, CR-298-L, CR-299-L, CR-300-L, CR-301-L, CR-1143-Pn, SLP.CR-319, CR-323; RM3, MNHN.F.CR- 941; two LDP3, MNHN.F.CR-535, CR-599; RDP3, MNHN.F.CR-224; RDP4, MNHN.F.CR-1438-Pn; RP2, MNHN.F.CR-38-Pn; five LP3s, MNHN.F.CR-595, CR-4372, CR-101-Ph, CR-144-Ph, CR-325-Ph; eight RP3s, MNHN.F.CR-283, CR-303, CR-508, CR-1184, CR-13759, CR-15954(Collier 553), CR-49-Pn, CR-327- Ph; two L dentaries, P4-M1, MNHN.F.CR-62-De, CR-73-MD; six LP4s, MNHN.F.CR-299, CR-519, CR-72-Pn, CR-689-Ph, SLP.CR-8, CR-398; six RP4s, MNHN.F. CR-1304, CR-4523, CR-17-Pn, CR-1462-Pn, CR-398-Ph, SLP.CR-345; ten M1s, MNHN.F.CR-53-Pn, CR-315-Pn, CR-317-Pn, CR-1134-Pn, CR-1711-Pn, CR-302-L, CR-303-L, CR-304-L, CR-305-L, C-418-W; eight RM1s, MNHN.F.CR-4514, CR-25-Pn, CR-46-Pn, CR-368-Pn, CR-1393-Pn, CR-1716-Pn, SLP.CR-316, CR-401; eight LM2s, MNHN.F.CR-342, CR-56-Pn, CR-1713-Pn, CR-15-Bn, CR-308-L, CR-309-L, SLP.CR- 318, CR-348; eight RM2s, MNHN.F.CR-86-Ph, CR-375- Ph, CR-356-Pn, CR-362-Pn, CR-1139-Pn, CR-306-L, CR-310-L, SLP.CR-322; seven LM3s, MNHN.F.CR-239, CR-11972, CR-324-Pn, CR-1153-Pn, CR-1732-Pn, CR-1761-Pn, CR-325-L; four RM3s, MNHN.F.CR-87-Pn, CR-1735-Pn, CR-326-L, CR-336-L; L dentary, DP3–4, MNHN.F.CR-14167; LDP3, MNHN.F.CR-590- Ph
Walbeckodon girardi n. sp.13 Hooker and Russell 2012 elephant shrew MNHN.F.CR-330 - type (left dentary fragment with M1–3); paratypes: RP3, MNHN.F.CR-134-Ph; RP4, MNHN. F.CR-4412; three LM1s, MNHN.F.CR-81-Pn, CR-7-Bn, CR-4442; 2 RM1s, MNHN.F.CR-1229, CR-329-Bn; RM2, MNHN.F.CR-116-Ph; RM3, MNHN.F.CR-334-L; L dentary with P4–M2, MNHN.F.CRL-825; R dentary with M1–2, MNHN.F.CR-331; LP4, MNHN.F.CR-1225; two RP4s, MNHN.F.CR-11906, CR-168-Bn; three LM1s, MNHN.F.CR-13778, CR-12-Pn, UCMP. 62075(CR-329); two RM1s, MNHN.F.CR-1753-Pn, UCMP.62003(CR-4409); three LM2s, MNHN.F.CR- 11992, CR-58-Pn, CR-73-Pn; four RM2s, MNHN.F.CR- 189, CR-11889, CR-109-Bn, NMB.CY-953; two RM3s, MNHN.F.CR-230-Bn, CR-951-Pn; referred material: LM2, MNHN.F.CR-661; RM2, MNHN.F.CR-188; L dentary with P4-M1, MNHN. F.CR-1753-Pn; 3 LM2s, MNHN.F.CR-474, CR-11910, CR-11992; RM2, MNHN.F.CR-4411
Louisina mirabilis13 Russell 1964 elephant shrew LP4, MNHN.F.CR-248-Bn; RM1, UCMP.62011(CR-239); LM3, MNHN.F.CR-4329; RM3, MNHN.F.CR-1285; LM3, MNHN.F.CR-1442-Pn; RM3, MNHN.F.CR-75-MD
Louisina marci n. sp.13 Hooker and Russell 2012 elephant shrew MNHN.F.CR-235 - type (left M1); paratypes: RP4, MNHN.F.CR-77-MD; four LM1s, MNHN.F.CR-4325, CR-11907, CR-571-Ph, UCMP. 62021(CR-4321); three RM1s, MNHN.F.CR-240, CR-54-Bn, CR-1140-Pn; five LM2s, MNHN.F.CR-247, CR-32-MD, UCMP.62015(CR-248), 62010(CR-1242), SLP.CR-315; 5 RM2s, MNHN.F.CR-216, CR-625, CR-951, CR-15929, UCMP.62013(CR-244); 3 LM3s, MNHN.F.CR-4336, CR-323-L, CR-324-L; seven RM3s, MNHN.F.CR-1143, CR-4323, CR-11904, CR-187-Ph, CR-572-Ph, CR-68-Bn, CR-322-L; LDP4, MNHN.F.CR- 4397; two RP3s, MNHN.F.CR-312-L, UCMP.62018(CR- 308); two LP4s, MNHN.F.CR-76-MD, SLP.CR-1-De; two RP4, MNHN.F.CR-233, CR-1117; LM1, FSL.1. 882; three RM1s, MNHN.F.CR-1-F, CR-25-MD, UCMP.62014(CR-949); four RM2s, MNHN.F.CRL-568, CRL-569, CR-17-MD, UCMP.62008(CR-4330); LM3, MNHN.F.CR-1704-Pn; two RM3s, MNHN.F.CR-773- Ph, CR-327-L; referred material: Two LM3, MNHN.F.CR-1272, CR-337-L; LM1, MNHN.F.CR-15936(Collier835); LM2, UCMP.62020(CR-233)
Berrulestes pellouini13 Hooker and Russell 2012 elephant shrew Paratypes: RM1, MNHN.F.CR-111-Ph; two LM3, MNHN.F.CR-377, CR-548; two RP4s, MNHN.F.CR- 1135-Pn, CR-39-MD; LM1, MNHN.F.CR-143-Pn; RM1, MNHN.F.CR-1154-Pn; LM2, MNHN.F.CR-1719-Pn; RM2, MNHN.F.CR-1425-Pn; LM3, MNHN.F.CR- 15928; referred material: RM1, MNHN.F.CR-149-Pn; LM2, MNHN.F.CR-4428; RM2, MNHN.F.CR-1126
Berrulestes phelizoni13 Hooker and Russell 2012 elephant shrew Paratypes: RP4, MNHN.F.CR-1693-Pn; four LM1s, MNHN.F.CR-213, CR-1230, CR-59-V, CR-20-MD; six RM1s, MNHN.F.CR-210, CR-937, CR-5487, CR-1390-Pn, CR-1699-Pn, UCMP.62006(CR-4419); RM2, MNHN.F.CR-3-Bn; LP4, MNHN.F.CR-34-MD; RP4, MNHN.F.CR-1448-Pn; three RM1s, MNHN.F. CR-42-Pn, CR-1415-Pn, CR-1707-Pn; three LM2s, MNHN.F.CR-281-Bn, CR-333-Pn, CR-17477; three RM2s, MNHN.F.CR-11901, CR-307-L, CR-94-Pn; five RM3s, MNHN.F.CR-341-Pn, CR-1730-Pn, CR-329-L, CR-330-L, CR-56-MD; RDP4, FSL.1.881; referred material: LP4, MNHN.F.CR-17476; RP4, MNHN.F.CR-237-Bn; 2 LM1s, MNHN.F.CR-384, CR-282-Bn; LM3, MNHN.F.CR-1424-Pn; doubtfully referred specimen: LDP4, MNHN.F.CR- 374
Berrulestes poirieri13 Hooker and Russell 2012 elephant shrew Paratypes: LP3, MNHN.F.CR-465-Ph; RM2, MNHN. F.CR-316-L; two LM2s, MNHN.F.CR-86-Bn, CR-371- Pn; five RM2s, MNHN.F.CR-1119, CR-99-Bn, CR-286- Bn, CR-1692-Pn, IRSNB.CR-101A-Wouters; RM3, MNHN.F.CR-208; referred material: Two LM2s, MNHN.F.CR-318-L, CR-320-L; two RM2s, MNHN.F.CR-319-L, CR-312-Bn
Thryptodon brailloni n. gen. n. sp.13 Hooker and Russell 2012 elephant shrew MNHN.F.CR-1268 - holotype (LM2); paratypes: RM2, MNHN.F.CR-311-L; RM1, MNHN. F.CR-55-Bn; LM2, MNHN.F.CR-317-L; two RM2s, MNHN.F.CR-90-Bn, CR-63-V; tentatively referred specimen: LP3, MNHN.F.CR-170- Ph
Gigarton sigogneauae13 Hooker and Russell 2012 elephant shrew Paratypes: LM1, MNHN.F.CR-78-MD; right M1, MNHN.F.CR-284-Bn; two RM3s, MNHN.F.CR-88-Ph, CR-335-L; LDP4, MNHN.F.CR-156-Ph; RDP4, MNHN.F.R1218; LP4, MNHN.F.CR-15937(Collier 836); LM1, CR-288-L; LM3, CR-321-L; referred material: LDP4, MNHN.F.CR-1206
Gigarton meyeri13 Hooker and Russell 2012 elephant shrew Paratypes: Two LP4s, MNHN.F.CR-11909, CR-6-Bn; LM1, MNHN.F.CR-318; RM1, CR-971-Pn; RDP4, MNHN.F.CR-340-Ph; LP3, MNHN.F.CR-1211; LP4, MNHN.F.CR-287-L; three RP4s, MNHN.F.CR-946, CR-1159, CR-1696-Pn; LM1, MNHN.F.CR-4331; RM1, MNHN.F.CR-13773; Doubtfully referred specimens: LP3?, MNHN.F.CR- 284-L; RP3?, CR-80-MD
Gigarton louisi n. sp.13 Hooker and Russell 2012 elephant shrew MNHN.F.CR-93-L - type (LM2); paratypes: LM1, MNHN.F.CR-291-L; four RM1s, MNHN.F.CR-435, CR-289-L, CR-8-MD, CR-1447-Pn; RM2, MNHN.F.CR-45-MD; LM3, MNHN.F.CR-129-Pn; LDP4, MNHN.F.CR-290-L
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Dissacus europaeus n. sp.22
Dissacus europaeus n. sp.22 Lemoine 1891 condylarth MNHN.F.CRL982 - type; referred material: MNHN.F.CRL1129, 1937-611, 14037, 17232
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Paschatherium dolloi13 Teilhard de Chardin 1927 odd-toed ungulate Left dentary fragment with P4–M3, MNHN.F.CRL-827; RM1, MNHN.F.CR-5-Bn
Teilhardimys brisswalteri n. sp.13 Hooker and Russell 2012 odd-toed ungulate MNHN.F.CR-4400 - type (LP4); paratypes: Two LP4s, MNHN.F.CR-11888, CR-38- Meniel; RP4, MNHN.F.CR-4521; L maxilla, M1–3, MNHN.F.CR-265-Bn; three RM1s, MNHN.F.CR-1281, CR-118-Pn, CR-33-MD; two LM2s, MNHN.F.CR- 568-Ph, CR-27-MD; LM3, MNHN.F.CR-538; RDP4, MNHN.F.CR-343-Ph; two RP3s, MNHN.F.CR-285-Bn, SLP.CR-352; three LP4s, MNHN.F.CR-1283, CR-4339, CR-546-Ph; two RP4s, MNHN.F.CR-238, CR-1264; five LM1s, MNHN.F.CR-234, CR-4324, CR-11891, CR-192- Ph, CR-690-Ph; five RM1s, MNHN.F.CR-358, CR- 191-Ph, CR-102-Pn, CR-358-Pn, SLP.CR-317; L dentary, M2–3, MNHN.F.CR-3-Bw; four LM2s, MNHN.F. CR-4332, CR-6-F, UCMP.62009(CR-4327), 62016(CR-1157); seven RM2s, MNHN.F.CR-316, CR-4328, CR-1180, CR-367-Pn; CR-372-Pn, CR-1703- Pn, SLP.CR-320; six LM3s, MNHN.F.CR-11890, CR-15930(Collier 699), CR-1724-Pn, CR-273-Bn, CR-84-MD, UCMP.62023(CR-1104); 13 RM3s, MNHN. F.CR-236, CR-237, CR-1146, CR-1251, CR-1722, CR-1396-Pn, CR-1705-Pn, CR-1718-Pn, CR-203-Ph, CR-328-L, CR-309-Bn, CR-83-MD, CR-85-MD; referred material: Two LP4s, MNHN.F.CR-207, CR-152-Bn; LM2, MNHN.F.CR-17479; two RM2s, MNHN.F.CR-294, CR-4322; two LM1s, MNHN.F.CR- 545-Ph, UCMP.62019(CR-4326); RM2, MNHN.F.CR- 619-Ph; RM3, MNHN.F.CR-11977
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Landenodon lavocati n. sp.21
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Remiculus deutschi n. gen. n. sp.23
Remiculus deutschi n. gen. n. sp.23 Russell 1964 placental from "Lemoine Quarry at Cernay-les-Reims" according to Szalay
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"Berruvius gingerichi n. sp." = Sarnacius gingerichi
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Adapisoriculidae indet.7, "Afrodon germanicus" = Bustylus germanicus8, Bustylus cernaysi n. gen. n. sp.9, Afrodon sp.8
Adapisoriculidae indet.7 Van Valen 1967 eutherian MNHM CR 1194 - originally considered a marsupial (Russell 1964) and later considered a probable eutherian (Clemens, 1986; Clemens and Marshall, 1976; Savage and Russell, 1983)
"Afrodon germanicus" = Bustylus germanicus8 Russell 1964 eutherian MNHM specimens; upper dentition: CR-18-Bn (right M2/ broken lingually), CR-19-Bn (right M2/), CR 181 Ph (left M2/); lower dentition: CR-1-Bn (right M/2?), CR-32-PN (left M/3), CR 399 (right M/l?), CR-354-Ph (right M/3), CY 590 (left M/2?), CR 685 (right M/l?), MNHM 16120 (right M/3) and MNHM 16121 (left M/3)
Bustylus cernaysi n. gen. n. sp.9 Gheerbrant and Russell 1991 eutherian MNHM CR 15950 - holotype (left m2); referred material too
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1Augé 2005; 2Broin 1977; 3Čerňanský et al. 2020; 4Codrea et al. 2014; 5Estes et al. 1967; 6Folie et al. 2013; 7Gheerbrant 1991; 8Gheerbrant and Russell 1989; 9Gheerbrant and Russell 1991; 10Gingerich 1973; 11Gingerich 1976; 12Hooker 2013; 13Hooker and Russell 2012; 14Hooker et al. 1999; 15Martin 1992; 16Martin et al. 2014; 17Mourer-Chauviré 1994; 18Pérez-García 2012; 19Pérez-García 2020; 20Rage 2003; 21Russell 1980; 22Solé et al. 2018; 23Szalay 1969; 24Vianey-Liaud 1979