Sansan (Miocene to of France)
Where: Midi-Pyrenees, France (43.5° N, 0.6° E: paleocoordinates 43.7° N, 0.6° W)
• coordinate based on nearby landmark
• outcrop-level geographic resolution
When: Langhian to Langhian (16.0 - 11.6 Ma)
• member-level stratigraphic resolution
Environment/lithology: lacustrine - small; unlithified, gray, blue claystone and marl
• The fossils come from different, superposed lithologies. Those include: purple clays, gray clays, marls, but also limestones and molasse. No distinction is made about the exact provenance of the fossils from these various levels.
Size classes: macrofossils, mesofossils
Primary reference: E. Lartet. 1836. [Lettre de E. Lartet a M. Michelin au sujet de la localité de Sansan (Gers), en lecture a la seance du 16 Mai 1836]. Bulletin de la Societe geologique de France 1(7):217-220 [E. Fara/E. Fara/E. Fara]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 32191: authorized by Emmanuel Fara, entered by Emmanuel Fara on 09.06.2003, edited by Susanna Kümmell, Chris Bell, Matthew Carrano, Patricia Holroyd, Evangelos Vlachos, Mark Uhen, Bethany Allen, Terri Cleary and Torsten Liebrecht
Creative Commons license: CC BY (attribution)
Taxonomic list
• The ref. Ginsburg (1964)is an update of previous works (Lartet 1836-1858) and Filhol (1891), but it was published as new excavations were taking place.
Show authors, comments, and common names
| unclassified | |
| Gastropoda | |
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Lymnaea aff. stagnalis snail | |
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Planorbis sp. Muller 1774 snail | |
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Ancilla sp. Lamarck 1799 olive snail | |
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Bulimus lubricus Müller 1774 snail | |
| Mammalia | |
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"Deinotherium levius" = Deinotherium giganteum7
"Deinotherium levius" = Deinotherium giganteum7 Kaup 1829 deinothere entered as "Dinotherium levius"
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"Trilophodon angustidens" = Tetrabelodon angustidens7
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"Chiroptera" "I nov. gen. nov. sp. (Ginsburg 1964)"7, "Chiroptera" "II ("Vespertilio" noctuloides Lartet)(Ginsburg 1964)(Ginsburg 1964)"7, "Chiroptera" "III ("Vespertilio" murinoides Lartet) (Ginsburg 1964)"7
"Chiroptera" "II ("Vespertilio" noctuloides Lartet)(Ginsburg 1964)(Ginsburg 1964)"7 Blumenbach 1779 bat | |
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Dicerorhinus sansaniensis7, Brachypotherium brachypus7, "Aceratherium tetradactylum" = Hoploaceratherium tetradactylum7
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Taxodon sansaniensis7, Mustelinae indet.7, "Lutra dubia" = Lartetictis dubia9, Melidellavus leptorhynchus7, Martes muncki7, Martes sansaniensis7, Ischyrictis zibethoides7, Mionictis dubia7
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Eumyarion medius8, Megacricetodon minor5, Cricetodon helveticus7, Cricetodon gaillardi7, Cricetodon cf. affine7, Cricetodon sansaniensis7, "Cricetodon minus" = Democricetodon minor7, "Cricetodon cf. breve" = Democricetodon brevis7
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| Reptilia | |
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? Scincidae sp.1 Gray 1825 skink 18 dentaries entered as ? Scincidae sp. 1; one dentary entered as ? Scincidae sp. 2
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Natrix sansaniensis1, Natrix longivertebrata1, Natrix (aff. Natrix) longivertebrata1, Paleoheterodon arcuatus1, Colubrinae indet.1, Texasophis meini1, Elaphe sp.1, Coluber pouchetii1, Coluber (aff. Coluber) pouchetii1, Natricinae indet.1, Neonatrix natricoides n. sp.1, Neonatrix europaea1, Palaeonatrix lehmani1, Palaeonatrix (aff. Palaeonatrix) lehmani1
Colubrinae indet.1 Oppel 1811 colubrid snake includes "Colubrinae incertae sedis nomen dubium" applied to Scaptophis Rochebrune, 1880; also includes 103 indeterminate vertebrae
Neonatrix natricoides n. sp.1 colubrid snake Holotype MNHN Sa 23683, one mid-trunk vertebra; 79 referred vertebrae
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Vipera aegertica n. sp.1 pit viper Holotype (Sa 23714) one trunk vertebra; also 20 referred vertebrae
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Lacerta sp.1 Linnaeus 1758 squamates 265 dentaries entered as Lacerta sp. 1; 54 dentaries entered as Lacerta sp. 2
Edlartetia sansaniensis n. gen.1 Augé and Rage 2000 squamates neotype designated as MNHN Sa 23595, an incomplete right dentary
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Gekkonidae indet.1 Gray 1825 gecko 17 maxilla or dentary fragments; 34 trunk vertebrae; 4 caudal vertebrae
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Anguimorpha indet.1 squamates ; published as Anguimorpha incertae sedis; 1 anterior left dentary fragment
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Ophisaurus sp.1 Daudin 1803 glass lizard 24 jaw elements (maxillae or dentary fragments) entered as Ophisaurus sp. 1; 5 jas elements (maxillae or dentary fragments) entered as Ophisaurus sp. 2
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"Trionyx sansaniensis n. sp." = Chelydropsis murchisoni2, Broilia robusta n. gen. n. sp.3, Leptochelys braneti n. gen. n. sp.3
Broilia robusta n. gen. n. sp.3 Bergounioux and Crouzel 1965 turtle MNHN SA uncat. (lectotype), a string of “caudal” vertebrae
Leptochelys braneti n. gen. n. sp.3 Bergounioux and Crouzel 1965 turtle MNHN SA2023 (holotype), a partial hyo/hypoplastron
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"Ptychogaster sansaniensis" = Ocadia sansaniensis6, "Ocadia sansaniensis n. sp." = Ocadia sansaniensis10, Clemmydopsis turnauensis6, Ptychogaster cf. batalleri6
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| Aves | |
"Anas sansaniensis" = Chenoanas sansaniensis16
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| Amphibia | |
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Albanerpeton inexpectatum14 Estes and Hoffstetter 1976 tetrapod Sa 23401 - Sa 23403, Sa 23404a, Sa 23405, Sa 23407 - Sa 23409
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| Angiospermae | |
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| Cyclostoma elegans | |
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Cyclostoma elegans Muller 1774 | |
1Augé and Rage 2000; 2Bergounioux 1935; 3Bergounioux and Crouzel 1965; 4Böhme 1999; 5Bollinger 1994; 6Broin 1977; 7Ginsburg 1964; 8Heißig 1989; 9Heizmann and Morlo 1998; 10Lartet 1851; 11Lydekker 1889; 12Mlíkovsky 1997; 13Rage and Bailon 2005; 14Rage and Hossini 2000; 15Tassy 1977; 16Zelenkov et al. 2018