Kotlovina, middle layer (Pliocene of Ukraine)

Also known as Kotlovyna

Where: Odessa, Ukraine (45.5° N, 28.6° E: paleocoordinates 45.6° N, 28.3° E)

• coordinate based on nearby landmark

When: MN 16 (3.2 - 2.6 Ma)

• "Upper Pliocene (Kuyalnick, MN 16)"

•According to the stratigraphic chart in the primary reference, the site is located in the Khaprovski faunistic complex.

Environment/lithology: terrestrial; unlithified, argillaceous sandstone

• "clayish sands with numerous bone remains"

Size classes: macrofossils, mesofossils

Primary reference: G. A. Zerova. 1993. Late Cainozoic localities of snakes and lizards of Ukraine. Revue de Paléobiologie, Volume spéciale 7:273-280 [J. Mueller/T. Liebrecht/T. Liebrecht]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 168020: authorized by Johannes Mueller, entered by Torsten Liebrecht on 12.04.2015

Creative Commons license: CC BY (attribution)

Taxonomic list

Reptilia
 Squamata - Boidae
Erycinae indet. Bonaparte 1825 sand boa
"trunk vertebrae of small erycines"
 Squamata - Crotalidae
Vipera (Pelias) sp. Merrem 1820 pit viper
trunk vertebrae, "compatible in size and morphology with that of V. (P.) berus"
 Squamata - Colubridae
Colubridae indet. Oppel 1811 colubrid snake
trunk and caudal vertebrae
Natricinae indet. Bonaparte 1838 colubrid snake
trunk and caudal vertebrae
 Squamata - Lacertidae
Lacertidae indet. Bonaparte 1831 squamates
"fragmentary parietal bone, dentaries and vertebrae"
 Squamata - Varanidae
Varanus sp. Merrem 1820 monitor lizard
caudal vertebrae
 Squamata - Anguidae
Pseudopus sp. Merrem 1820 legless lizard
osteoscutes