Where: Leiria, Portugal (39.7° N, 8.8° W: paleocoordinates 33.7° N, 1.9° E)
• coordinate estimated from map
• local area-level geographic resolution
When: Alcobaça Formation, Kimmeridgian (157.3 - 152.1 Ma)
• two lignite coal layers wth vertebrate fossils. Dated as Oxfordian based on microflora, Kimmeridgian based on ostracods. Recent work (Schudack 1993) favors Kimmeridgian.
• group of beds-level stratigraphic resolution
Environment/lithology: lagoonal; coal
Size classes: macrofossils, mesofossils, microfossils
Preservation: mold/impression
Collection methods: surface (in situ), mechanical,
Primary reference: C. Brauckmann. 1978. Beitrag zur Flora der Grube Guimarota (Ober-Jura; Mittel-Portugal). Geologica et Palaeontologica 12:213-222 [A. Rees/A. Rees/A. Rees]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 24563: authorized by Allister Rees, entered by Allister Rees on 09.08.2002, edited by Meg Hall, Matthew Carrano, Kaitlin Maguire, Roger Benson, Philip Mannion, Oliver Rauhut and Terri Cleary
Creative Commons license: CC BY (attribution)
Taxonomic list
unclassified | |
Foraminifera | |
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Demospongiae | |
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Chondrichthyes | |
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Amphibia | |
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Mammalia | |
Paulchoffatiidae "gen. A"10, Paulchoffatiidae "gen. B"10, Paulchoffatiidae "gen. C"10, Paulchoffatiidae "gen. D sp. 1"10, Paulchoffatiidae "gen. D sp. 2"10, Paulchoffatiidae "gen. D sp. 3"10, Paulchoffatiidae "gen. D sp. 4"10, Meketichoffatia sp.10, Meketichoffatia krausei n. gen. n. sp.6, Meketichoffatia ? krausei10, Paulchoffatia delgadoi n. gen. n. sp.6, Henkelodon sp.10, Henkelodon naias n. gen. n. sp.5, Kuehneodon sp. n. sp.5, Kuehneodon simpsoni6, Kuehneodon uniradiculatus n. sp.6, Kuehneodon dietrichi n. gen. n. sp.6, Kuehneodon guimarotensis n. sp.6, Meketibolodon robustus9, Guimarotodon leiriensis n. gen. n. sp.6, Pseudobolodon krebsi n. sp.8, "Kuehneodon dryas n. sp." = Pseudobolodon dryas5, Pseudobolodon oreas n. gen. n. sp.5, Pseudobolodon oreas7, Renatodon amalthea n. gen. n. sp.7, Bathmochoffatia hapax n. gen. n. sp.10, Kielanodon hopsoni n. gen. n. sp.6, Xenachoffatia oinopion n. gen. n. sp.10, "Parachoffatia thoas n. gen. n. sp." = Plesiochoffatia thoas10, "Parachoffatia peperethos n. sp." = Plesiochoffatia10, Parachoffatia staphylos n. sp.10
Paulchoffatiidae "gen. A"10 Hahn 1969 multituberculate V.J. 385-155 and IPFUB A(m2)-1-5 (five m2 teeth)
Paulchoffatiidae "gen. D sp. 1"10 Hahn 1969 multituberculate V.J. 163-155 (a left m2), IPFUB D1(m2)-1-5 (five m2 teeth)
Paulchoffatiidae "gen. D sp. 3"10 Hahn 1969 multituberculate V.J. 381-155 (left m2 tooth), 3880155 (right m2 tooth), 389-155 (right m2 tooth); IPFUB D3(m2)-1-12 (twelve m2 teeth)
Meketichoffatia krausei n. gen. n. sp.6 Hahn 1993 multituberculate V.J. 110-155 (holotype skull), 443-155, 444-155, 445-155, 446-155 (skulls); plus an m1 referred by Hahn (1998) (IPFUB M(m1)-1)
Meketichoffatia ? krausei10 Hahn 1993 multituberculate V.J. 390-155-395-135 (6 x m1 referred to Paulchoffatia sp. by Hahn (1969))
Paulchoffatia delgadoi n. gen. n. sp.6 Kuhne 1961 multituberculate V.J. 1-155 (holotype mandible), 2-155, 3-155, 402-155, 403-155 (mandibles); plus specimens referred by Hahn (1998): V.J. 95-155, 96-155, IPFUB de(m2)-1, de(m2)-2 (total of four m2 teeth)
Kuehneodon uniradiculatus n. sp.6 Hahn 1978 multituberculate V.J. 427-155 (holotype mandible), 428-155, 429-155, 431-155, 442-155 (mandibles)
Kuehneodon dietrichi n. gen. n. sp.6 Hahn 1969 multituberculate V.J. 4-155 (holotype mandible), 410-155 to 425-155, 429-155, 439-155, 441-155 (mandibles)
Kuehneodon guimarotensis n. sp.6 Hahn 1969 multituberculate V.J. 5-155 (holotype mandible), 6-155, 11-155(questionably), 440-155 (mandibles)
Meketibolodon robustus9 Hahn 1978 multituberculate V.J. 408-155 (holotype mandible), 407-155, 409-155, 430-155 (mandibles)
Guimarotodon leiriensis n. gen. n. sp.6 Hahn 1969 multituberculate V.J. 396-155 (holotype mandible), 461-155, 462-155 (mandibles)
Pseudobolodon krebsi n. sp.8 Hahn and Hahn 1994 multituberculate V.J. 447-155 (holotype skull fragments), plus 451-155, 357-155
Renatodon amalthea n. gen. n. sp.7 Hahn 2001 multituberculate V.J. 457-155 (holotype right maxilla), 452-155 (paratype)
Kielanodon hopsoni n. gen. n. sp.6 Hahn 1987 multituberculate V.J. 463-155 (holotype right maxilla), 464-155 (right maxilla); plus a right p3 reported by Hahn & Hahn (1998) V.J. 382-155
Xenachoffatia oinopion n. gen. n. sp.10 Hahn and Hahn 1998 multituberculate IPFUB oi(m2)-1 (holotype left m2), -2-3 (paratype right m2s)
"Parachoffatia thoas n. gen. n. sp." = Plesiochoffatia thoas10 Hahn and Hahn 1998 multituberculate IPFUB th(m2)-1 (holotype left m2), -2-3 (paratype right m2s)
"Parachoffatia peperethos n. sp." = Plesiochoffatia10 multituberculate IPFUB pe(m2)-1 (holotype right m2)
Parachoffatia staphylos n. sp.10 Hahn and Hahn 1998 multituberculate IPFUB st(m2)-1 (holotype left m2)
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Proalbionbaatar plagiocyrtus n. gen. n. sp.11
Proalbionbaatar plagiocyrtus n. gen. n. sp.11 Hahn and Hahn 1998 multituberculate IPFUB pl(m1)-1-2 (holotype right m1 and paratype left m1)
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Nanolestes drescherae n. gen. n. sp.21
Nanolestes drescherae n. gen. n. sp.21 Martin 2002 mammal The holotype (Gui Mamm 1002) and the following specimens: lower teeth (molars if not otherwise indicated): Gui Mam 1000 (left); 1001 (left); 1003 (left); 1004 (left); 1005 (dp?2-3, left); 1006 (right); 1007 (sectioned for enamel analysis); 1008 (left); 1009 (left); 1010 (left); 1011 (left); 1012 (right); 1013 (dp?3- 5, right); 1014 (left); 1015/1 (right) and 1015/2 (right), most probably from the same individual; 1016 (right); 1017 (right); 1018 (right); 1019 (dp?3-5, right); 1020 (right); 1021 (left); 1022 (left); 1040 (left); anterior dentary fragment with p3-5, broken roots of p2 and alveoli for il-4, c, and pl (Gui Mam 66/79); right dentary with p2 and alveoli for il-4, p1-5, and ml-5 (Guimarota 19; "cf. Peramus" of Ktihne, 1968a); upper teeth (molars if not otherwise indicated): Gui Mam 1023/1 (P4 or 5, left), 1023/2 (left), 1023/3 (left), most probably from the same individual; 1024 (left); 1025 (left); 1026 (left); 1027 (DP? 3-5, left); 1029 (left); 1030 (left); 1031 (right); 1032 (right); 1033 (right); 1034 (DP? 3-5, right); 1035 (right); 1036 (left); 1037 (right); 1038 (right); 1039 (DP? 3-5, right, double-rooted, roots broken, tooth was not shed); 1041 (left); left maxilla fragment consisting of two maxillary fragments and five isolated teeth (C, P1-P4) and a possible molar fragment (Gui Mam 176/ 75).
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Dryolestidae indet.15, Henkelotherium guimarotae27, Dryolestes leirensis n. sp.20, Dryolestes cf. leirensis20, Guimarotodus inflatus n. gen. n. sp.20, Krebsotherium lusitanicum n. gen. n. sp.20
Dryolestes leirensis n. sp.20 Martin 1999 mammal Gui Mam 130/74 (holotype left mandible) plus 81 other specimens. Includes Butterigale sp. of Kuhne 1968
Guimarotodus inflatus n. gen. n. sp.20 Martin 1999 mammal IPFUB Gui Mam 121/75 (holotype right mandible) plus ten other specimens
Krebsotherium lusitanicum n. gen. n. sp.20 Martin 1999 mammal IPFUB Gui Mam 9/75 (holotype left mandible) plus 78 other specimens
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Drescheratherium acutum n. gen. n. sp.16 Krebs 1998 mammal IPFUB Gui Mam 4/73 (holotype maxilla), 34/A6, 70/74, 1/76
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Osteichthyes | |
Haldanodon exspectatus n. sp.18
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Reptilia | |
"Chelonia indet." = Testudines1
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"Peltochelys duchastelli" = Peltochelys duchastelii13
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Lisboasaurus estesi n. gen. n. sp.29 Seiffert 1975 crocodilian FUB Giu 37, near-complete right maxilla with one tooth; plus FUB field batch number Jun 18:13:1
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Goniopholis baryglyphaeus n. sp.26 Schwarz 2002 crocodilian IPFUB Gui Croc 1, holotype: an articulated partial skeleton, including a nearly complete dorso-ventrally ¯attened skull with major portions of the lower jaw; four cervical vertebrae, six thoracic vertebrae, one sacral vertebra and three caudal vertebrae; the right and left coracoids, the left ischium and portions of the right femur, tibia and ®bula, a metacarpal, and several dorsal and ventral osteoderms
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"Theriosuchus guimarotae n. sp." = Knoetschkesuchus guimarotae28
"Theriosuchus guimarotae n. sp." = Knoetschkesuchus guimarotae28 Schwarz and Salisbury 2005 crocodilian IPFUB Gui Croc 7308
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Machimosaurus hugii14 Meyer 1837 marine crocodile Fragments of the snout, part of the frontal, the left postorbital, the pterygoids left and the occiput, also two dorsal vertebrae, two sacral vertebrae, a tail vertebrae, ribs, left fibula, and some armor plates; found in 1961
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Titanosauriformes indet.19, cf. Ceratosaurus sp.22, Tetanurae indet.22, Aviatyrannis jurassica n. gen. n. sp.24, Aviatyrannis jurassica24
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Aves | |
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Reptilia | |
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Portugalophis lignites n. gen. n. sp.2
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"Introrsisaurus pollicidens n. gen. n. sp." = Dorsetisaurus purbeckensis29
"Introrsisaurus pollicidens n. gen. n. sp." = Dorsetisaurus purbeckensis29 Hoffstetter 1967 squamates | |
Saurillus henkeli n. sp.29, Saurillus cf. obtusus29, "Saurillus proraformis n. sp." = Saurillodon proraformis29, "Becklesisaurus hoffstetteri n. sp." = Becklesius hoffstetteri29
"Becklesisaurus hoffstetteri n. sp." = Becklesius hoffstetteri29 Seiffert 1975 squamates FUB Gui.A56
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"Cteniogenys reedi n. sp." = Cteniogenys antiquus29
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Actinopteri | |
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Ostracoda | |
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Theriosynoecum hemigymnon30, Theriosynoecum wyomingense30, Bisulcocypris sp.30, Timiriasevia mackerrowi30
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Bivalvia | |
"Lamellibranchia indet." = Autobranchia30
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Gastropoda | |
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Polypodiopsida | |
? Schizoneura sp. Schimper and Mougeot 1844
Equisetum lusitanicum horsetail | |
Ginkgoopsida | |
Ginkgo sp. Linnaeus 1771 ginkgo | |
? Caytonia sp. Thomas 1925 | |
Otozamites sp. Braun 1843 | |
Charophyceae | |
Brachyphyllum sp. Brongniart 1828 |