PBDB Collection

Ikovo (Eocene of Ukraine)

Also known as Novopskov

Where: Luhansk, Ukraine (49.5° N, 39.1° E: paleocoordinates 47.5° N, 33.5° E)

• coordinate stated in text

• small collection-level geographic resolution

When: Lutetian (47.8 - 41.3 Ma)

• early Lutetian

• group of beds-level stratigraphic resolution

Environment/lithology: marine; sandstone

Size class: macrofossils

Collected in 2010–2013

Primary reference: G. Mayr and E. A. Zvonok. 2011. Middle Eocene Pelagornithidae and Gaviiformes (Aves) from the Ukrainian Paratethys. Palaeontology 54(6):1347-1359 [M. Uhen/M. Uhen/O. Alfaro]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 120885: authorized by Mark Uhen, entered by Mark Uhen on 23.11.2011, edited by Evangelos Vlachos and Philip Mannion

Creative Commons license: CC BY (attribution)

Taxonomic list

• Redeposited from past epochs:

•Megasqualus sp.

•Paraorthacodus sp.

•Isurolamna cf. inflata

•Anacoracidae indet.

Show authors, comments, and common names

unclassified
-	"Clerolamna" sp.⁴, "Tobolamna" sp.⁴, Sphyrenodus sp.⁴ "Clerolamna" sp.⁴ "Tobolamna" sp.⁴ «Tobolamna» sp. Sphyrenodus sp.⁴
Osteichthyes
Osteichthyes -	Osteichthyes indet.⁸ Osteichthyes indet.⁸ Huxley 1880 bony fish
Acipenseriformes -	Cylindracanthus sp.⁴ Cylindracanthus sp.⁴ Leidy 1856 ray-finned fish
Actinopteri
Labriformes - Labridae	Labrus sp.⁴ Labrus sp.⁴ Linnaeus 1758
Tetraodontiformes - Triodontidae	Triodon sp.⁴ Triodon sp.⁴ Cuvier 1829
Tetraodontiformes - Eotrigonodontidae	Eotrigonodon sp.⁴ Eotrigonodon sp.⁴ Weiler 1929
Scombriformes - Scombridae	Cybium sp.⁴ Cybium sp.⁴ Cuvier 1829
Scombriformes - Trichiuridae	Eutrichiurides sp.⁴ Eutrichiurides sp.⁴ Casier 1944
Reptilia
Testudines -	Testudinoidea indet.¹ Testudinoidea indet.¹ Gray 1825 turtle
Testudines - Trionychidae	Trionychidae indet.⁵, Trionyx ikoviensis n. sp.¹ Trionychidae indet.⁵ Gray 1825 softshell turtle Trionyx ikoviensis n. sp.¹ Danilov et al. 2011 softshell turtle
Testudines - Cheloniidae	Cheloniidae indet.⁶, Eochelone sp.¹⁰, "Puppigerus nessovi" = Puppigerus camperi¹ Cheloniidae indet.⁶ Bonaparte 1832 sea turtle The material of Cheloni idae gen. indet. (Fig. 2) includes the following ele ments: specimens ZIN PH, nos. 18/145 and 19/145, two postorbitals; specimen ZIN PH, no. 20/145, frag mentary right half of upper jaw; specimen ZIN PH, no. 21/145, costal plate IV; specimen ZIN PH, no.22/145, costal plate VII; specimens ZIN PH, nos. 23–27/145, fragments of five costal plates; speci men ZIN PH, no. 28/145, right peripheral plate II (?); specimen ZIN PH, no. 29/145, left peripheral plate II (?); specimen ZIN PH, no. 30/145, peripheral plate VI or VII; specimen ZIN PH, no. 31/145, posterior part of left epiplastron; specimen ZIN PH, no. 32/145, medial part of left hyoplastron; specimen ZIN PH, no. 33/145, medial part of right hypoplastron; speci men ZIN PH, no. 34/145, proximal part of left humerus; specimens ZIN PH, nos. 35/145 and 36/145, two right tibiae Eochelone sp.¹⁰ Dollo 1903 sea turtle "Puppigerus nessovi" = Puppigerus camperi¹ Gray 1831 sea turtle
Testudines - Pancheloniidae	"cf. Glossochelys sp." = Euclastes¹⁰, Argillochelys sp.⁵ "cf. Glossochelys sp." = Euclastes¹⁰ Cope 1870 turtle Argillochelys sp.⁵ Lydekker 1889 turtle
Crocodylia -	Crocodylia indet.² Crocodylia indet.² Owen 1842 crocodilian ZIN PH 2/145, 59/145, 61/145–74/145
Crocodylia - Gavialidae	Tomistominae indet.⁸ Tomistominae indet.⁸ crocodilian ZIN PH 1/145 (anterior fragment of a right dentary)
Crocodylia -	cf. Diplocynodon sp.² cf. Diplocynodon sp.² Pomel 1847 crocodilian ZIN PH 52/145, 53/145, 55/145, 56/145, 57/145, 58/145, 60/145
Aves
-	Aves indet. Aves indet. Linnaeus 1758 bird
Charadriiformes -	Charadriiformes indet.⁸ Charadriiformes indet.⁸ Huxley 1867 shorebird
Gaviiformes - Gaviidae	Colymbiculus udovichenkoi n. gen. n. sp. Colymbiculus udovichenkoi n. gen. n. sp. Mayr and Zvonok 2011 loon
Procellariiformes -	Kievornis rogovitshi⁷ Kievornis rogovitshi⁷ Averianov et al. 1990 bird
Odontopterygiformes - Pelagornithidae	Lutetodontopteryx tethyensis n. gen. n. sp.³, ? Dasornis sp. Lutetodontopteryx tethyensis n. gen. n. sp.³ Mayr and Zvonok 2012 bird ? Dasornis sp. Owen 1870 bird
Chondrichthyes
Myliobatiformes - Aetobatidae	Aetobatus sp.⁴ Aetobatus sp.⁴ Blainville 1816 stingray
Myliobatiformes - Myliobatidae	Myliobatis sp.⁴, Burnhamia sp.⁴ Myliobatis sp.⁴ Cuvier 1816 eagle ray Myliobatis sp. 1, 2 Burnhamia sp.⁴ Cappetta 1976 eagle ray
Hexanchiformes - Hexanchidae	Notorhynchus sp.⁹ Notorhynchus sp.⁹ Ayres 1855 sevengill shark
Heterodontiformes - Heterodontidae	Heterodontus sp.⁴, "Heterodontus cf. vincenti" = Cestracion vincenti⁴ Heterodontus sp.⁴ de Blainville 1816 bullhead shark "Heterodontus cf. vincenti" = Cestracion vincenti⁴ Leriche 1905 bullhead shark
Squaliformes - Dalatiidae	Isistius trituratus⁴ Isistius trituratus⁴ Winkler 1874 shark
Squatiniformes - Squatinidae	Squatina cf. prima⁴ Squatina cf. prima⁴ Winkler 1874 angel shark
Carcharhiniformes - Scyliorhinidae	Abdounia lapierri⁴, Abdounia cf. minutissima⁴ Abdounia lapierri⁴ cat shark Abdounia cf. minutissima⁴ Winkler 1874 cat shark
Carcharhiniformes - Carcharhinidae	"Rhizoprionodon gantourensis" = Rhizoprionodon ganntourensis⁴, Carcharhinus sp.⁴, Physogaleus minor⁴, Physogaleus secundus⁴ "Rhizoprionodon gantourensis" = Rhizoprionodon ganntourensis⁴ Arambourg 1952 sharpnose shark Carcharhinus sp.⁴ de Blainville 1816 requiem shark Physogaleus minor⁴ requiem shark Physogaleus secundus⁴ Winkler 1876 requiem shark
Carcharhiniformes - Triakidae	Galeorhinus sp.⁴, Galeorhinus cf. ypresiensis⁴ Galeorhinus sp.⁴ de Blainville and Prévost 1816 school shark Galeorhinus cf. ypresiensis⁴ Casier 1946 school shark
Lamniformes - Lamnidae	"Carcharocles auriculatus" = Carcharodon auriculatus⁹, Isurus praecursor⁴ "Carcharocles auriculatus" = Carcharodon auriculatus⁹ Blainville 1818 white shark Isurus praecursor⁴ Thurmond and Jones 1981 mackerel shark
Lamniformes -	Isurolamna cf. bajarunasi⁴ Isurolamna cf. bajarunasi⁴ mackerel shark
Lamniformes - Odontaspididae	Jaekelotodus aff. borystenicus⁴, Odontaspis winkleri⁴, Synodontaspis vincenti⁴, Hypotodus verticalis⁴ Jaekelotodus aff. borystenicus⁴ sand tiger shark Odontaspis winkleri⁴ Leriche 1905 sand tiger shark Synodontaspis vincenti⁴ sand tiger shark Hypotodus verticalis⁴ Agassiz 1843 sand tiger shark
Lamniformes - Mitsukurinidae	Striatolamia macrota⁴ Striatolamia macrota⁴ Agassiz 1838 goblin shark
Chimaeriformes - Edaphodontidae	Edaphodon sp.⁹ Edaphodon sp.⁹ Buckland 1838 chimaera

¹Danilov et al. 2011; ²Kuzmin and Zvonok 2021; ³Mayr and Zvonok 2012; ⁴Udovichenko 2009; ⁵Zvonok et al. 2010; ⁶Zvonok et al. 2013; ⁷Zvonok et al. 2015; ⁸Zvonok and Skutschas 2011; ⁹Zvonok and Удовиченко 2011; ¹⁰Звонок 2011