Villarroya (Pliocene of Spain)

Where: La Rioja, Spain (42.1° N, 2.1° W: paleocoordinates 42.2° N, 2.4° W)

• coordinate based on political unit

• local area-level geographic resolution

When: Pliocene (5.3 - 2.6 Ma)

• "Pliocene" according to Villalta (1952), "Villafranquiense" according to Villalta (1956)

Environment/lithology: terrestrial; red, sandy mudstone and red, sandy mudstone

• Upper: maroon sandy mudstone; Lower: light red sandy mudstone

Size class: macrofossils

Primary reference: J. F. Villalta Comella. 1952. Contribución al conocimiento de la fauna de mamíferos fósiles del Plioceno de Villaroya (Logroño). Boletín del Instituto Geológico y Minero de España 64:1-201 [J. Alroy/J. Wertheim/J. Wertheim]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 38892: authorized by John Alroy, entered by Jill Wertheim on 13.05.2004, edited by Hannah O'Regan

Creative Commons license: CC BY (attribution)

Taxonomic list

Mammalia
 Rodentia - Hystricidae
Hystrix primigenia2 Wagner 1848 porcupine
 Perissodactyla - Rhinocerotidae
Stephanorhinus etruscus4 Falconer 1868 rhinoceros
 Perissodactyla - Equidae
Hipparion rocinantis2 Hernández Pacheco 1921 hipparionine horse
 Artiodactyla - Cervidae
""Cervus" perrieri" = Praeelaphus perrieri1 Croizet and Jobert 1828 deer
Arvernoceros ardei2 Croizet and Jobert 1828 deer
Croizetoceros ramosus1 Croizet and Jobert 1828 deer
 Artiodactyla - Bovidae
Leptobos elatus1 antelope
Megalovis sp.2 Schaub 1924 antelope
"Hesperoceras merlae n. gen. n. sp." = Pliotragus ardeus5 Deperet 1883 antelope
 Carnivora - Felidae
Megantereon cultridens2 Cuvier 1824 cat
Homotherium crenatidens4 Fabrini 1890 cat
Acinonyx pardinensis1 Croizet and Jobert 1828 cheetah
Acinonyx pardinensis2 Croizet and Jobert 1828 cheetah
 Carnivora - Hyaenidae
Chasmaporthetes lunensis3 Del Campana 1914 hyaena
Hyaena perrieri2 Croizet and Jobert 1828 hyaena
 Carnivora - Mustelidae
Enhydrictis ardea2 Bravard 1828 mustelid carnivore
 Carnivora - Canidae
Canidae indet.2 Fischer de Waldheim 1817 canid
Nyctereutes megamastoides Pomel 1842 raccoon dog
Vulpes alopecoides2 Forsyth Major 1877 fox
 Proboscidea - Gomphotheriidae
Anancus arvernensis1 Croizet and Jobert 1828 gomphothere