Where: Worcestershire, United Kingdom (52.3° N, 2.1° W: paleocoordinates 52.3° N, 2.1° W)
• coordinate stated in text
• outcrop-level geographic resolution
When: Late/Upper Pleistocene (0.1 - 0.0 Ma)
• The Upton Warren site was radiocarbon dated at 41,500 +/-1200 (GRO 595) and 41,900 +/- 800 (GRO 1245). Both dates are in close agreement and together with a dated humate extract indicate that the organic horizon was deposited approximately 42,000 years B.P.
• bed-level stratigraphic resolution
Environment/lithology: terrestrial; unlithified peat
Size class: macrofossils
Collected in the 1950s
Primary reference: G. R. Coope, F. W. Shotton, I. Strachan and S. P. Dance. 1961. A Late Pleistocene fauna and flora from Upton Warren, Worcestershire. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences 244:379-421 [M. Clapham/M. Clapham]more details
Purpose of describing collection: general faunal/floral analysis
PaleoDB collection 163070: authorized by Matthew Clapham, entered by Matthew Clapham on 11.10.2014
Creative Commons license: CC BY (attribution)
Taxonomic list
unclassified | |
Insecta | |
"Deltocephalus ocellaris" = Errastunus ocellaris, "Strongylocephalus agrestis" = Stroggylocephalus agrestis, "Aphrodes bifasciatus" = Planaphrodes bifasciatus
"Deltocephalus ocellaris" = Errastunus ocellaris Fallen 1806 leafhopper
"Strongylocephalus agrestis" = Stroggylocephalus agrestis Fallen 1806 leafhopper
"Aphrodes bifasciatus" = Planaphrodes bifasciatus Linnaeus 1758 leafhopper | |
Saldula saltatoria Linnaeus 1758 shore bug
Salda littoralis Linnaeus 1758 shore bug | |
Hymenoptera indet. Linnaeus 1758 winged insect | |
"Galesus sp." = Psilus
"Galesus sp." = Psilus Panzer 1801 wasp | |
Formicidae indet. Latreille 1802 ant | |
Dolerus sp. Panzer 1801 wood wasp | |
Limnephilidae indet. Kolenati 1848 caddisfly | |
Phryganea obsoleta Hagen 1864 large caddisfly | |
"Nebria gyllenhali" = Nebria rufescens, Notiophilus aquaticus, Pelophila borealis, Agonum versutum, Agonum sahlbergi, Agonum sexpunctatum, Loricera pilicornis, Dyschirius sp., "Dyschirius arenosus" = Dyschirius thoracicus, "Dyschirius septentrionum" = Dyschiriodes nigricornis, "Dyschirius globosus" = Dyschiriodes globosus, Carabus sp., Carabus hortensis, "Feronia adstricta" = Pterostichus (Bothriopterus) adstrictus, "Feronia nigrita" = Pterostichus nigrita, Feronia diligens, "Harpalus cordatus" = Ophonus (Metophonus) cordatus, Harpalus sp., Calathus melanocephalus, Amara spp., Amara (Cyrtonotus) torrida, Amara interstitialis, Amara erratica, Amara municipalis, "Amara quenseli" = Amara (Paracelia) quenseli, Cymindis angularis, Patrobus septentrionis, Patrobus assimilis, Bembidion spp., Bembidion obliquum, Bembidion aeneum, "Bembidion lampros" = Bembidion (Metallina) lampros, Bembidion lunatum, "Bembidion bipunctatum" = Bembidion (Testedium) bipunctatum, Bembidion femoratum, Bembidion virens, Elaphrus cupreus, Elaphrus riparius, Elaphrus lapponicus, "Helobium multipunctatum" = Blethisa multipunctata
"Nebria gyllenhali" = Nebria rufescens Strøm 1768 ground beetle
Notiophilus aquaticus Linnaeus 1758 ground beetle
Pelophila borealis Paykull 1790 ground beetle
Agonum versutum Sturm 1824 ground beetle
Agonum sahlbergi Chaudoir 1850 ground beetle
Agonum sexpunctatum Linnaeus 1758 ground beetle
Loricera pilicornis Fabricius 1775 ground beetle
Dyschirius sp. Bonelli 1810 ground beetle
"Dyschirius arenosus" = Dyschirius thoracicus Rossi 1790 ground beetle
"Dyschirius septentrionum" = Dyschiriodes nigricornis Motschulsky 1844 ground beetle
"Dyschirius globosus" = Dyschiriodes globosus Herbst 1784 ground beetle
Carabus sp. Linnaeus 1758 ground beetle
Carabus hortensis Linnaeus 1758 ground beetle
"Feronia adstricta" = Pterostichus (Bothriopterus) adstrictus Eschscholtz 1823 ground beetle
"Feronia nigrita" = Pterostichus nigrita Paykull 1790 ground beetle
Feronia diligens Sturm 1824 ground beetle
"Harpalus cordatus" = Ophonus (Metophonus) cordatus Duftschmid 1812 ground beetle
Harpalus sp. Latreille 1802 ground beetle
Calathus melanocephalus Linnaeus 1758 ground beetle
Amara spp. Bonelli 1810 ground beetle
Amara (Cyrtonotus) torrida Panzer 1797 ground beetle
Amara interstitialis Dejean 1828 ground beetle
Amara erratica Duftschmid 1812 ground beetle
Amara municipalis Duftschmid 1812 ground beetle
"Amara quenseli" = Amara (Paracelia) quenseli Schönherr 1806 ground beetle
Cymindis angularis Gyllenhal 1810 ground beetle
Patrobus septentrionis Dejean 1828 ground beetle
Patrobus assimilis Chaudoir 1844 ground beetle
Bembidion spp. Latreille 1802 ground beetle
Bembidion obliquum Sturm 1825 ground beetle
Bembidion aeneum Germar 1824 ground beetle
"Bembidion lampros" = Bembidion (Metallina) lampros Herbst 1784 ground beetle
Bembidion lunatum Duftschmid 1812 ground beetle
"Bembidion bipunctatum" = Bembidion (Testedium) bipunctatum Linnaeus 1761 ground beetle
Bembidion femoratum Sturm 1825 ground beetle
Bembidion virens Gyllenhal 1827 ground beetle
Elaphrus cupreus Duftschmid 1812 ground beetle
Elaphrus riparius Linnaeus 1758 ground beetle
Elaphrus lapponicus Gyllenhal 1810 ground beetle
"Helobium multipunctatum" = Blethisa multipunctata Linnaeus 1758 ground beetle | |
Gyrinus minutus Fabricius 1798 whirligig beetle | |
Haliplus obliquus Fabricius 1787 crawling water beetle
Haliplus ruficollis De Geer 1774 crawling water beetle | |
Ilybius sp., Ilybius subaeneus, "Agabus nebulosus" = Agabus (Gaurodytes) nebulosus, "Agabus bipustulatus" = Agabus (Gaurodytes) bipustulatus, "Agabus congener" = Agabus (Acatodes) congener, "Agabus sturmii" = Agabus (Acatodes) sturmii, "Agabus arcticus" = Agabus (Acatodes) arcticus, Deronectes sp., "Deronectes griseostriatus" = Boreonectes griseostriatus, Hydroporus spp., Hydroporus planus, "Hydroporus arcticus" = Hydroporus notabilis, Hydroporus palustris, "Hygrotus impressopunctatus" = Hygrotus (Leptolambus) impressopunctatus, "Hygrotus cf. novemlineatus" = Hygrotus (Leptolambus) novemlineatus, Dytiscus marginalis, "Rantus sp." = Rhantus, "Rantus exsoletus" = Rhantus exsoletus, Colymbetes paykulli, Colymbetes fuscus
Ilybius subaeneus Erichson 1837 predaceous diving beetle
"Agabus nebulosus" = Agabus (Gaurodytes) nebulosus Forster 1771 predaceous diving beetle
"Agabus bipustulatus" = Agabus (Gaurodytes) bipustulatus Linnaeus 1767 predaceous diving beetle
"Agabus congener" = Agabus (Acatodes) congener Thunberg 1794 predaceous diving beetle
"Agabus sturmii" = Agabus (Acatodes) sturmii Gyllenhal 1808 predaceous diving beetle
"Agabus arcticus" = Agabus (Acatodes) arcticus Paykull 1798 predaceous diving beetle
"Deronectes griseostriatus" = Boreonectes griseostriatus De Geer 1774 predaceous diving beetle
Hydroporus spp. Clairville 1806 predaceous diving beetle
Hydroporus planus Fabricius 1781 predaceous diving beetle
"Hydroporus arcticus" = Hydroporus notabilis LeConte 1850 predaceous diving beetle
Hydroporus palustris Linnaeus 1761 predaceous diving beetle
"Hygrotus impressopunctatus" = Hygrotus (Leptolambus) impressopunctatus Schaller 1783 predaceous diving beetle
"Hygrotus cf. novemlineatus" = Hygrotus (Leptolambus) novemlineatus Stephens 1829 predaceous diving beetle
Dytiscus marginalis Linnaeus 1758 predaceous diving beetle
"Rantus sp." = Rhantus Dejean 1833 predaceous diving beetle
"Rantus exsoletus" = Rhantus exsoletus Forster 1771 predaceous diving beetle
Colymbetes paykulli Erichson 1837 predaceous diving beetle
Colymbetes fuscus Linnaeus 1758 predaceous diving beetle | |
Limnebius truncatellus Thunberg 1794 minute moss beetle
Ochthebius dilatatus Stephens 1829 minute moss beetle | |
Leiodes spp. Latreille 1796 round fungus beetle
Hydnobius punctatus Sturm 1807 round fungus beetle
Catops sp. Paykull 1798 round fungus beetle | |
Xantholinus linearis, Quedius spp., "Staphylinus olens" = Ocypus olens, "Staphylinus pubescens" = Dinothenarus pubescens, Staphylinus sp., Philonthus spp., Philonthus linki, Philonthus rotundicollis, Thanatophilus dispar, Aclypea opaca, Bledius spp., Bledius littoralis, Platystethus spp., Platystethus nitens, Anotylus gibbulus2, Olophrum spp., Olophrum aff. boreale, Geodromicus plagiatus, Arpedium sp., Acidota crenata, Omalium sp., Tachinus "sp. A", Tachinus spp.
Xantholinus linearis Olivier 1795 rove beetle
Quedius spp. Stephens 1829 rove beetle
"Staphylinus olens" = Ocypus olens Müller 1764 rove beetle
"Staphylinus pubescens" = Dinothenarus pubescens De Geer 1774 rove beetle
Philonthus spp. Curtis 1829 rove beetle
Philonthus linki Solsky 1867 rove beetle
Philonthus rotundicollis Menetries 1832 rove beetle
Thanatophilus dispar Herbst 1793 rove beetle
Aclypea opaca Linnaeus 1758 rove beetle
Bledius spp. Leach 1819 rove beetle
Bledius littoralis Heer 1839 rove beetle
Platystethus spp. Mannerheim 1830 rove beetle
Platystethus nitens Sahlberg 1832 rove beetle
Olophrum spp. Erichson 1839 rove beetle
Olophrum aff. boreale Paykull 1792 rove beetle
Geodromicus plagiatus Fabricius 1798 rove beetle
Arpedium sp. Erichson 1839 rove beetle
Acidota crenata Fabricius 1792 rove beetle
Omalium sp. Gravenhorst 1802 rove beetle
Tachinus "sp. A" Gravenhorst 1802 rove beetle
Tachinus spp. Gravenhorst 1802 rove beetle | |
Geotrupes sp. Latreille 1796 earth-boring dung beetle | |
Aphodius sp., Aphodius spp., Aphodius plagiatus, Aphodius rufipes, Aphodius fimitarius, Aphodius fossor, Aphodius holdereri1, Aphodius cf. distinctus, "Aegialia sabuleti" = Aegialia (Psammoporus) sabuleti
Aphodius spp. Hellwig 1798 scarab beetle
Aphodius plagiatus Linnaeus 1767 scarab beetle
Aphodius rufipes Linnaeus 1758 night-flying dung beetle
Aphodius fimitarius Linnaeus 1758 scarab beetle
Aphodius fossor Linnaeus 1758 scarab beetle
Aphodius cf. distinctus Müller 1776 scarab beetle
"Aegialia sabuleti" = Aegialia (Psammoporus) sabuleti Panzer 1797 scarab beetle | |
Georissus crenulatus, Sphaeridium sp., Cercyon spp., Cercyon melanocephalus, Cercyon tristis, Cercyon ustulatus, Cryptopleurum minutum, Hydrobius fuscipes, Laccobius minutus
Georissus crenulatus Rossi 1794 water scavenger beetle
Cercyon spp. Leach 1817 water scavenger beetle
Cercyon melanocephalus Linnaeus 1758 water scavenger beetle
Cercyon tristis Illiger 1801 water scavenger beetle
Cercyon ustulatus Preyssler 1790 water scavenger beetle
Cryptopleurum minutum Fabricius 1775 water scavenger beetle
Hydrobius fuscipes Linnaeus 1758 water scavenger beetle
Laccobius minutus Linnaeus 1758 water scavenger beetle | |
"Helophorus aquaticus" = Helophorus (Meghelophorus) aquaticus, Helophorus nubilus, Helophorus brevipalpis
"Helophorus aquaticus" = Helophorus (Meghelophorus) aquaticus Linnaeus 1758 water scavenger beetle
Helophorus nubilus Fabricius 1777 water scavenger beetle
Helophorus brevipalpis Bedel 1881 water scavenger beetle | |
Caenocara bovistae Hoffmann 1803 spider beetle | |
"Cantharis figurata" = Cantharis (Cantharis) figurata, "Cantharis rufa" = Cantharis (Cantharis) rufa, "Cantharis cf. paludosa" = Cantharis (Cantharis) paludosa
"Cantharis figurata" = Cantharis (Cantharis) figurata Mannerheim 1843 soldier beetle
"Cantharis rufa" = Cantharis (Cantharis) rufa Linnaeus 1758 soldier beetle
"Cantharis cf. paludosa" = Cantharis (Cantharis) paludosa Fallen 1807 soldier beetle | |
Corymbites melancholicus Fabricius 1798 click beetle
Hypnoidus riparius Fabricius 1792 click beetle | |
"Simplocaria semistriata" = Simplocaria (Simplocaria) semistriata, Morychus aeneus, Cytilus sericeus, Byrrhus sp., Byrrhus pustulatus, Byrrhus pilula, Syncalypta sp.
"Simplocaria semistriata" = Simplocaria (Simplocaria) semistriata Fabricius 1794 pill beetle
Morychus aeneus Fabricius 1775 pill beetle
Cytilus sericeus Forster 1771 pill beetle
Byrrhus pustulatus Forster 1771 pill beetle
Byrrhus pilula Linnaeus 1758 pill beetle
Syncalypta sp. Dillwyn 1829 pill beetle | |
Dryops ernesti des Gozis 1886 long toed water beetle | |
Heterocerus fusculus Kiesenwetter 1843 Variegated mud-loving beetle
Heterocerus hispidulus Kiesenwetter 1843 Variegated mud-loving beetle
Heterocerus fenestratus Thunberg 1784 Variegated mud-loving beetle | |
Kateretes rufilabris Latreille 1807 beetle | |
Antherophagus pallens Linnaeus 1758 silken fungus beetle | |
Anthicus sp. Paykull 1798 ant like flower beetle | |
Hippodamia sp., "Coccinella undecimpunctata" = Coccinella (Spilota) undecimpunctata, "Coccinella septempunctata" = Coccinella (Coccinella) septempunctata
"Coccinella undecimpunctata" = Coccinella (Spilota) undecimpunctata Linnaeus 1758 ladybird beetle
"Coccinella septempunctata" = Coccinella (Coccinella) septempunctata Linnaeus 1758 ladybird beetle | |
Phyllobius spp., Otiorrhynchus rugifrons, Otiorrhynchus ligneus, Otiorrhynchus arcticus, Sitona spp., Sitona lepidus, "Litodactylus leucogaster" = Phytobius leucogaster, Rhinoncus castor, "Ceuthorhynchus sp." = Ceutorhynchus, Bagous tempestivus, Rhynchaenus foliorum, "Notaris bimaculatus" = Tournotaris bimaculatus, Grypus equiseti, Notaris aethiops, Notaris acridulus, Mecinus collaris, Orobitis cyaneus, Phytonomus dauci
Phyllobius spp. Schoenher 1824 snout beetle
Otiorrhynchus rugifrons Gyllenhal 1813 snout beetle
Otiorrhynchus ligneus Olivier 1807 snout beetle
Otiorrhynchus arcticus Fabricius 1780 snout beetle
Sitona spp. Germar 1817 snout beetle
Sitona lepidus Gyllenhal 1834 snout beetle
"Litodactylus leucogaster" = Phytobius leucogaster Marsham 1802 snout beetle
Rhinoncus castor Fabricius 1792 snout beetle
"Ceuthorhynchus sp." = Ceutorhynchus Germar 1824 snout beetle
Bagous tempestivus Herbst 1795 snout beetle
Rhynchaenus foliorum Müller 1764 snout beetle
"Notaris bimaculatus" = Tournotaris bimaculatus Fabricius 1787 snout beetle
Grypus equiseti Fabricius 1775 snout beetle
Notaris aethiops Fabricius 1792 snout beetle
Notaris acridulus Linnaeus 1758 snout beetle
Mecinus collaris Germar 1821 snout beetle
Orobitis cyaneus Linnaeus 1758 snout beetle
Phytonomus dauci Olivier 1807 snout beetle | |
Apion spp. Herbst 1797 weevil | |
Galeruca tanaceti, Crepidodera interpunctata, Chaetocnema spp., Donacia thalassina, "Phaedon (Neophaedon) pyritosus" = Neophaedon pyritosus, Chrysolina spp., Chrysolina staphylea, Chrysolina marginata, Chrysolina limbata, Gastrophysa viridula, Entomoscelis adonidis, Phaedon armoraciae
Galeruca tanaceti Linnaeus 1758 leaf beetle
Crepidodera interpunctata Motschulsky 1859 flea beetle
Chaetocnema spp. Stephens 1831 flea beetle
Donacia thalassina Germar 1811 leaf beetle
"Phaedon (Neophaedon) pyritosus" = Neophaedon pyritosus Rossi 1792 leaf beetle
Chrysolina spp. Moltschulsky 1860 leaf beetle
Chrysolina staphylea Linnaeus 1758 leaf beetle
Chrysolina marginata Linnaeus 1758 leaf beetle
Chrysolina limbata Fabricius 1775 leaf beetle
Gastrophysa viridula De Geer 1775 leaf beetle
Entomoscelis adonidis Pallas 1771 leaf beetle
Phaedon armoraciae Linnaeus 1758 leaf beetle | |
| |
Sialis sp. Latreille 1802 alderfly | |
Diptera indet. Linnaeus 1758 fly | |
Dilophus sp. Meigen 1803 march fly | |
| |
Glyptotendipes sp. Kieffer 1913 non biting midge
Chironomus sp. Meigen 1803 non biting midge
Cryptochironomus defectus Kieffer 1913 non biting midge | |
Tipulidae indet. Latreille 1802 crane fly | |
Arachnida | |
"Pelops sp." = Eupelops
"Pelops sp." = Eupelops Ewing 1917 oribatid mite | |
Erigone sp. Audouin 1826 money spider/sheet weaver | |
Lycosidae indet. Sundevall 1833 wolf spider | |
Acari indet. Leach 1817 mite | |
Ostracoda | |
Cypris pubera Müller 1776 ostracod
Erpetocypris reptans Baird 1835 ostracod | |
Ilyocypris bradyi Sars 1890 ostracod
Ilyocypris gibba Ramdohr 1808 ostracod | |
Candona candida Müller 1776 ostracod
Candona neglecta Sars 1887 ostracod | |
Bivalvia | |
Pisidium casertanum Poli 1791 pea clam
Pisidium nitidum Jenyns 1832 pea clam | |
Gastropoda | |
Montia fontana Linnaeus 1753 volute | |
Planorbis leucostoma Millet 1813 snail
Planorbis crista Linnaeus 1758 snail
Planorbis laevis Alder 1838 snail | |
Lymnaea peregra, "Lymnaea truncatula" = Galba truncatula
Lymnaea peregra Müller 1774 snail
"Lymnaea truncatula" = Galba truncatula Müller 1774 snail | |
Pupilla muscorum Linnaeus 1758 snail | |
Mammalia | |
Microtus sp. Schrank 1798 vole
Dicrostonyx henseli lemming | |
Amphibia | |
Rana temporaria Linnaeus 1758 common frog | |
Actinopteri | |
Gasterosteus aculeatus Linnaeus 1758 three-spined stickleback | |
Pinopsida | |
Pinus sylvestris pine | |
Angiospermae | |
Zannichellia palustris Linnaeus 1753 horned pondweed | |
Triglochin maritima Linnaeus 1753 arrowgrass | |
Betula sp. Linnaeus 1753 birch | |
Salix herbacea willow
Salix cf. phylicifolia willow | |
Potentilla palustris cinquefoil
Potentilla sterilis cinquefoil
Potentilla spp. Linnaeus 1753 cinquefoil
Potentilla anserina Linnaeus 1753 cinquefoil
Potentilla reptans Linnaeus 1753 cinquefoil | |
| |
Capsella bursapastoris, "Cruciferae indet." = Brassicaceae
"Cruciferae indet." = Brassicaceae Burnett 1835 | |
Dicotyledoneae | |
Primula sp. Linnaeus 1753 primrose
Primula cf. farinosa Linnaeus 1753 primrose | |
Hippuris vulgaris Linnaeus 1753 mare's tail | |
Angiospermae | |
Thalictrum sp., Thalictrum alpinum, Thalictrum minus, Ranunculus (Batrachium) spp., Ranunculus repens, Ranunculus acris, Ranunculus sceleratus
Thalictrum sp. Linnaeus 1753 meadow-rue
Thalictrum alpinum Linnaeus 1753 meadow-rue
Thalictrum minus Linnaeus 1753 meadow-rue
Ranunculus (Batrachium) spp. Linnaeus 1753 buttercup
Ranunculus repens Linnaeus 1753 buttercup
Ranunculus acris Linnaeus 1753 buttercup
Ranunculus sceleratus Linnaeus 1753 buttercup | |
Polygonum aviculare knotweed
Polygonum amphibium Linnaeus 1753 knotweed
Rumex acetosella Linnaeus 1753 dock | |
Atriplex sp. Linnaeus 1753 orache | |
Caryophyllaceae indet. Jussieu 1789 carnation
Dianthus cf. carthusianorum Linnaeus 1753 carnation
Viscaria alpina carnation
Cerastium arvense Linnaeus 1753 chickweed | |
Campanula cf. rotundifolia Linnaeus 1753 bellflower | |
Artemisia sp. Linnaeus 1753 wormwood
"Tubuliflorae indet." = Asteraceae Berchtold and Presl 1820 daisy
Taraxacum officinale dandelion | |
Pedicularis palustris Linnaeus 1753 lousewort | |
Parnassia palustris Linnaeus 1753 grass of Parnassus | |
Plantago maritima Linnaeus 1753 plantain | |
Myriophyllum alternifolium water milfoil
Myriophyllum spicatum water milfoil
Myriophyllum sp. Linnaeus 1753 water milfoil | |
Cyperaceae indet., Schoenoplectus lacustris, Carex dioica, Carex flava, Carex rostrata, Carex spp., Eleocharis palustris, Eleocharis quinqueflora, Eleocharis uniglumis
Cyperaceae indet. Jussieu 1789 sedge
Schoenoplectus lacustris sedge
Carex dioica sedge
Carex flava sedge
Carex rostrata sedge
Carex spp. Linnaeus 1753 sedge
Eleocharis palustris sedge
Eleocharis quinqueflora sedge
Eleocharis uniglumis sedge | |
"Gramineae indet." = Poaceae
"Gramineae indet." = Poaceae Barnhart 1895 true grass | |
Linum | |
Linum praecursor flax | |
Scleranthus annuus | |
Scleranthus annuus Linnaeus 1753 knotweed | |
Leontodon | |
Leontodon autumnalis hawkbit | |
Lycopodiopsida | |
Selaginella selaginoides spikemoss | |
Isoetes cf. hystrix quillwort | |
Draba incana | |
Draba incana Linnaeus 1753 Whitlow grass | |
Glaux maritima | |
Glaux maritima Linnaeus 1753 | |
Viola | |
Viola sp. Linnaeus 1753 violet | |
Scabiosa columbaria | |
Scabiosa columbaria Linnaeus 1753 pincushion flower | |
Rhinanthus | |
cf. Rhinanthus sp. Linnaeus 1753 | |
Crypticus quisquilius | |
Crypticus quisquilius Linnaeus 1761 |