Shanwang Flora, Linqu (Miocene of China)

Also known as Locality N-4

Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)

• From He et al. 2011: "40Ar/39Ar ages obtained from basalts of the Niushan and Yaoshan Formations, which underlie and overlie the Shanwang Formation, are 21.0 ± 2.5 Ma (2σ, full external error) and 17.3 ± 1.5 Ma (2σ, full external error), respectively. The 40Ar/39Ar age of the basalt in the Shanwang Formation is 17–18 Ma. Given the age constraints of the basalts of the Yaoshan and Shanwang Formations, the age of the Shanwang biota is estimated to be ca. 17 Ma, late Burdigalian of the Early Miocene,". Yu et al. 2021 support this, stating "Paleomagnetic and radio isotope dating data shows that the diatomaceous shale was deposited during Chron C5En, which places it at approximately 18.5 Ma during the Burdigalian stage of the Early Miocene, rather than in the Middle Miocene as previously thought."

• group of beds-level stratigraphic resolution

Environment/lithology: marine; diatomite

Size class: macrofossils

Preservation: mold/impression, adpression

Reposited in the NIGPAS

Primary reference: Y.-S. Liu, S.-X. Guo, and D. K. Ferguson. 1996. A catalogue of Cenozoic megafossil plants in China. Palaeontographica Abteilung B 238:141-179 [B. Allen/B. Allen]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 223803: authorized by Bethany Allen, entered by Bethany Allen on 01.12.2021

Creative Commons license: CC BY (attribution)

Taxonomic list

 Polypodiales - Polypodiidae
Pteris sp. Linnaeus 1753
 Ceratophyllales - Ceratophyllaceae
 Poales - Cyperaceae
Cyperacites sp. Schimper 1870 sedge
 Poales - Poaceae
Graminites sp. Geinitz 1865 grass
 Laurales - Lauraceae
 Proteales - Sabiaceae
 Violales - Flacourtiaceae
Carrierea calycina Franchet 1896
 Saxifragales - Saxifragaceae
 Saxifragales - Hamamelidaceae
 Caryophyllales - Polygonaceae
 Cornales - Cornaceae
 Dipsacales - Caprifoliaceae
Lonicera hispida Pallas 1819 honeysuckle
Lonicera cf. japonica Thunberg 1784 honeysuckle
 Apiales - Araliaceae
 Lamiales - Oleaceae
 Lamiales - Bignoniaceae
Catalpa szei trumpet vine
 Lamiales - Scrophulariaceae
 Ericales - Ebenaceae
Diospyros miokaki Awasthi and Prasad 1990
 Vitales - Vitaceae
Vitis romanetii Romanet du Caillaud 1883 grapevine
 Rosids - Moraceae
 Fabales - Fabaceae
Albizia miokalkora Hu and Chaney 1940 legume
"Podogonium knorrii" = Podocarpium podocarpum Braun 1845 legume
 Fagales - Fagaceae
 Fagales - Juglandaceae
 Fagales - Betulaceae
 Rosales - Ulmaceae
Celtis angusta hackberry
Celtis bungeana Blume 1852 hackberry
Zelkova ungeri Kovats 1865 zelkova
Ulmus macrocarpa Hance 1868 elm
 Rosales -
 Rosales - Pomaceae
 Rosales - Rhamnaceae
"Berchemia miofloribunda" = Berhamniphyllum miofloribundum Hu and Chaney 1940 buckthorn
 Rosales - Papilionaceae
 Rosales - Cannabaceae
 Rosales - Rosaceae
 Malpighiales - Salicaceae
Salix angusta Braun 1851 willow
Populus balsamoides Göppert 1855 poplar
Populus latior Braun 1835 poplar
Populus simonii Carrière 1867 poplar
Populus glandulifera Heer 1856 poplar
 Malpighiales - Euphorbiaceae
 Celastrales - Celastraceae
 Crossosomatales - Stachyuraceae
 Huerteales - Tapisciaceae
 Malvales - Malvaceae
 Malvales - Tiliaceae
 Malvales - Sterculiaceae
 Sapindales - Sapindaceae
Koelreuteria macrocarpa golden rain tree
Acer nordenskioeldi Nathorst 1883 maple
Acer subpictum Saporta 1873 maple
Acer miofranchetii Hu and Chaney 1940 maple
Acer diabolicum Blume 1864 maple
 Sapindales - Rutaceae
 Sapindales - Meliaceae
 Sapindales - Simaroubaceae
 Sapindales - Anacardiaceae
 Alismatales - Potamogetonaceae
Potamogeton sp. Linnaeus 1753 pondweed
 Magnoliales - Magnoliaceae
Magnolia miocenica Hu and Chaney 1938 magnolia
 Pinales - Pinaceae
Keteleeria ezoana Tanai 1961
 Hypnales - Neckeraceae
Neckera shanwanica n. sp. Wu and Feng 1978