Where: Basque County, Spain (42.7° N, 2.9° W: paleocoordinates 41.3° N, 6.1° W)
• coordinate based on nearby landmark
When: MP 18 (37.2 - 33.9 Ma)
• biozone MP 18, early middle Ludian (Priabonian)
•European Headonian Land Mammal Age corresponding to the reference level MP 18 (Astibia et al., 2000; Badiola, 2004; Badiola et al., 2009b).
• group of beds-level stratigraphic resolution
Environment/lithology: lacustrine; carbonaceous marl and limestone
•The depositional setting was a shallow, low-gradient freshwater lake margin with a peripheral swamp (Iriarte et al., 2003; Badiola et al., 2009a).
•the fossiliferous beds, which consist of palustrine- lacustrine coal-bearing marls and marlstones, are located in one of the oldest lacustrine systems in the Miranda-Treviño Basin (Basque-Cantabrian Region) (Astibia et al., 2000)
Size classes: macrofossils, mesofossils
Primary reference: A. Badiola, H. Astibia, A. Aranburu, X. Pereda Suberbiola, X. Murelaga, C. Sese, M. A. Cuesta, S. Moya-Sola, J. I. Baceta and M. Kohler. 1999. A new continental vertebrate locality from the Upper Eocene of Zambrana (Miranda-Trevino Basin, Alava, Basque Country). IV European Workshop on vertebrate Paleontology, Albarracin, Spain 19 [J. Alroy/E. Leckey/E. Leckey]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 38817: authorized by John Alroy, entered by Erin Leckey on 06.05.2004, edited by Richard Butler and Grace Varnham
Creative Commons license: CC BY (attribution)
Taxonomic list
Mammalia | |
Microchoerus aff. erinaceus6 Wood 1844 tarsier Right mandible fragment with p3-m3 (MCNA 14518); right mandible fragment with p4-m3 (MCNA 14519).
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"Paranchilophus remyi" = Anchilophus (Paranchilophus) remyi6, Pachynolophus zambranensis n. sp.5, Iberolophus arabensis n. gen. n. sp.4, Leptolophus "new species"6, Leptolophus sp.2, Leptolophus franzeni n. sp.7, Leptolophus cuestai n. sp.7, Palaeotherium "new species"
"Paranchilophus remyi" = Anchilophus (Paranchilophus) remyi6 Casanovas and Santafé 1989 odd-toed ungulate fragment of mandible with teeth
Pachynolophus zambranensis n. sp.5 Badiola et al. 2005 odd-toed ungulate MCNA 10657 (holotype). MCNA 10658-10661 (paratypes). 21 referred specimens
Iberolophus arabensis n. gen. n. sp.4 Badiola and Cuesta 2008 odd-toed ungulate MCNA 11953 (holotype). Paratypes: MCNA 10183, skull fragment, showing the right and left M1–M3 molar series; MCNA 11888, left P3–4; MCNA 11906, left P1–2; MCNA 13372, left p4; and MCNA 13373, right p1. Referred material: —MCNA 11904, right P3–4; MCNA 11887, 11893, left P3–4; MCNA 13375, fragment of the left P3–4; MCNA 11905, fragment of the left P3–4; MCNA 11950, right P1-2 (incomplete lingually); MCNA 13374, fragment of the right upper premolar; MCNA 11951, fragment of the upper premolar; MCNA 11952, fragment of the lower premolar
Leptolophus franzeni n. sp.7 Perales-Gogenola et al. 2021 odd-toed ungulate Holotype: UPV/EHU-Z4.1092, cranium fragment with right and left C–P2–P3–P4–M1–M2–(M3) series. Fragmented right P2 and P4, and left P2; MCNA 11848, left M3; MCNA 11849, right M3; UPV/EHU- Z4.1041 fragment of right hemimandible with a par- tially erupted m3.
Leptolophus cuestai n. sp.7 Perales-Gogenola et al. 2021 odd-toed ungulate Holotype: MCNA 9948, right M2; MCNA 9947, right M1; MCNA 9002, incomplete left hemimandible preserving the m1–(m2)–(m3) series; MCNA 9949, incomplete right hemimandible showing the p4– m1–(m2)–(m3) series
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Amphibia | |
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