Also known as Hambach 6C
Where: Germany (50.9° N, 6.5° E: paleocoordinates 51.1° N, 5.0° E)
• coordinate based on nearby landmark
• outcrop-level geographic resolution
When: Frimmersdorf Member (Ville Formation), MN 5 (16.0 - 13.7 Ma)
• biozone MN 5, in the middle part of the main lignite seam, later authors list horizon as 6Ca/b
•JA: "6C" entered as local bed, with no local section name
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•Hambach 6C local fauna can be correlated with the upper part of mammalian Neogene unit MN 5, indicating an absolute age-range of 16.0–15.2 Ma for this late Orleanian, early middle Miocene fauna (Mörs et al. 2000; Mo€rs 2002; Mörs & Kalthoff 2004).
• bed-level stratigraphic resolution
Environment/lithology: paralic; lignite and lenticular, sandy claystone
•
•channel fill and floodplain deposits (horizon 6C according to the local lithostratigraphy) within the middle Miocene Frimmersdorf lignite seam (Mo€rs et al. 2000; Sch€afer et al. 2004).
Size classes: macrofossils, mesofossils
Collection methods: this collection almost certainly comes from the same unit and same general area as Tiffney's collections Hambach 8251-13349
•
•Macaluso et al. 2021: The new taxon is described based on material stored in the Steinmann Institute of the Rheinische Friedrich-Wilhelms-Universität Bonn, Germany.
Primary reference: T. Mors, F. Hocht, and B. Wutzler. 2000. Die erst Wirbeltierfauna aus der miozanen Braunkohle der Niederrheinischen Bucht (Ville-Schichten, Tagebau Hambach) [The first vertebrate fauna from the Miocene Ville Series of the Lower Rhine Embayment (Hambach open cast mine, western Germany)]. Paläontologische Zeitschrift 74(1/2):145-170 [J. Alroy/E. Leckey/J. Alroy]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 46955: authorized by John Alroy, entered by Erin Leckey on 27.01.2005, edited by Terri Cleary, Grace Varnham and Mark Uhen
Creative Commons license: CC BY (attribution)
Taxonomic list
Actinopteri | |
Barbus sp. Cuvier and Cloquet 1816 carp-like fish
Palaeoleuciscus sp. Obrhelová 1969 carp-like fish
Palaeotinca sp. Obrhelová 1969 carp-like fish | |
Sparidae indet. Cuvier 1817 | |
Atractoscion sp. Gill 1862 | |
Aves | |
Aves indet. Linnaeus 1758 bird | |
Reptilia | |
Diplocynodon sp. Pomel 1847 crocodilian | |
Anguidae "indet. 1"1 Gray 1825 squamates IPB-HaH 3700, twenty osteoderms; IPB-HaH 3709, caudal vertebra
Ophisaurus sp. Daudin 1803 glass lizard
Pseudopus cf. ahnikoviensis1 Klembara 2012 legless lizard IPB-HaH 3706, right dentary; IPB-HaH 3707, left dentary
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Lacertidae "indet. 1"1 Bonaparte 1831 squamates IPB-HaH 3701, left maxilla; IPB-HaH 3702 and 3703, two left dentaries
Lacertidae "indet. 2"1 Bonaparte 1831 squamates IPB-HaH 3704, premaxilla; IPB-HaH 3705, left dentary
Lacerta sp. Linnaeus 1758 squamates | |
Chameleontidae indet., Chamaeleonidae "tooth morphotype 1"1, Chamaeleonidae "tooth morphotype 2"1, Chamaeleo aff. andrusovi1
Chamaeleonidae "tooth morphotype 1"1 Gray 1825 chameleon IPB-HaH 3901, right maxilla; IPB-HaH 3902, left maxilla
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Bavarioboa sp.1 Szyndlar and Schleich 1993 boa IPB-HaH 3825, 3807, 3013, 3814, 3818, five trunk vertebrae
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Eoanilius sp.1 Rage 1974 snake IPB-HaH 3801, 3802 and 3824, five trunk vertebrae (remainder prefixed by latter number)
Colubroidea indet.1 Oppel 1811 snake IPB-HaH 3829; 3836; 3840; 3848; 3851; 3856, thirty-six caudal vertebrae
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Vipera ""oriental viper" sp."1 Laurenti 1768 pit viper IPB-HaH 3846; 3809; 3811; 3812; 3819; 3860, eight trunk vertebrae
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"Natricinae" indet.1, "Colubrinae" "indet. 1"1, "Coluber" sp.1, Texasophis sp.1, Telescopus sp.1, Natrix "sp. 1"1
"Natricinae" indet.1 Bonaparte 1838 colubrid snake IPB-HaH 3827; 3834; 3837; 3849; 3854; 3821, nine trunk vertebrae
"Colubrinae" "indet. 1"1 Oppel 1811 colubrid snake IPB-HaH 3828; 3835; 3838; 3844; 3805; 3806; 3855, fifty seven trunk vertebrae
"Coluber" sp.1 Linnaeus 1758 racer IPB-HaH 3800; 3826; 3831; 3803; 3804; 3852; 3808; 3857; 3815; 3816; 3817; 3820; 3822; 3823, forty two trunk vertebrae
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Mammalia | |
Amphiperatherium frequens von Meyer 1846 marsupial | |
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cf. Prosantorhinus sp. Heissig 1973 rhinoceros
Brachypotherium brachypus Lartet 1848 rhinoceros | |
Anchitherium aurelianense Cuvier 1825 anchitheriine horse | |
Cetacea indet. Brisson 1762 whale | |
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Dorcatherium guntianum von Meyer 1846 chevrotain
Dorcatherium naui Kaup 1833 chevrotain | |
Amphimoschus pontileviensis Bourgeois 1873 ruminant | |
Lagomeryx ruetimeyeri Thenius 1949 ruminant
Dicrocerus elegans Lartet 1837 ruminant | |
Orygotherium escheri von Meyer 1838 antelope | |
Hyotherium soemmeringi von Meyer 1834 pig | |
Taucanamo sansaniense Lartet 1851 even-toed ungulate | |
? Protictitherium sp. Kretzoi 1938 hyaena | |
Amphicyonidae indet. Haeckel 1886 bear-dog
Pseudocyon steinheimensis Fraas 1885 bear-dog | |
"Ursavus elmensis" = Ballusia elmensis
"Ursavus elmensis" = Ballusia elmensis Stehlin 1917 bear | |
Amphictis sp. Pomel 1853 red panda | |
Mustelidae indet. Fischer 1817 mustelid carnivore
Trocharion albanense Forsyth-Major 1903 mustelid carnivore | |
Potamotherium miocenicum Peters 1868 carnivoran | |
"Talpini indet." = Talpidae, "Talpidae indet." = Talpidae7, Desmaninae indet., Mygalea cf. antiqua, Proscapanus sansaniensis
"Talpini indet." = Talpidae Fischer de Waldheim 1817 mole
Desmaninae indet. Thomas 1912 mole
Mygalea cf. antiqua Pomel 1848 mole
Proscapanus sansaniensis Lartet 1851 mole | |
Soricidae indet. Fischer 1814 shrew
Dinosorex aff. zapfei Engesser 1975 shrew
Heterosorex sp. Gaillard 1915 shrew
Allosorex gracilidens Viret and Zapfe 1951 red-toothed shrew | |
Plesiosorex germanicus Seemann 1938 placental | |
"Mioechinus" sansaniensis Lartet 1851 hedgehog
Lanthanotherium aff. sansaniense hedgehog | |
Chainodus aff. intercedens Müller 1967 placental
Plesiodimylus chantrei Gaillard 1899 placental | |
Democricetodon mutilus, Karydomys wigharti n. sp.6, Megacricetodon collongensis, Fahlbuschia koenigswaldi, Eumyarion cf. weinfurteri, Eumyarion cf. bifidus, Lartetomys cf. zapfei
Democricetodon mutilus Fahlbusch 1964 mouse
Megacricetodon collongensis Mein 1958 mouse
Fahlbuschia koenigswaldi Freudenthal 1963 mouse
Eumyarion cf. weinfurteri Schaub and Zapfe 1953 mouse
Eumyarion cf. bifidus Fahlbusch 1964 mouse
Lartetomys cf. zapfei Mein and Freudenthal 1971 mouse | |
Neocometes cf. similis Fahlbusch 1966 mouse | |
Eomyidae indet. Winge 1887 rodent
Eomyops cf. catalaunicus rodent
Keramidomys carpathicus Schaub and Zapfe 1953 rodent | |
Steneofiber depereti Mayet 1908 beaver
Anchitheriomys sp. Roger 1885 beaver
Anchitheriomys suevicus Schlosser 1884 beaver | |
Myoglis meini, Glirudinus sp., Glirudinus undosus, "Eomuscardinus sp." = Muscardinus, Eomuscardinus sansaniensis, Bransatoglis cf. astaracensis, Paraglirulus werenfelsi, Paraglirulus cf. agelakisi, Microdyromys cf. legidensis, Miodyromys aegercii
Myoglis meini de Bruijn 1966 dormouse
Glirudinus sp. de Bruijn 1966 dormouse
Glirudinus undosus Mayr 1979 dormouse
"Eomuscardinus sp." = Muscardinus Kaup 1829 dormouse
Eomuscardinus sansaniensis Lartet 1851 dormouse
Bransatoglis cf. astaracensis Baudelot 1970 dormouse
Paraglirulus werenfelsi Engesser 1972 dormouse
Paraglirulus cf. agelakisi Van der Meulen and De Bruijn 1982 dormouse
Microdyromys cf. legidensis Daams 1981 dormouse
Miodyromys aegercii Baudelot 1972 dormouse | |
Anomalomys minor Fejfar 1972 rodent | |
Hylopetes sp., Spermophilinus aff. bredai, Blackia miocaenica, Palaeosciurus sutteri, ? Ratufa sp., Miopetaurista lappi, Heteroxerus sp.
Hylopetes sp. Thomas 1908 squirrel
Spermophilinus aff. bredai von Meyer 1848 squirrel
Blackia miocaenica Mein 1970 squirrel
Palaeosciurus sutteri Ziegler and Fahlbusch 1986 squirrel
? Ratufa sp. Gray 1867 Asian giant squirrel
Miopetaurista lappi Mein 1958 squirrel
Heteroxerus sp. Stehlin and Schaub 1951 squirrel | |
Ochotonidae indet. Thomas 1897 pika | |
Pliopithecidae indet. Zapfe 1961 monkey
Pliopithecus antiquus Blainville 1839 monkey | |
Rhinolophus cf. delphinensis Gaillard 1899 horseshoe bat | |
Eptesicus cf. campanensis Baudelot 1970 house bat | |
Amphibia | |
Latonia seyfriedi von Meyer 1843 frog | |
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Andrias scheuchzeri, Proteidae indet., Euronecturus grogu n. sp.5, Salamandridae indet., Chelotriton paradoxus, cf. Salamandra sp.
Andrias scheuchzeri Tschudi 1837 giant salamander
Proteidae indet. Hogg 1838 mudpuppy
Euronecturus grogu n. sp.5 Macaluso et al. 2021 mudpuppy Holotype: IPB-HaH 2121, an almost complete atlas; IPB-HaH 2120, atlas; IPB-HaH 2150, atlas; IPB-HaH 2119, atlas; IPB-HaH 2165, atlas
Chelotriton paradoxus Pomel 1853 newt
cf. Salamandra sp. Laurenti 1768 salamander | |
Chondrichthyes | |
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Scyliorhinus distans Probst 1879 cat shark | |
Physogaleus sp. Cappetta 1980 requiem shark | |
Carcharias cf. reticulata, "Odontaspis vorax" = Carcharias acutissima
Carcharias cf. reticulata Probst 1879 sand shark
"Odontaspis vorax" = Carcharias acutissima Agassiz 1843 sand shark |