PBDB Collection

Florissant, Station 14 (Eocene of the United States)

Where: Teller County, Colorado (38.9° N, 105.3° W: paleocoordinates 40.4° N, 97.0° W)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Florissant Formation, Chadronian (37.2 - 33.9 Ma)

Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous shale

• "shales"

Size class: mesofossils

Preservation: adpression

Collected by W. P. Cockerell, G. N. Rohwer; reposited in the AMNH, UCM, YPM

Primary reference: T. D. A. Cockerell. 1906. Fossil Saw-flies from Florissant Colorado. Bulletin of the American Museum of Natural History 22:499-501 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 106381: authorized by John Alroy, entered by John Alroy on 13.03.2011, edited by Amy Delelli, Jered Karr and Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Blattodea - Kalotermitidae	"Parotermes fodinae" = Proelectrotermes fodinae³⁷ "Parotermes fodinae" = Proelectrotermes fodinae³⁷ Scudder 1883 termite
Mantodea - Chaeteessidae	Lithophotina floccosa n. gen. n. sp.²⁰ Lithophotina floccosa n. gen. n. sp.²⁰ Cockerell 1908 mantis
Embioptera - Embiidae	"Embia florissantensis n. sp." = Lithembia florissantensis¹⁷ "Embia florissantensis n. sp." = Lithembia florissantensis¹⁷ Cockerell 1908 webspinners
Orthoptera - Tettigoniidae	Anabrus caudelli n. sp.²¹ Anabrus caudelli n. sp.²¹ Cockerell 1908 mormon cricket G. N. Rohwer, 1907
Orthoptera - Gryllidae	Lithogryllites lutzii n. sp.²¹ Lithogryllites lutzii n. sp.²¹ Cockerell 1908 true crickets S. A. Rohwer, 1907
Orthoptera - Eumastacidae	"Eobanksia bittaciformis" = Taphacris bittaciformis²⁷ "Eobanksia bittaciformis" = Taphacris bittaciformis²⁷ Cockerell 1909 monkey grasshopper Yale cotype (YPM 34983)
Dermaptera -	"Labiduromma gurneyi" = Petrolabis gurneyi² "Labiduromma gurneyi" = Petrolabis gurneyi² Brown 1984 earwig AMNH 18912
Coleoptera - Carabidae	Harpalus maceratus n. sp.⁵⁹ Harpalus maceratus n. sp.⁵⁹ Wickham 1911 ground beetle UCM
Coleoptera - Dytiscidae	Acilius florissantensis n. sp.⁵⁷, Miodytiscus hirtipes n. gen. n. sp.⁵⁹, Agabus charon n. sp.⁶⁰, "Coelambus miocenus n. sp." = Hygrotus miocenus⁶⁰ Acilius florissantensis n. sp.⁵⁷ Wickham 1909 predaceous diving beetle Miodytiscus hirtipes n. gen. n. sp.⁵⁹ Wickham 1911 predaceous diving beetle AMNH, coll. 1907 Agabus charon n. sp.⁶⁰ Wickham 1912 predaceous diving beetle W.P. Cockerell or S.A. Rohwer "Coelambus miocenus n. sp." = Hygrotus miocenus⁶⁰ Wickham 1912 predaceous diving beetle coll G.N. Rohwer
Coleoptera - Coccinellidae	Adalia subversa⁶⁰ Adalia subversa⁶⁰ Scudder 1900 ladybird beetle
Coleoptera - Lymexylidae	"Lymexylon lacustrinum n. sp." = Melittomma lacustrinum⁵⁹ "Lymexylon lacustrinum n. sp." = Melittomma lacustrinum⁵⁹ Wickham 1911 ship-timber beetle UCM
Coleoptera - Anthicidae	Macratria gigantea n. sp.⁵⁸ Macratria gigantea n. sp.⁵⁸ Wickham 1910 ant like flower beetle
Coleoptera - Meloidae	Nemognatha exsecta n. sp.⁶⁰ Nemognatha exsecta n. sp.⁶⁰ Wickham 1912 blister beetle W.P. Cockerell
Coleoptera - Kateretidae	Amartus petrefactus n. sp.⁶⁰ Amartus petrefactus n. sp.⁶⁰ Wickham 1912 beetle
Coleoptera - Cerambycidae	Leptura petrorum n. sp.⁶⁰, Phymatodes volans n. sp.²¹, Dryobius miocenicus n. sp.²¹, Elaphidion fracticorne n. sp.⁵⁹ Leptura petrorum n. sp.⁶⁰ Wickham 1912 longhorned beetle W.P. Cockerell Phymatodes volans n. sp.²¹ Beutenmüller and Cockerell 1908 longhorned beetle S. A. Rohwer, 1907 Dryobius miocenicus n. sp.²¹ Beutenmüller and Cockerell 1908 longhorned beetle W. P. Cockerell, 1907 Elaphidion fracticorne n. sp.⁵⁹ Wickham 1911 longhorned beetle UCM, Florissant 1906 no. 217
Coleoptera - Chrysomelidae	Bruchus henshawi n. sp.⁶⁰, "Bruchus haywardi n. sp." = Oligobruchus haywardi⁶⁰, "Bruchus scudderi n. sp." = Oligobruchus scudderi⁶⁰, "Donacia primaeva n. sp." = Plateumaris primaeva⁶⁰, Colaspis aetatis n. sp.⁵⁹ Bruchus henshawi n. sp.⁶⁰ Wickham 1912 leaf beetle "Bruchus haywardi n. sp." = Oligobruchus haywardi⁶⁰ Wickham 1912 palm bruchid Geo. N. Rohwer "Bruchus scudderi n. sp." = Oligobruchus scudderi⁶⁰ Wickham 1912 palm bruchid S.A. Rohwer "Donacia primaeva n. sp." = Plateumaris primaeva⁶⁰ Wickham 1912 reed leaf beetle S.A. Rohwer Colaspis aetatis n. sp.⁵⁹ Wickham 1911 leaf beetle UCM, Florissant 1906 no. 61
Coleoptera - Attelabidae	Eugnamptidea tertiaria n. gen. n. sp.⁶¹, Docirhynchus terebrans⁶¹, "Docirhynchus culex" = Pseudomesauletes (Pseudomesauletes) culex⁶¹, Isothea alleni⁶¹, Masteutes saxifer⁶¹, "Apion exanimale" = Prodeporaus exanimale⁶¹ Eugnamptidea tertiaria n. gen. n. sp.⁶¹ Wickham 1912 tooth-nosed snout weevil Docirhynchus terebrans⁶¹ Scudder 1893 tooth-nosed snout weevil "Docirhynchus culex" = Pseudomesauletes (Pseudomesauletes) culex⁶¹ Scudder 1893 tooth-nosed snout weevil Isothea alleni⁶¹ Scudder 1893 tooth-nosed snout weevil Masteutes saxifer⁶¹ Scudder 1893 tooth-nosed snout weevil "Apion exanimale" = Prodeporaus exanimale⁶¹ Scudder 1893 tooth-nosed snout weevil
Coleoptera - Curculionidae	"Coniatus differens n. sp." = Listroderes differens⁶¹, Cleonus foersteri⁶¹, Scyphophorus tertiarius n. sp.⁵⁹, Scyphophorus fossionis⁶¹, Otiorhynchites florissantensis n. sp.⁵⁹, Geralophus pumiceus⁶¹, Geralophus repositus⁶¹, Ophryastes championi⁶¹, Geralophus saxuosus⁶¹, Geralophus antiquarius⁵⁹, Geralophus fossicius⁶¹, Geralophus lassatus⁵⁹, Geralophus occultus⁵⁹, Geralophus scudderi n. sp.⁵⁹, Paussopsis secunda n. sp.⁶⁰, "Balaninus restrictus" = Curculio restrictus⁵⁹, "Balaninus minusculus" = Curculio minusculus⁶¹, "Balaninus minusculoides n. sp." = Curculio minusculoides⁵⁹, "Sibynes whitneyi" = Sibinia whitneyi⁶¹, Conotrachelus florissantensis n. sp.⁶¹, "Baris hoveyi n. sp." = Geraeus hoveyi⁶¹, Ophryastites miocenus n. sp.⁶¹, Ophryastites cinereus⁶¹, Cleonus estriatus n. sp.⁶¹, Cleonus primoris⁶¹, Cleonus rohweri n. sp.⁵⁹, Cleonus degeneratus⁶¹, Cryptorhynchus fallii n. sp.⁶¹, Cryptorhynchus coloradensis n. sp.⁶¹, Cryptorhynchus kerri⁶¹ "Coniatus differens n. sp." = Listroderes differens⁶¹ Wickham 1912 snout beetle Cleonus foersteri⁶¹ Theobald 1937 snout beetle Scyphophorus tertiarius n. sp.⁵⁹ Wickham 1911 snout beetle AMNH, Florissant 1906 no. 110(159) Scyphophorus fossionis⁶¹ Scudder 1893 snout beetle Otiorhynchites florissantensis n. sp.⁵⁹ Wickham 1911 snout beetle UCM, Florissant 1906 no. 213(215) Geralophus pumiceus⁶¹ Scudder 1893 snout beetle Geralophus repositus⁶¹ Scudder 1893 snout beetle Ophryastes championi⁶¹ Wickham 1912 snout beetle Geralophus saxuosus⁶¹ Scudder 1893 snout beetle Geralophus antiquarius⁵⁹ Scudder 1893 snout beetle no. 85, 104, 177, 180, 251 Geralophus fossicius⁶¹ Scudder 1893 snout beetle Geralophus lassatus⁵⁹ Scudder 1893 snout beetle no. 6 Geralophus occultus⁵⁹ Scudder 1876 snout beetle no. 43 (Wickham 1912 mentions four specimens) Geralophus scudderi n. sp.⁵⁹ Wickham 1911 snout beetle Florissant 1906, no. 11 Paussopsis secunda n. sp.⁶⁰ Wickham 1912 snout beetle Geo. N. Rohwer "Balaninus restrictus" = Curculio restrictus⁵⁹ Scudder 1893 snout beetle Florissant 1906 no. 74 (Wickham 1912 mentions a second specimen) "Balaninus minusculus" = Curculio minusculus⁶¹ Scudder 1893 snout beetle "Balaninus minusculoides n. sp." = Curculio minusculoides⁵⁹ Wickham 1911 snout beetle UCM, Florissant 1906 no. 34 "Sibynes whitneyi" = Sibinia whitneyi⁶¹ Scudder 1893 snout beetle Conotrachelus florissantensis n. sp.⁶¹ Wickham 1912 snout beetle "Baris hoveyi n. sp." = Geraeus hoveyi⁶¹ Wickham 1912 snout beetle AMNH Ophryastites miocenus n. sp.⁶¹ Wickham 1912 snout beetle Ophryastites cinereus⁶¹ Scudder 1893 snout beetle Cleonus estriatus n. sp.⁶¹ Wickham 1912 snout beetle AMNH Cleonus primoris⁶¹ Scudder 1893 snout beetle Cleonus rohweri n. sp.⁵⁹ Wickham 1911 snout beetle UCM, Florissant 1906 no. 71(88) Cleonus degeneratus⁶¹ Scudder 1893 snout beetle Cryptorhynchus fallii n. sp.⁶¹ Wickham 1912 snout beetle Cryptorhynchus coloradensis n. sp.⁶¹ Wickham 1912 snout beetle Cryptorhynchus kerri⁶¹ Scudder 1893 snout beetle
Coleoptera - Brentidae	"Apion refrenatum" = Toxorhynchus refrenatum⁶¹ "Apion refrenatum" = Toxorhynchus refrenatum⁶¹ Scudder 1893 weevil
Coleoptera - Erotylidae	"Tritoma materna n. sp." = Triplax materna⁶⁰, "Tritoma submersa n. sp." = Triplax submersa⁶⁰ "Tritoma materna n. sp." = Triplax materna⁶⁰ Wickham 1912 pleasing fungus beetle S.A. Rohwer "Tritoma submersa n. sp." = Triplax submersa⁶⁰ Wickham 1912 pleasing fungus beetle
Coleoptera - Melyridae	Eudasytites listriformis n. gen. n. sp.⁶⁰ Eudasytites listriformis n. gen. n. sp.⁶⁰ Wickham 1912 soft-winged flower beetle S.A. Rohwer
Coleoptera - Scarabaeidae	Ataenius patescens⁵⁸, Aphodius laminicola n. sp.⁵⁸, Macrodactylus propheticus n. sp.⁶⁰, Serica antediluviana n. sp.⁶⁰ Ataenius patescens⁵⁸ Scudder 1900 scarab beetle Aphodius laminicola n. sp.⁵⁸ Wickham 1910 scarab beetle Type in YPM, other in UCM; additional specimen in Wickham, 1912 Macrodactylus propheticus n. sp.⁶⁰ Wickham 1912 scarab beetle Serica antediluviana n. sp.⁶⁰ Wickham 1912 scarab beetle W.P. Cockerell
Coleoptera - Trogidae	Trox antiquus n. sp.⁵⁷ Trox antiquus n. sp.⁵⁷ Wickham 1909 hide beetle BMNH
Coleoptera - Glaphyridae	"Amphicoma defuncta n. sp." = Lichnanthe defuncta⁵⁸ "Amphicoma defuncta n. sp." = Lichnanthe defuncta⁵⁸ Wickham 1910 bumble bee scarab beetle
Coleoptera - Hydrophilidae	Hydrobius prisconatator n. sp.⁵⁹ Hydrobius prisconatator n. sp.⁵⁹ Wickham 1911 water scavenger beetle UCM, bearing numbers 175(185), coll. 1906
Coleoptera - Staphylinidae	Quedius mortuus n. sp.⁶⁰, Bledius osborni⁶⁰, Deleaster grandiceps n. sp.⁶⁰, Miosilpha necrophiloides n. gen. n. sp.⁶⁰, Tachinus sommatus⁶⁰, Mycetoporus demersus⁶⁰, Necrodes primaevus n. sp.²¹ Quedius mortuus n. sp.⁶⁰ Wickham 1912 rove beetle S.A. Rohwer Bledius osborni⁶⁰ Scudder 1900 rove beetle W.P. Cockerell Deleaster grandiceps n. sp.⁶⁰ Wickham 1912 rove beetle Geo. N. Rohwer Miosilpha necrophiloides n. gen. n. sp.⁶⁰ Wickham 1912 rove beetle Coll. W.P. Cockerell Tachinus sommatus⁶⁰ Scudder 1900 rove beetle Mycetoporus demersus⁶⁰ Scudder 1900 rove beetle W.P. Cockerell Necrodes primaevus n. sp.²¹ Beutenmüller and Cockerell 1908 rove beetle S. A. Rohwer 1907
Coleoptera - Byrrhidae	Nosotetocus vespertinus⁶⁰ Nosotetocus vespertinus⁶⁰ Scudder 1900 pill beetle S.A. Rohwer
Coleoptera - Lutrochidae	Lutrochites lecontei n. gen. n. sp.⁶⁰ Lutrochites lecontei n. gen. n. sp.⁶⁰ Wickham 1912 robust marsh loving beetle W.P. Cockerell
Coleoptera - Dryopidae	"Dryops tenuior n. sp." = Helichus tenuior⁶⁰ "Dryops tenuior n. sp." = Helichus tenuior⁶⁰ Wickham 1912 long toed water beetle W.P. Cockerell
Coleoptera - Buprestidae	Brachyspathus curiosus n. gen. n. sp.⁶³, Anthaxia exhumata⁶⁰, Melanophila cockerellae n. sp.⁶⁰, "Acmaeodera schaefferi n. sp." = Ptosima schaefferi⁶⁰ Brachyspathus curiosus n. gen. n. sp.⁶³ Wickham 1917 jewel beetle UCM 184 collected by Cockerell 1906 Anthaxia exhumata⁶⁰ Wickham 1913 jewel beetle Geo. N. Rohwer Melanophila cockerellae n. sp.⁶⁰ Wickham 1912 jewel beetle W.P. Cockerell "Acmaeodera schaefferi n. sp." = Ptosima schaefferi⁶⁰ Wickham 1912 jewel beetle W.P. Cockerell
Coleoptera - Elateridae	Cardiophorus lithographicus⁶², Cardiophorus cockerelli⁶², "Corymbites granulicollis n. sp." = Ctenicera granulicollis⁵⁶, "Corymbites submersus n. sp." = Ctenicera submersa⁶², "Corymbites restructus n. sp." = Ctenicera restructa⁶², "Paranomus laevissimus n. sp." = Eanus laevissimus⁶², "Paranomus heeri n. sp." = Eanus heeri⁶², Athous fractus n. sp.⁶², Limonius florissantensis⁶², Limonius volans n. sp.⁶², Limonius shoshonis n. sp.⁶², Melanactes cockerelli n. sp.⁵⁶, Elater rohweri n. sp.⁶² Cardiophorus lithographicus⁶² Wickham 1916 click beetle UCM Cardiophorus cockerelli⁶² Wickham 1916 click beetle UCM, bearing numbers 151, 178(179), 209(258) "Corymbites granulicollis n. sp." = Ctenicera granulicollis⁵⁶ Wickham 1908 click beetle YPM 1, coll S.A. Rohwer "Corymbites submersus n. sp." = Ctenicera submersa⁶² Wickham 1916 click beetle UCM "Corymbites restructus n. sp." = Ctenicera restructa⁶² Wickham 1916 click beetle UCM "Paranomus laevissimus n. sp." = Eanus laevissimus⁶² Wickham 1916 click beetle UCM "Paranomus heeri n. sp." = Eanus heeri⁶² Wickham 1916 click beetle UCM Athous fractus n. sp.⁶² Wickham 1916 click beetle UCM Limonius florissantensis⁶² Wickham 1916 click beetle UCM Limonius volans n. sp.⁶² Wickham 1916 click beetle Limonius shoshonis n. sp.⁶² Wickham 1916 click beetle UCM, Cockerell 58 Melanactes cockerelli n. sp.⁵⁶ Wickham 1908 click beetle YPM 3, coll. W.P. Cockerell Elater rohweri n. sp.⁶² Wickham 1916 click beetle UCM, Cockerell 192(211)
Coleoptera - Eucnemidae	Microrhagus miocenicus n. sp.⁶², "Deltometopus fossilis n. sp." = Microrhagus fossilis⁶² Microrhagus miocenicus n. sp.⁶² Wickham 1916 false click beetle UCM "Deltometopus fossilis n. sp." = Microrhagus fossilis⁶² Wickham 1916 false click beetle UCM
Coleoptera - Lampyridae	"Pyropyga prima n. sp." = Lucidota prima⁶¹ "Pyropyga prima n. sp." = Lucidota prima⁶¹ Wickham 1912 beetle UCM-28047
Coleoptera - Dermestidae	Dermestes tertiarius n. sp.⁶⁰, Orphilus dubius⁶⁰ Dermestes tertiarius n. sp.⁶⁰ Wickham 1912 skin beetle W.P. Cockerell Orphilus dubius⁶⁰ Wickham 1912 skin beetle W.P. Cockerell
Coleoptera - Bostrichidae	Xylobiops lacustre n. sp.⁶⁰ Xylobiops lacustre n. sp.⁶⁰ Wickham 1912 false powderpost beetle S.A. Rohwer
Raphidioptera - Raphidiidae	"Raphidia mortua n. sp." = Florissantoraphidia mortua⁵⁵ "Raphidia mortua n. sp." = Florissantoraphidia mortua⁵⁵ Rohwer 1909 snakefly UCM-4512
Neuroptera - Osmylidae	Pseudosmylidia relicta n. gen. n. sp.⁴⁷ Pseudosmylidia relicta n. gen. n. sp.⁴⁷ Makarkin 2017 lance lacewing UCM 85899
Neuroptera - Ithonidae	"Polystoechotes piperatus n. sp." = Polystoechotites piperatus²¹ "Polystoechotes piperatus n. sp." = Polystoechotites piperatus²¹ Cockerell 1908 moth lacewing AMNH 34871 G. N. Rohwer 1907
Neuroptera - Chrysopidae	Archaeochrysa paranervis n. sp.¹, "Palaeochrysa concinnula n. sp." = Lithochrysa concinnula³⁰, Cimbrochrysa americana n. sp.⁴⁸ Archaeochrysa paranervis n. sp.¹ Adams 1967 green lacewing UCM 4419 "Palaeochrysa concinnula n. sp." = Lithochrysa concinnula³⁰ Cockerell 1909 green lacewing S.A. Rohwer Cimbrochrysa americana n. sp.⁴⁸ Makarkin et al. 2022 green lacewing UCM 33497
Trichoptera - Limnephilidae	"Platyphylax (Eopteryx) florissantensis n. sp." = Tricheopteryx florissantensis¹³ "Platyphylax (Eopteryx) florissantensis n. sp." = Tricheopteryx florissantensis¹³ Cockerell 1907 caddisfly "station in doubt, may have come from station 13"
Trichoptera - Polycentropodidae	Derobrochus typharum n. sp.³¹ Derobrochus typharum n. sp.³¹ Cockerell 1910 caddisfly
Lepidoptera -	"Tortrix florissantana n. sp." = Paleolepidopterites florissantanus¹² "Tortrix florissantana n. sp." = Paleolepidopterites florissantanus¹² Cockerell 1907 moth
Lepidoptera - Nymphalidae	Nymphalites scudderi n. sp.²¹ Nymphalites scudderi n. sp.²¹ Beutenmüller and Cockerell 1908 brush-footed butterfly S. A. Rohwer
Lepidoptera - Psychidae	Adelopsyche frustrans n. gen. n. sp.⁴³ Adelopsyche frustrans n. gen. n. sp.⁴³ Cockerell 1926 bagworm moth
Lepidoptera -	Phylledestes vorax n. gen. n. sp.⁸ Phylledestes vorax n. gen. n. sp.⁸ Cockerell 1907 moth UCM 4608
Mecoptera - Panorpidae	Panorpa arctiiformis n. sp.⁹ Panorpa arctiiformis n. sp.⁹ Cockerell 1907 common scorpionfly W. P. Cockerell
Mecoptera - Eomeropidae	Eomerope tortriciformis n. gen. n. sp.²⁷ Eomerope tortriciformis n. gen. n. sp.²⁷ Cockerell 1909 scorpionfly Doubtless from station 14, but the specimens is not marked
Mecoptera - Holcorpidae	Holcorpa maculosa⁷ Holcorpa maculosa⁷ Scudder 1878 scorpionfly UCM 4494
Hymenoptera - Ichneumonidae	Acoenites defunctus n. sp.⁴, Orthocentrus primus n. sp.⁴, Rhyssa petiolata n. sp.⁴, Tryphon explanatum n. sp.⁴¹ Acoenites defunctus n. sp.⁴ Brues 1906 ichneumon wasp Orthocentrus primus n. sp.⁴ Brues 1906 ichneumon wasp Rhyssa petiolata n. sp.⁴ Brues 1906 ichneumon wasp Tryphon explanatum n. sp.⁴¹ Cockerell 1919 ichneumon wasp UCM 8607
Hymenoptera - Braconidae	Microgaster primordialis n. sp.⁴, Bracon resurrectus n. sp.⁶, Bracon cockerelli n. sp.⁶ Microgaster primordialis n. sp.⁴ Brues 1906 wasp Bracon resurrectus n. sp.⁶ Brues 1910 wasp A137/8 Bracon cockerelli n. sp.⁶ Brues 1910 wasp AMNH A72, A31, A35, A128/129, A41 (some from station 17)
Hymenoptera - Crabronidae	Mellinus handlirschi n. sp.⁵¹, Crabro longaevus n. sp.³², Pison cockerellae n. sp.⁵³ Mellinus handlirschi n. sp.⁵¹ Rohwer 1908 apoid wasp Crabro longaevus n. sp.³² Cockerell 1910 moustache wasp Pison cockerellae n. sp.⁵³ Rohwer 1908 apoid wasp
Hymenoptera - Megachilidae	Heriades saxosus n. sp.³⁶, Lithanthidium pertriste n. gen. n. sp.³³, Megachile praedicta n. sp.¹⁸ Heriades saxosus n. sp.³⁶ Cockerell 1913 mason bee UCM 18638 Lithanthidium pertriste n. gen. n. sp.³³ Cockerell 1911 mason bee UCM 18738 Megachile praedicta n. sp.¹⁸ Cockerell 1908 mason bee UCM 4542
Hymenoptera - Halictidae	"Cyrtapis anomalus n. gen. n. sp." = Cyrtapis anomala²⁴ "Cyrtapis anomalus n. gen. n. sp." = Cyrtapis anomala²⁴ Cockerell 1908 sweat bee UCM 19217 (W.P. Cockerell, 1907)
Hymenoptera - Apidae	Protomelecta brevipennis n. gen. n. sp.²⁴ Protomelecta brevipennis n. gen. n. sp.²⁴ Cockerell 1908 bee UCM 18639 (W.P. Cockerell, 1907)
Hymenoptera - Sphecidae	Chalybion mortuum n. sp.¹³ Chalybion mortuum n. sp.¹³ Cockerell 1907 digger wasp
Hymenoptera - Chrysididae	Chrysis rohweri n. sp.¹³, Chrysis miocenica n. sp.⁵⁵ Chrysis rohweri n. sp.¹³ Cockerell 1907 cuckoo wasp Chrysis miocenica n. sp.⁵⁵ Rohwer 1909 cuckoo wasp UCM-4523
Hymenoptera - Tiphiidae	"Paratiphia praefracta n. sp." = Philoponites praefractus¹¹ "Paratiphia praefracta n. sp." = Philoponites praefractus¹¹ Cockerell 1907 wasp
Hymenoptera - Vespidae	Odynerus wilmattae n. sp.³⁹ Odynerus wilmattae n. sp.³⁹ Cockerell 1915 potter wasp Coll. W.P. Cockerell
Hymenoptera - Pompilidae	"Salius laminarum n. sp." = Pepsinites laminarum⁵⁴, "Salius senex n. sp." = Pompilites senex⁵⁴, "Agenia saxigena n. sp." = Agenioideus saxigenus¹⁷ "Salius laminarum n. sp." = Pepsinites laminarum⁵⁴ Rohwer 1909 spider wasp UCM 8597 "Salius senex n. sp." = Pompilites senex⁵⁴ Rohwer 1909 spider wasp UCM 8594 "Agenia saxigena n. sp." = Agenioideus saxigenus¹⁷ Cockerell 1908 spider wasp
Hymenoptera - Aulacidae	Pristaulacus rohweri n. sp.⁶, "Aulacites secundus n. gen. n. sp." = Pristaulacus secundus⁴⁰ Pristaulacus rohweri n. sp.⁶ Brues 1910 wasp no # "Aulacites secundus n. gen. n. sp." = Pristaulacus secundus⁴⁰ Cockerell 1916 wasp USNM 61458 (Univ. Colorado expedition)
Hymenoptera -	Pantoclis deperdita n. sp.⁴, Pteronus prodigus⁵² Pantoclis deperdita n. sp.⁴ Brues 1906 wasp Pteronus prodigus⁵² Brues 1908 wood wasp W.P. Cockerell, 1907
Hymenoptera - Tenthredinidae	Tenthredo submersa n. sp.¹³, Fenusa primula n. sp.⁵², "Lithoryssus parvus n. gen. n. sp." = Fenusa parva⁴, Dineura laminarum⁵², "Hemichroa eophila n. sp." = Eohemichroa eophila, Selandria sapindi n. sp.³¹ Tenthredo submersa n. sp.¹³ Cockerell 1907 wood wasp Fenusa primula n. sp.⁵² Rohwer 1908 wood wasp S.A. Rohwer, 1907 "Lithoryssus parvus n. gen. n. sp." = Fenusa parva⁴ Brues 1906 wood wasp Dineura laminarum⁵² Brues 1908 wood wasp W.P. Cockerell, 1907 "Hemichroa eophila n. sp." = Eohemichroa eophila Cockerell 1906 wood wasp S. A. Rohwer July, 1906 Selandria sapindi n. sp.³¹ Cockerell 1910 wood wasp
Hymenoptera - Cimbicidae	Trichiosomites obliviosus⁵², "Perga coloradensis n. sp." = Phenacoperga coloradensis⁹ Trichiosomites obliviosus⁵² Brues 1908 cimbicid sawfly W.P. Cockerell, 1907 "Perga coloradensis n. sp." = Phenacoperga coloradensis⁹ Cockerell 1907 cimbicid sawfly W. P. Cockerell
Hymenoptera - Athaliidae	"Eriocampa wheeleri n. sp." = Athalia wheeleri "Eriocampa wheeleri n. sp." = Athalia wheeleri Cockerell 1906 wood wasp W. M. Wheeler July 1906 (additional specimen by Geo. N. Rohwer, 1907)
Hemiptera - Cicadidae	Platypedia primigenia n. sp.¹⁴ Platypedia primigenia n. sp.¹⁴ Cockerell 1908 cicada YPM (W.P. Cockerell, 1907)
Hemiptera - Cercopidae	Palecphora praevalens¹⁵, Palecphora communis¹⁵ Palecphora praevalens¹⁵ Scudder 1890 froghopper Palecphora communis¹⁵ Scudder 1890 froghopper
Hemiptera - Aphrophoridae	Palaphrodes irregularis¹⁵ Palaphrodes irregularis¹⁵ Scudder 1890 froghopper
Hemiptera - Nepidae	"Zaitha vulcanica n. sp." = Nepa vulcanica¹⁷ "Zaitha vulcanica n. sp." = Nepa vulcanica¹⁷ Cockerell 1908 true bug
Hemiptera - Pentatomidae	Teleoschistus rigoratus²⁹, Poliocoris amnesis n. gen. n. sp.⁴⁶ Teleoschistus rigoratus²⁹ Scudder 1890 stink bug S. A. Rohwer, 1907 Poliocoris amnesis n. gen. n. sp.⁴⁶ Kirkaldy 1910 stink bug
Ephemeroptera - Ephemeridae	Ephemera howarthi n. sp.¹⁷ Ephemera howarthi n. sp.¹⁷ Cockerell 1908 burrowing mayfly
Odonata - Gomphaeschnidae	"Lithaeschna needhami n. gen. n. sp." = Oligoaeschna needhami¹⁰ "Lithaeschna needhami n. gen. n. sp." = Oligoaeschna needhami¹⁰ Cockerell 1907 true dragonfly W. P. Cockerell July 7, 1906
Odonata - Aeshnidae	"Aeschna solida" = Aeshna solida¹⁶, "Oplonaeschna separata" = Oligaeschna separata¹⁹ "Aeschna solida" = Aeshna solida¹⁶ Scudder 1890 hawker dragonfly "Oplonaeschna separata" = Oligaeschna separata¹⁹ Scudder 1890 darner dragonfly
Odonata - Calopterygidae	Agrion telluris²¹ Agrion telluris²¹ Scudder 1890 broad-winged damselfly G. N. Rohwer, 1907
Odonata - Dysagrionidae	Phenacolestes mirandus n. gen. n. sp.²¹, Phenacolestes parallelus n. sp.²¹ Phenacolestes mirandus n. gen. n. sp.²¹ Cockerell 1908 damselfly S. A. Rohwer Phenacolestes parallelus n. sp.²¹ Cockerell 1908 damselfly W. P. Cockerell
Odonata - Coenagrionidae	"Hesperagrion praevolans n. sp." = Amphiagrion praevolans¹⁰, Enallagma oblisum n. sp.⁴², Enallagma florissantella n. sp.¹⁹ "Hesperagrion praevolans n. sp." = Amphiagrion praevolans¹⁰ Cockerell 1907 narrow-winged damselfly AMNH FI 39556 Enallagma oblisum n. sp.⁴² Cockerell 1925 bluet Enallagma florissantella n. sp.¹⁹ Cockerell 1908 bluet YPM-26172
Odonata - Argiolestidae	Melanagrion nigerrimum n. sp.¹⁹, Lithagrion hyalinum¹⁰ Melanagrion nigerrimum n. sp.¹⁹ Cockerell 1908 damselfly W. P. Cockerell Holotype in YPM-26168 Lithagrion hyalinum¹⁰ Scudder 1890 damselfly AMNH FI 39479
Diptera - Xylophagidae	Dialysis revelata n. sp.²⁵ Dialysis revelata n. sp.²⁵ Cockerell 1908 fly W.P. Cockerell, 1907; and AMNH 26495 (coll Wheeler) in Melander 1949
Diptera - Rhagionidae	Symphoromyia subtrita n. sp.³⁴, Rhagio wheeleri n. sp.⁵⁰ Symphoromyia subtrita n. sp.³⁴ Cockerell 1911 snipe fly AMNH-FI-18867 Rhagio wheeleri n. sp.⁵⁰ Melander 1949 snipe fly AMNH 26496:1, 3, 4, 4A (coll Wheeler)
Diptera - Stratiomyidae	Moyamyia limigena n. gen. n. sp.⁵⁰, "Oxycera rohweri n. sp." = Cyphomyia rohweri⁴⁰ Moyamyia limigena n. gen. n. sp.⁵⁰ Melander 1949 soldier fly AMNH 26494 "Oxycera rohweri n. sp." = Cyphomyia rohweri⁴⁰ Cockerell 1916 soldier fly (G.N. Rohwer)
Diptera - Nemestrinidae	"Hirmoneura melanderi n. sp." = Neorhynchocephalus melanderi²³, "Hirmoneura vulcanica n. sp." = Neorhynchocephalus vulcanicus²³, "Hirmoneura occultator n. sp." = Neorhynchocephalus occultator²² "Hirmoneura melanderi n. sp." = Neorhynchocephalus melanderi²³ Cockerell 1908 tangle-veined fly YPM 14172 (W.P. Cockerell, 1907) "Hirmoneura vulcanica n. sp." = Neorhynchocephalus vulcanicus²³ Cockerell 1908 tangle-veined fly YPM 14173 (collected by Mrs Charlotte Hill "Hirmoneura occultator n. sp." = Neorhynchocephalus occultator²² Cockerell 1908 tangle-veined fly UCM 18631
Diptera - Empididae	Acallomyia probolaea n. sp.⁵⁰, Rhamphomyia interita n. sp.⁵⁰, Rhamphomyia aeterna n. sp.⁵⁰, Rhamphomyia tumulata n. sp.⁵⁰, Rhamphomyia spodites n. sp.⁵⁰ Acallomyia probolaea n. sp.⁵⁰ Melander 1949 dagger fly AMNH 26603 Rhamphomyia interita n. sp.⁵⁰ Melander 1949 dagger fly AMNH 26605 (coll Wheeler) Rhamphomyia aeterna n. sp.⁵⁰ Melander 1949 dagger fly AMNH 26604 Rhamphomyia tumulata n. sp.⁵⁰ Melander 1949 dagger fly AMNH 26608 Rhamphomyia spodites n. sp.⁵⁰ Melander 1949 dagger fly AMNH 26607
Diptera - Syrphidae	Syrphus willistoni n. sp.²⁶, Platycheirus lethaeus⁵⁰, "Chilosia miocenica" = Cheilosia miocenica²⁹ Syrphus willistoni n. sp.²⁶ Cockerell 1909 hover fly in AMNH Platycheirus lethaeus⁵⁰ Melander 1949 hover fly AMNH 26610 (S.A. Rohwer) "Chilosia miocenica" = Cheilosia miocenica²⁹ Cockerell 1909 hoverfly S. A. Rohwer, 1907
Diptera - Glossinidae	"Glossina oligocena" = Glossina oligocenus⁹ "Glossina oligocena" = Glossina oligocenus⁹ Scudder 1892 tsetse fly G. N. Rohwer 1907
Diptera - Agromyzidae	Agromyza praecursor n. sp.⁵⁰ Agromyza praecursor n. sp.⁵⁰ Melander 1949 leaf-miner fly AMNH 26619 (coll Wheeler)
Diptera - Ulidiidae	Melieria calligrapha n. sp.⁵⁰ Melieria calligrapha n. sp.⁵⁰ Melander 1949 picture-winged fly AMNH 26615 (coll Wheeler)
Diptera - Phoridae	Phora tumbae n. sp.⁵⁰, "Phora laminarum n. sp." = Chaetopleurophora laminarum⁵ Phora tumbae n. sp.⁵⁰ Melander 1949 scuttle fly AMNH 26612 "Phora laminarum n. sp." = Chaetopleurophora laminarum⁵ Brues 1908 scuttle fly W. P. Cockerell, and AMNH 26611 in Melander 1949
Diptera - Bombyliidae	"Lithocosmus coquilletti" = Paracosmus (Paracosmus) coquilletti³⁴, "Geronites stigmalis n. gen. n. sp." = Poecilognathus stigmalis³⁸, Melanderella glossalis³⁴, Apolysis magister n. sp.⁴⁹, "Megacosmus mirandus n. gen. n. sp." = Comptosia miranda²⁶, Alepidophora cockerelli n. sp.⁵⁰ "Lithocosmus coquilletti" = Paracosmus (Paracosmus) coquilletti³⁴ Cockerell 1909 bee fly found by SA Rohwer "Geronites stigmalis n. gen. n. sp." = Poecilognathus stigmalis³⁸ Cockerell 1914 bee fly UCM coll Melanderella glossalis³⁴ Cockerell 1909 bee fly found by SA Rohwer Apolysis magister n. sp.⁴⁹ Melander 1946 bee fly "Megacosmus mirandus n. gen. n. sp." = Comptosia miranda²⁶ Cockerell 1909 bee fly in AMNH Alepidophora cockerelli n. sp.⁵⁰ Melander 1949 bee fly AMNH 26499
Diptera - Asilidae	Senoprosopis eureka n. sp.⁵⁰, Asilus peritulus³⁶, Asilus florissantinus n. sp.⁴⁵, Philonicus saxorum n. sp.⁴⁵, Microstylum wheeleri n. sp.²⁵, Taracticus contusus n. sp.³¹, "Psilocephala hypogaea" = Taracticus hypogaeus²⁹, Lestomyia miocenica n. sp.⁴⁵ Senoprosopis eureka n. sp.⁵⁰ Melander 1949 robber fly AMNH 26602 Asilus peritulus³⁶ Cockerell 1909 robber fly Geo. N. Rohwer Asilus florissantinus n. sp.⁴⁵ James 1939 robber fly W. P. Cockerell Philonicus saxorum n. sp.⁴⁵ James 1939 robber fly W. P. Cockerell Microstylum wheeleri n. sp.²⁵ Cockerell 1908 robber fly T.D.A. Cockerell, July 8, 1906 Taracticus contusus n. sp.³¹ Cockerell 1910 robber fly "Psilocephala hypogaea" = Taracticus hypogaeus²⁹ Cockerell 1909 robber fly G. N. Rohwer Lestomyia miocenica n. sp.⁴⁵ James 1939 robber fly W. P. Cockerell
Diptera - Therevidae	"Psilocephala scudderi n. sp." = Palaeopherocera scudderi²⁶ "Psilocephala scudderi n. sp." = Palaeopherocera scudderi²⁶ Cockerell 1909 stiletto fly 1907, in AMNH; USNM 61456 (G.N. Rohwer) in Cockerell, 1916
Diptera - Mydidae	Mydas miocenicus n. sp.³⁵ Mydas miocenicus n. sp.³⁵ Cockerell 1913 mydas fly
Diptera - Bibionidae	Bibio cockerelli⁵⁰, Bibio wickhami⁵⁰, Bibio vetus n. sp.⁴⁴ Bibio cockerelli⁵⁰ James 1937 march fly AMNH 26484/2 Bibio wickhami⁵⁰ Cockerell 1915 march fly AMNH 26490/3 (10 specimens from four stations, undivided) Bibio vetus n. sp.⁴⁴ James 1937 march fly W. P. Cockerell, and AMNH 26488 in Melander 1949
Diptera - Mycetophilidae	"Lasiosoma mirandula n. sp." = Sciophila mirandula²⁸ "Lasiosoma mirandula n. sp." = Sciophila mirandula²⁸ Cockerell 1909 fungus gnat in YPM
Diptera - Keroplatidae	Proapemon infernus n. gen. n. sp.⁵⁰ Proapemon infernus n. gen. n. sp.⁵⁰ Melander 1949 predatory fungus gnat AMNH 26463
Diptera - Sciaridae	Sciara requieta n. sp.⁵⁰ Sciara requieta n. sp.⁵⁰ Melander 1949 dark-winged fungus gnat AMNH 26465
Diptera - Cecidomyiidae	? Cecidomyia pontaniiformis²¹ ? Cecidomyia pontaniiformis²¹ Cockerell 1908 gall midge
Diptera - Tipulidae	Tipula wilmatteae n. sp.⁵⁰, Tipula florissanta⁵⁰, Tipula maclurei⁵⁰, Tipula lapillescens⁵⁰, "Tipula (Micrapsis) decorata n. sp." = Tipula fmartinbrowni³, Tipula heilprini⁵⁰ Tipula wilmatteae n. sp.⁵⁰ Melander 1949 crane fly AMNH 26452 Tipula florissanta⁵⁰ Scudder 1894 crane fly AMNH 26453/1 (coll Wheeler) Tipula maclurei⁵⁰ Scudder 1894 crane fly AMNH 26456:1, 2 (coll Brues) Tipula lapillescens⁵⁰ Scudder 1894 crane fly AMNH 26455:1, 2 "Tipula (Micrapsis) decorata n. sp." = Tipula fmartinbrowni³ Evenhuis 1994 crane fly UCM 29988(30067) Tipula heilprini⁵⁰ Scudder 1894 crane fly AMNH 26454
Diptera - Limoniidae	Limnocema marcescens⁵⁰ Limnocema marcescens⁵⁰ Scudder 1894 crane fly AMNH 26461
Diptera - Chironomidae	Chironomus requiescens⁵⁰, Chironomus pristinus n. sp.⁵⁰, Chironomus scudderiellus⁵⁰ Chironomus requiescens⁵⁰ Melander 1949 non biting midge AMNH 26474/2:1, 2:2 (G. Rohwer) Chironomus pristinus n. sp.⁵⁰ Melander 1949 non biting midge AMNH 26470:1, 2 Chironomus scudderiellus⁵⁰ Cockerell 1916 non biting midge AMNH 26472/2
Arachnida
Opiliones - Sclerosomatidae	"Leptobunus atavus n. sp." = Amauropilio atavus¹³ "Leptobunus atavus n. sp." = Amauropilio atavus¹³ Cockerell 1907 harvestman
Acariformes - Eriophyidae	? Eriophyes beutenmulleri²¹ ? Eriophyes beutenmulleri²¹ Cockerell 1908 gall mite
Myriapoda
- Iulidae	Iulus florissantellus n. sp.¹³ Iulus florissantellus n. sp.¹³ Cockerell 1907 millipede

Coniferales - Cupressaceae	"Heyderia coloradensis n. sp." = Libocedrus coloradensis⁹ "Heyderia coloradensis n. sp." = Libocedrus coloradensis⁹ Cockerell 1907 cypress from "Florissant"; placed here because it was collected in the same field season as three other species from Station 14 described in the same paper
Angiospermae
Fagales - Myricaceae	Myrica drymeja²¹ Myrica drymeja²¹ Lesquereux 1873 bayberry

¹Adams 1967; ²Brown 1984; ³Brown 1985; ⁴Brues 1906; ⁵Brues 1908; ⁶Brues 1910; ⁷Carpenter 1931; ⁸Cockerell 1907; ⁹Cockerell 1907; ¹⁰Cockerell 1907; ¹¹Cockerell 1907; ¹²Cockerell 1907; ¹³Cockerell 1907; ¹⁴Cockerell 1908; ¹⁵Cockerell 1908; ¹⁶Cockerell 1908; ¹⁷Cockerell 1908; ¹⁸Cockerell 1908; ¹⁹Cockerell 1908; ²⁰Cockerell 1908; ²¹Cockerell 1908; ²²Cockerell 1908; ²³Cockerell 1908; ²⁴Cockerell 1908; ²⁵Cockerell 1908; ²⁶Cockerell 1909; ²⁷Cockerell 1909; ²⁸Cockerell 1909; ²⁹Cockerell 1909; ³⁰Cockerell 1909; ³¹Cockerell 1910; ³²Cockerell 1910; ³³Cockerell 1911; ³⁴Cockerell 1911; ³⁵Cockerell 1913; ³⁶Cockerell 1913; ³⁷Cockerell 1913; ³⁸Cockerell 1914; ³⁹Cockerell 1915; ⁴⁰Cockerell 1916; ⁴¹Cockerell 1919; ⁴²Cockerell 1925; ⁴³Cockerell 1926; ⁴⁴James 1937; ⁴⁵James 1939; ⁴⁶Kirkaldy 1910; ⁴⁷Makarkin 2017; ⁴⁸Makarkin et al. 2022; ⁴⁹Melander 1946; ⁵⁰Melander 1949; ⁵¹Rohwer 1908; ⁵²Rohwer 1908; ⁵³Rohwer 1908; ⁵⁴Rohwer 1909; ⁵⁵Rohwer 1909; ⁵⁶Wickham 1908; ⁵⁷Wickham 1909; ⁵⁸Wickham 1910; ⁵⁹Wickham 1911; ⁶⁰Wickham 1912; ⁶¹Wickham 1912; ⁶²Wickham 1916; ⁶³Wickham 1917