Radoboj (Grätz Collection) (Miocene of Croatia)
Where: Croatia (46.2° N, 15.9° E: paleocoordinates 45.7° N, 15.6° E)
• coordinate based on nearby landmark
• outcrop-level geographic resolution
When: Sarmatian (12.7 - 11.6 Ma)
• Likely early Sarmatian (MN7) in age, based on foraminifera (Bajraktarevic, 1984) and siliceous microfossils (Galovic, 2003) from localities in the vicinity of Krapina.
Environment/lithology: lagoonal or restricted shallow subtidal; lithified limestone
• The deposits of Radoboj are marine deposits accumulated in a shallow epicontinental sea, possibly in a lagoon. In addition to insects, marine fish and algae have been found in Radoboj. The formation of the Pannonian Basin System commenced in the Early and the Middle Miocene as the consequence of the continental collision of the African (=Apulian) and European plates, and deposition in the entire basin was influenced by important extensional tectonics.
Size classes: macrofossils, mesofossils
Preservation: mold/impression, adpression
Collection methods: Repository: Sammlung zu Grätz, in Naturalienkabinet der Johanneums, sent to Heer by Unger. Includes most (?all) material described and illustrated by Charpentier, 1843.
Primary reference: F. Unger. 1841. Fossile Insecten. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 19(2):415-428 [M. Clapham/M. Clapham]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 117246: authorized by Matthew Clapham, entered by Matthew Clapham on 18.09.2011, edited by Jered Karr
Creative Commons license: CC BY (attribution)
Taxonomic list
Show authors, comments, and common names
| Insecta | |
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"Rhipidia maior n. sp." = Tipula maior Unger 1841 crane fly | |
Rhipidia extincta n. sp.
Rhipidia extincta n. sp. Unger 1841 crane fly | |
"Protomyia lygaeoides n. sp." = Plecia lygaeoides5, "Protomyia latipennis n. sp." = Plecia lygaeoides5, Bibio enterodelus n. sp., "Bibio giganteus n. sp." = Bibio giganteus, "Bibio maculatus" = Bibio giganteus5, Bibio firmus n. sp.5, "Bibio pinguis n. sp." = Dilophus pinguis5, "Protomyia longa" = Penthetria longa5, "Bibio gracilis n. sp." = Penthetria gracilis, "Bibiopsis cimicoides n. sp." = Penthetria cimicoides5, "Bibio murchisonis n. sp." = Penthetria murchisonii
"Protomyia lygaeoides n. sp." = Plecia lygaeoides5 Heer 1849 love bug Bibio lignarius in Unger (Pl 72, fig 5B)
Bibio enterodelus n. sp. Unger 1841 march fly
"Bibio giganteus n. sp." = Bibio giganteus Unger 1841 march fly
"Bibio pinguis n. sp." = Dilophus pinguis5 Heer 1849 march fly SLJG 77473; Bibio pinguis radobojanus
"Bibio gracilis n. sp." = Penthetria gracilis Unger 1841 march fly
"Bibiopsis cimicoides n. sp." = Penthetria cimicoides5 Heer 1849 march fly SLJG, no catalogue number (Skartveit & Krizmanic, 2020)
"Bibio murchisonis n. sp." = Penthetria murchisonii Unger 1841 march fly | |
Leptogaster hellii n. sp.
Leptogaster hellii n. sp. Unger 1841 grass fly | |
"Tephritis antiqua n. sp." = Chetostoma antiqua5
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Bibio ungeri n. sp.5
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Anthomyia morio n. sp.5
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"Libellula platyptera n. sp." = Croatocordulia platyptera1
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"Lestes coloratus n. sp." = Phenacolestes coloratus3
"Lestes coloratus n. sp." = Phenacolestes coloratus3 Hagen 1848 damselfly Calopteryx sp. in Charpentier, 1841. Specimen lost
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Heterogaster radobojanus n. sp.6
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Aradus antediluvianus n. sp.6
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Coreites oblongus n. sp.6, Coreites crassus n. sp.6, Palaeocoris spectabilis n. gen. n. sp.6, Spartocerus maculatus n. sp.6
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Tingis obscura n. sp.6
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"Cercopis ungeri n. sp." = Aphrophora ungeri6
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Cercopis longicollis n. sp.6, "Hylotoma cineracea n. sp." = Cercopis charpentieri1, "Cercopis fasciata" = Cercopis oswaldi6, "Cercopis haidingeri" = Locrites haidingeri6
"Hylotoma cineracea n. sp." = Cercopis charpentieri1 Heer 1853 froghopper 6 in Gratz and Wien collections (arbitrarily divided in half); unnecessarily replaced by Cercopis charpentieri Heer
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"Acocephalus curtulus n. sp." = Aphrodes curtulus6
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Lachnus pectorosus n. sp.6, "Aphis pallescens" = Greenidea pallescens6, "Aphis morloti" = Greenidea morloti6
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"Myrmeleon brevipenne n. sp." = Gryllacris brevipennis1
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Oedipoda melanosticta n. sp.1
Oedipoda melanosticta n. sp.1 Charpentier 1843 spur-throated grasshopper VGB 73, 78 in Zeuner, 1941; Heer mentions three but Charpentier illustrates five
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"Termes pristinus n. sp." = Macrotermes pristinus1
"Termes pristinus n. sp." = Macrotermes pristinus1 Charpentier 1843 termite 5 (1 questionable) by Heer; three were illustrated by Charpentier
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"Termes (Termopsis) procerus n. sp." = Miotermes procerus5 Heer 1849 termite Includes T. pristinus Charpentier Pl 23, fig 4
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"Formica longipennis n. sp." = Lasius longipennis5, "Formica globularis n. sp." = Lasius globularis5, "Formica capito" = Lasius globularis7, "Formica ophthalmica n. sp." = Lasius ophthalmicus5, "Lasius ophthalmicus" = Lasius ophthalmicus2, "Formica occultata n. sp." = Lasius occultatus5, "Formica obliterata n. sp." = Lasius occultatus5, "Formica minutula n. sp." = Lasius occultatus5, "Formica pumila n. sp." = Lasius occultatus5, "Poneropsis anthracina" = Lasius anthracinus8, "Formica longaeva n. sp." = Lasius longaevus5, Formica obscura5, "Formica ungeri n. sp." = Formica ungeri5, "Ponera elongatula n. sp." = Formica ungeri5, "Formica kollari n. sp." = Formica ungeri8, "Formica acuminata n. sp." = Formica ungeri5, "Formica oblita n. sp." = Formica ungeri8, "Formica aemula n. sp." = Formica ungeri8, "Formica redtenbacheri n. sp." = Formica ungeri5, "Formica indurata n. sp." = Camponotus induratus5, "Camponotus induratus" = Camponotus induratus2, Attopsis longipennis n. sp.5, Attopsis longipes n. sp.8, Attopsis longipes8, Formica fragilis n. sp.5, "Formica obesa n. sp." = Oecophylla obesa5, "Attopsis nigra n. sp." = Oecophylla obesa5, "Formica pinguis" = Oecophylla obesa5, "Attopsis anthracina n. sp." = Oecophylla obesa5, "Myrmica bremii n. sp." = Gesomyrmex bremii5, "Formica ocella n. sp." = Emplastus ocellus5, Emplastus miocenicus2, Emplastus dubius n. sp.2, Dolichoderus heeri n. sp.2, "Formica imhoffii n. sp." = Liometopum imhoffii5, "Liometopum imhoffii" = Liometopum imhoffii2, "Ponera affinis n. sp." = Liometopum imhoffii5, "Poneropsis livida n. sp." = Liometopum imhoffii8, "Poneropsis lugubris n. sp." = Liometopum imhoffii8, "Poneropsis morio n. sp." = Liometopum imhoffii8, "Formica schmidtii n. sp." = Liometopum imhoffii5, "Formica gracilis n. sp." = Ponerites gracilior8, "Formica atavina n. sp." = Ponerites atavinus5, "Ponera nitida n. sp." = Ponerites nitidus5, "Poneropsis tenuis n. sp." = Ponerites tenuis8, Formica obtecta n. sp.5, Formica obvoluta n. sp.5, "Formica longiventris n. sp." = Casaleia longiventris5, "Myrmica pusilla n. sp." = Myrmecites pusillus5, "Myrmica tertiaria" = Paraphaenogaster tertiaria5, "Myrmica bicolor n. sp." = Paraphaenogaster tertiaria8, "Myrmica jurinei n. sp." = Paraphaenogaster jurinei5, "Myrmica concinna n. sp." = Paraphaenogaster jurinei8, "Formica freyeri n. sp." = Lonchomyrmex freyeri8, Formica oculata n. sp.5
"Formica longipennis n. sp." = Lasius longipennis5 Heer 1849 ant UMJG&P 77.503 (surviving syntype), 77.644a (F. pumila in Heer, pl 11, fig 15a), 77.646a (F. pumila in Heer, pl 10, fig 9h)
"Formica ophthalmica n. sp." = Lasius ophthalmicus5 Heer 1849 ant UMJG&P 77.646e (only of Heer's syntypes belonging to this species), 77.492, 77.496, 77.508, 77.517, 77.557, 77.588; Heer 1867 lists 32 specimens.
"Lasius ophthalmicus" = Lasius ophthalmicus2 Heer 1849 ant ssp. radobojana; UMJG&P 77.539 (fig. in 1867); Heer 1867 lists 41 specimens.
"Formica occultata n. sp." = Lasius occultatus5 Heer 1849 ant UMJG&P 77.499, 77.505, 77.512, 77.514, 77.529, 77.579, 77.587, 211 030 (Heer listed 549 specimens but most are other species or unidentifiable)
"Formica obliterata n. sp." = Lasius occultatus5 Heer 1849 ant UMJG&P 77.565 (Heer's two syntypes lost)
"Poneropsis anthracina" = Lasius anthracinus8 Heer 1867 ant Heer's four syntypes lost; UMJG&P 77.497, 77.506, 77.590 in Dlussky & Putyatina 2014
"Formica longaeva n. sp." = Lasius longaevus5 Heer 1849 ant UMJG&P 77.580 (surviving Heer syntype), 77.504, 77.511, 77.570, 77.575, 211 031 (Heer 1867 mentions 16)
Formica obscura5 Heer 1849 ant UMJG&P 77.536 (Heer also included three abdomens, which are unidentifiable: Dlussky & Putyatina 2014)
"Formica ungeri n. sp." = Formica ungeri5 Heer 1849 ant UMJG&P 77.566, 77.548, 77.502, 77.516 (F. ophthalmica in Heer; pl 9, fig 9b), 77.526, 77.531, 77.532, 77.533, 77.542, 77.551, 211 029, 211 033, 211 027, 211 034 (Heer mentions 10 in 1849 and 50 in 1867, but many are other species or unidentifiable: Dlussky & Putyatina 2014)
"Ponera elongatula n. sp." = Formica ungeri5 Heer 1849 ant UMJG&P 77.641 (in Dlussky & Putyatina 2014; different from Heer's figured syntype)
"Formica aemula n. sp." = Formica ungeri8 Heer 1849 ant UMJG&P 77.519 (Heer described five, but three are unidentifiable: Mayr 1867 and Dlussky & Putyatina 2014)
"Formica redtenbacheri n. sp." = Formica ungeri5 Heer 1849 ant UMJG&P 77.510 (Heer mentions 12 in 1849 and 62 in 1867, but many are other species or unidentifiable: Dlussky & Putyatina 2014)
"Camponotus induratus" = Camponotus induratus2 Heer 1849 ant ssp. radobojana; UMJG&P 77.644c, 645h, 645i, 646b, 646g
Dolichoderus heeri n. sp.2 Dlussky and Putyatina 2014 ant UMJG&P 77.507a, 77.568 (both F. occultata in Heer), 77.569a (F. redtenbacheri in Heer), 77.507b, 77.569b
"Formica imhoffii n. sp." = Liometopum imhoffii5 Heer 1849 ant 3 males in Heer; gyne UMJG&P 77.491 in Dlussky & Putyatina 2014 (6 more from unspecified coll. in Heer 1867)
"Liometopum imhoffii" = Liometopum imhoffii2 Heer 1849 ant ssp. radobojana UMJG&P 592(593), 77.637 (Heer 1867 mentions 5 specimens from unspecified collections)
"Formica schmidtii n. sp." = Liometopum imhoffii5 Heer 1849 ant UMJG&P 77.594, 77.520, 77.577, 77.585, 77.623 (Heer 1867 mentions 39 specimens from unspecified collections)
"Ponera nitida n. sp." = Ponerites nitidus5 Heer 1849 ant Heer's holotype lost; neotype UMJG&P 77.622
Formica obtecta n. sp.5 Heer 1849 ant UMJG&P 77.646c (Heer also included an isolated gaster, which is unidentifiable beyond family level)
"Myrmica pusilla n. sp." = Myrmecites pusillus5 Heer 1849 ant Heer's holotype lost (presumably from this collection)
"Myrmica tertiaria" = Paraphaenogaster tertiaria5 Heer 1849 ant Specimens examined by Heer and Mayr (not lost)
"Myrmica jurinei n. sp." = Paraphaenogaster jurinei5 Heer 1849 ant Heer's holotype lost; neotype UMJG&P 211 032 (ID as Attopsis anthracina by Heer)
"Formica freyeri n. sp." = Lonchomyrmex freyeri8 Heer 1867 ant Heer had a single holotype and Mayr examined another; both now lost (presumably type at least from this collection)
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Myrmeleon reticulatum n. sp.1
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"Colymbetes ungeri n. sp." = Agabus ungeri4
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Hydrobius longicollis n. sp.4
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Nitidula radobojana n. sp.4
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Acanthoderes phrixi n. sp.4
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"Sphinx atavus n. sp." = Nymphalites atavus1
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