Where: Japan (35.5° N, 134.3° E: paleocoordinates 38.0° N, 131.7° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Fuganji Formation, Middle Miocene (16.0 - 11.6 Ma)

• bed-level stratigraphic resolution

Environment/lithology: lacustrine; lithified mudstone

Size class: macrofossils

Preservation: adpression

Collected by M. Ota, Y. Okazaki in 31, 1980

Primary reference: K. Ueda. 1989. A Miocene fossil of long-armed scarabaeid beetle from Tottori, Japan. Bulletin of the Kitakyushu Museum of Natural History 9:105-110 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 123353: authorized by Matthew Clapham, entered by Matthew Clapham on 19.01.2012

Creative Commons license: CC BY (attribution)