Shanwang, Linqu County, collection s (Shandong Provincial Museum) (Miocene of China)

Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)

• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone

• Two minor volcanic faulted basins, the Linqu and Shanwang Basins, are present in the study area. The former lies near the county town of Linqu, and the latter in the Shanwang National Geopark, east of this town. The faulted basins are the result of intense Mesozoic-Cenozoic tectonic extension and rifting; the development of these basins was closely related in time with the nearby NE-SW and NW-SE trending faults in the Yishu Fault Zone.

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Shandong Provincial Museum. This collection contains specimens labeled with five-digit numbers prefaced by letter s.

Primary reference: J. F. Zhang. 1989. Fossil Insects From Shanwang, Shandong, China 1-459 [D. Smith/E. Leckey/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 128552: authorized by Matthew Clapham, entered by Matthew Clapham on 11.06.2012

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hemiptera - Fulgoridae
Limois shanwangensis Hong 1979 fulgorid planthopper
s82668, s82671
 Hemiptera - Aphididae
"Cinara limnogena n. sp." = Cinara (Cinara) limnogena, "Cinara elegans n. sp." = Cinara (Cinara) elegans
"Cinara limnogena n. sp." = Cinara (Cinara) limnogena Zhang 1989 aphid
s82673
"Cinara elegans n. sp." = Cinara (Cinara) elegans Zhang 1989 aphid
s82674
 Hemiptera - Coreidae
Stictopleurus breviantennus n. sp. Zhang 1989 leaf-footed bug
s82703
 Hemiptera - Urostylididae
? Urochela sp. Dallas 1851 true bug
s82674
Urochela falsirubra n. sp. Zhang 1989 true bug
s82677
Urochela pardalina n. sp. Zhang 1989 true bug
s82678
 Hemiptera - Pentatomidae
Carbula aff. crassiventris Dallas 1849 stink bug
s82676
Carbula pustulosa n. sp. Zhang 1989 stink bug
s82675
Stollia rubibrunneus n. sp. Zhang 1989 stink bug
s82716
 Hemiptera - Tingidae
Agramma tertiarium n. sp. Zhang 1989 lace bug
s82730
 Hemiptera - Nepidae
Laccotrephes cf. robustus Stal 1870 true bug
s82717
 Hemiptera - Corixidae
Corixa triquetra n. sp. Zhang 1989 water boatman
s82679
 Blattodea - Blattidae
Periplaneta lacera n. sp. Zhang 1989 cockroach
s82696
 Blattodea - Kalotermitidae
"Kalotermes shandongianus n. sp." = Glyptotermes shandongianus
"Kalotermes shandongianus n. sp." = Glyptotermes shandongianus Zhang 1989 termite
s82767
 Dermaptera - Forficulidae
Allodahlia shanwangensis Zhou 1986 common earwig
s82727, s82729
 Orthoptera - Acrididae
Tumearolium humifuse n. gen. n. sp.
Tumearolium humifuse n. gen. n. sp. Zhang 1989 slant-faced grasshopper
s82697
 Hymenoptera - Vespidae
"Vespa magnifica" = Vespa mandarinia magnifica
"Vespa magnifica" = Vespa mandarinia magnifica Smith 1852 Asian giant hornet
s82640
 Hymenoptera - Tiphiidae
Tiphia brevala n. sp. Zhang 1989 wasp
s82641
Tiphia varia n. sp. Zhang 1989 wasp
s82633
Tiphia mediovena n. sp. Zhang 1989 wasp
s82642
 Hymenoptera - Formicidae
Formica ovala Zhang 1989 ant
s82624
Formica linquensis Zhang 1989 ant
s82623
Camponotus ambon n. sp. Zhang 1989 ant
s82621, s82622
Camponotus ampullosus n. sp. Zhang 1989 ant
s82603
Camponotus microthoracus Zhang 1989 ant
s82620
Camponotus festinus Smith 1857 ant
s82602
Camponotus gracilis n. sp. Zhang 1989 ant
s82606
Camponotus shanwangensis Hong 1983 ant
s82601, s82604, s82605
Myopopone sinensis n. sp. Zhang 1989 ant
s82628
Elaphrodites scutulatus n. gen. n. sp. Zhang 1989 ant
s82636
Dolichoderus lacinius n. sp. Zhang 1989 ant
s82627
Dolichoderus evolans n. sp. Zhang 1989 ant
s82625
Technomyrmex septentrionalis n. sp. Zhang 1989 ant
s82626
Alloiomma changweiensis n. sp. Zhang 1989 ant
s82629
 Hymenoptera - Megachilidae
Megachile shanwangae n. sp. Zhang 1989 mason bee
s82638
 Hymenoptera - Melittidae
"Dasypoda basaltica n. sp." = Macropis basaltica
"Dasypoda basaltica n. sp." = Macropis basaltica Zhang 1989 bee
s82637
 Hymenoptera - Apidae
Xylocopa diatoma n. sp.2 Zhang 1990 large carpenter bee
s82772
Bombus luianus n. sp.2 Zhang 1990 bumble bee
s82771
Apis longitibia n. sp.2 Zhang 1990 long-legged honey bee
s82773
 Hymenoptera - Ichneumonidae
Exetastes areolus n. sp. Zhang 1989 ichneumon wasp
s82645
Corynobatopsis anfracta n. gen. n. sp. Zhang 1989 ichneumon wasp
s82649
Agrothereutes townesi n. sp. Zhang 1989 ichneumon wasp
s82648
? Ovipterygium flexuosum n. sp. Zhang 1989 ichneumon wasp
s82647
Mesoleptus stigmus n. sp. Zhang 1989 ichneumon wasp
s82646
Yezoceryx brachycoleos n. sp. Zhang 1989 ichneumon wasp
s82644
 Hymenoptera - Scelionidae
"Protelenomus miocenicus n. gen. n. sp." = Sinoprotelenomus miocenicus
"Protelenomus miocenicus n. gen. n. sp." = Sinoprotelenomus miocenicus Zhang 1989 wasp
s82643
 Hymenoptera - Tenthredinidae
Selandria perelegans n. sp. Zhang 1989 wood wasp
s82715
 Coleoptera - Dascillidae
Dascillus sp. Latreille 1796 soft bodied plant beetle
s82750
Dascillus shandongianus Zhang 1989 soft bodied plant beetle
s82732, s82733, s82740, s82741, s82742
Dascillus flatus n. sp. Zhang 1989 soft bodied plant beetle
s82734, s82735, s82736
 Coleoptera - Cantharidae
"Curticantharis sp." = Themus
"Curticantharis sp." = Themus Motschulsky 1858 soldier beetle
s82753
 Coleoptera - Elateridae
Elater mitrus n. sp. Zhang 1989 click beetle
s82713
Elater asmodeus n. sp. Zhang 1989 click beetle
s82747
 Coleoptera - Hydrophilidae
Enochrus lapsus n. sp., Helochares mydaleus n. sp., "Hydrous pergrandis n. sp." = Hydrophilus pergrandis
Enochrus lapsus n. sp. Zhang 1989 water scavenger beetle
s82714
Helochares mydaleus n. sp. Zhang 1989 water scavenger beetle
s82720
"Hydrous pergrandis n. sp." = Hydrophilus pergrandis Zhang 1989 water scavenger beetle
s82719
 Coleoptera - Staphylinidae
Medon stenus n. sp. Zhang 1989 rove beetle
s82699
Lathrobium gensium n. sp. Zhang 1989 rove beetle
s82698
 Coleoptera - Tenebrionidae
Dascillus exiguus n. sp. Zhang 1989 comb-clawed beetle
s82731
 Coleoptera - Cleridae
Cleridae indet. Latreille 1802 checkered beetle
s82748
 Coleoptera - Endomychidae
"Engonius alviolatus" = Gerstaeckerus alviolatus
"Engonius alviolatus" = Gerstaeckerus alviolatus Hong 1983 handsome fungus beetle
s82743, s82744, 82745, s82746
 Coleoptera - Coccinellidae
Scymnus (Pullus) aff. kawamurai Ohta 1929 ladybug or ladybird
s82722
 Coleoptera - Cerambycidae
Stromatium cf. barbatum Fabricius 1775 longhorned beetle
s82708
Mesosa viaria n. sp. Zhang 1989 longhorned beetle
s82709
Delostromatium curvum n. gen. n. sp. Zhang 1989 longhorned beetle
s82705
Alifasciola sirdena n. gen. n. sp. Zhang 1989 longhorned beetle
s82693
Trosites foveolatus n. gen. n. sp. Zhang 1989 longhorned beetle
s82695
Protaxis chinensis n. sp. Zhang 1989 longhorned beetle
s82694
 Coleoptera - Chrysomelidae
Chrysomelidae indet. Latreille 1802 leaf beetle
s82749
Chrysothoracus tropicus n. gen. n. sp. Zhang 1989 leaf beetle
s82701
 Coleoptera - Curculionidae
Hylobius turfaceus n. sp. Zhang 1989 snout beetle
s82691
Hylobius plenus Zhang 1989 snout beetle
s82690
 Coleoptera - Attelabidae
Oedeuops ephemera n. gen. n. sp.
Oedeuops ephemera n. gen. n. sp. Zhang 1989 leaf rolling weevil
s82689
 Coleoptera - Brentidae
"Balanobius edentis n. sp." = Longifistulia edentis, "Balanobius crassipedalis n. sp." = Longifistulia crassipedalis
"Balanobius edentis n. sp." = Longifistulia edentis Zhang 1989 weevil
s82683, s82684, s82685, s82687
"Balanobius crassipedalis n. sp." = Longifistulia crassipedalis Zhang 1989 weevil
s82688
 Coleoptera - Scarabaeidae
Onthophagus sp. Latreille 1802 scarab beetle
s82761
Anomala sp. Samouelle 1819 scarab beetle
s82762, s82763
Anomala furva Hong 1983 scarab beetle
s82755
Anomala scia n. sp. Zhang 1989 scarab beetle
s82710
? Phalops sp. Erichson 1848 scarab beetle
s82760
 Coleoptera -
Scelocopris enertheus n. gen. n. sp.
Scelocopris enertheus n. gen. n. sp. Zhang 1989 scarab beetle
s82754
 Coleoptera - Dytiscidae
Laccophilus corystes n. sp. Zhang 1989 predaceous diving beetle
s82711
Laccophilus pusulatus n. sp. Zhang 1989 predaceous diving beetle
s82721
 Coleoptera - Carabidae
Calosoma cf. maderae Fabricius 1775 ground beetle
s82751
Bembidion strenum n. sp. Zhang 1989 ground beetle
s82706
 Diptera - Chironomidae
Bythomyia oryctes n. gen. n. sp.1
Bythomyia oryctes n. gen. n. sp.1 Zhang 1989 non biting midge
s32469
 Diptera - Pallopteridae
Palloptera hypolithica n. sp. Zhang 1989 fly
s82665
 Diptera - Asilidae
Asilus brunneitibius n. sp. Zhang 1989 robber fly
s82664
Asilus angustialius n. sp. Zhang 1989 robber fly
s82663
 Diptera - Bibionidae
Plecia villosa n. sp. Zhang 1989 love bug
s82650, s82766
Plecia spinula n. sp. Zhang 1989 love bug
s82652, s82675
Plecia aculeolata n. sp. Zhang 1989 love bug
s82651
Bibio colossoceps n. sp. Zhang 1989 march fly
s82658
"Bibio abdominalis" = Bibio brunettii Hardy 1973 march fly
s82659
Bibio expansus n. sp. Zhang 1989 march fly
s82654
Bibio tranopterus Zhang 1989 march fly
s82653
Bibio ventricosus Zhang 1989 march fly
s82655
Bibio varineurus Zhang 1989 march fly
s82718
 Lepidoptera - Hepialidae
Oiophassus nycterus n. gen. n. sp.
Oiophassus nycterus n. gen. n. sp. Zhang 1989 moth
s82702
 Ephemeroptera - Heptageniidae
Heptagenia sp. Walsh 1863 stream mayfly
s82707, s82712, s82724
 Odonata - Coenagrionidae
Agriocnemis lacteola Selys 1877 wisp
s82682
 Odonata - Aeshnidae
"Mediaeschna lucida n. sp." = Epiaeschna lucida
"Mediaeschna lucida n. sp." = Epiaeschna lucida Zhang 1989 darner dragonfly
s82681
 Odonata - Corduliidae
Epophthalmia zotheca n. sp. Zhang 1989 green-eyed skimmer
s82725/6
Arachnida
 Araneae - Thomisidae
"Ectona brunnea n. sp." = Ecotona brunnea3, Miothomisus subnudus n. sp.3
"Ectona brunnea n. sp." = Ecotona brunnea3 Zhang et al. 1994 crab spider
s85611
Miothomisus subnudus n. sp.3 Zhang et al. 1994 crab spider
s83401
 Araneae - Araneidae
Araneus luianus n. sp.3 Zhang et al. 1994 orb-weaving spider
s82704