Where: Shandong, China (36.6° N, 118.7° E: paleocoordinates 36.5° N, 116.3° E)
• coordinate based on nearby landmark
• local area-level geographic resolution
When: Shanwang Formation, Burdigalian (20.4 - 16.0 Ma)
• Paleomagnetic and radiometric evidence correlates the Shanwang Formation to subchron 5En of Chron 5. The main part of the Shanwang Formation covers 18.524 Ma to 18.056 Ma, with the Shanwang Formation as a whole covering 18.5 ma to 17.5 Ma. Recent paleontological results show that the Shanwang Formation can be correlated with the European mammalian zone MN instead of the previous MN5.
• group of beds-level stratigraphic resolution
Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous mudstone
Size classes: macrofossils, mesofossils
Preservation: adpression
Collection methods: Repository: Beijing Museum of Natural History. This collection contains specimens labeled with five-digit numbers starting with 79... or 80...
Primary reference: Y. C. Hong. 1984. Fossil insects in the diatoms of Shanwang. Bulletin of the Tianjin Institute of Geology and Mineral Resources 8:1-15 [D. Smith/E. Leckey/M. Clapham]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 128550: authorized by Matthew Clapham, entered by Matthew Clapham on 11.06.2012, edited by Erin Leckey
Creative Commons license: CC BY (attribution)
Taxonomic list
Arachnida | |
"Retina robusta" = Heteropoda robusta6
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"Eocryphoeca parva" = Tetragnatha parva6
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Insecta | |
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Jagoria sinica n. sp.5
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"Oncotympana lapidescens n. sp." = Tanyocicada lapidescens5
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Tarsonecta mecopoda n. gen. n. sp.5
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Coroides xiejiaheense n. gen. n. sp.
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Manocerus stagnans n. gen. n. sp.5
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Cloresmus shanwangi n. sp.5, Gonocerus shandongensis n. sp.5, "Coriopsis parva n. sp." = Coreiopsis parva3, Coreiopsis punctata n. gen. n. sp., "Coriopsis miocenica n. sp." = Coreiopsis miocenica3, Shanwangicoris longa n. gen. n. sp.3
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"Cinara reconditivenia n. sp." = Cinara (Cinara) reconditivenia5
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"Oxycephala xiejiaheensis" = Limois shanwangensis5, "Fulgoropsis fusca n. gen. n. sp." = Limois shanwangensis, Aphaena lithoecia n. sp.5
"Oxycephala xiejiaheensis" = Limois shanwangensis5 Hong 1979 fulgorid planthopper 79026, 79079, 79080
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Apanechura asecta n. sp.5
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"Megaxyela xiejiaheensis n. sp." = Xyelecia xiejiaheensis
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"Procimbex shanwangensis n. sp." = Abia shanwangensis, Abia paurocephala n. sp.5, "Procimbex shandongense n. sp." = Abia shandongensis1
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Luia lacaris n. gen. n. sp.5
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"Sinostigma spinalata n. sp." = Pemphredon spinalatum
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Apis miocenica n. sp.
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Paraphaenogaster shanwangensis n. sp., "Heteromyrmex atopogaster n. gen. n. sp." = Zhangidris atopogaster5, Formica linquensis n. sp.5, Camponotus mitis5, "Camponotus curviansatus n. sp." = Camponotites curviansatus5, "Rabidia xiejiaheensis n. sp." = Camponotites xiejiaheensis, "Camponotus shanwangense" = Camponotites shanwangensis, "Camponotus microthoracus n. sp." = Camponotites microthoracus5, "Camponotus longus n. sp." = Camponotites longus5, ? Lasius truncatus n. sp.5, Lasius vetulus5, Lasius validus n. sp.5, ? Tapinoma baculum n. sp.5, "Liometopum jiaoyanshanense n. sp." = Dolichoderus jiaoyanshanensis5, "Shanwangella palaeoptera n. gen. n. sp." = Liometopum palaeopterum5, Liometopum potamophilum n. sp.5
"Heteromyrmex atopogaster n. gen. n. sp." = Zhangidris atopogaster5 Zhang 1989 ant 79054 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 3)
Formica linquensis n. sp.5 Zhang 1989 ant 79058 (Paraphaenogaster shanwangense in Hong, 1985 - pl 23, fig 4)
"Camponotus curviansatus n. sp." = Camponotites curviansatus5 Zhang 1989 ant 79042 (Camponotus shanwangensis in Hong, 1985 - pl 27, fig 1)
"Camponotus shanwangense" = Camponotites shanwangensis Hong 1984 ant 79031; 79043, 79076 (Liometopum jiaoyanshanense in Hong, 1985); ten specimens including 80179 in Hong & Wang 1985
Lasius validus n. sp.5 Zhang 1989 ant 79121 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 4)
"Liometopum jiaoyanshanense n. sp." = Dolichoderus jiaoyanshanensis5 Hong 1985 ant 79087; 80196 in Hong & Wang 1985
"Shanwangella palaeoptera n. gen. n. sp." = Liometopum palaeopterum5 Zhang 1989 ant 79051 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 2)
Liometopum potamophilum n. sp.5 Zhang 1989 ant 79036 (Liometopum jiaoyanshanense in Hong, 1985 - pl 26, fig 6)
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Elis pyrula n. sp.5
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"Brachyopteris shanwangensis n. sp." = Tiphia shanwangensis
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"Pseudoperla shanwangensis n. sp." = Eumenes shanwangensis
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Sinocistela siphla n. gen. n. sp.5
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? Epicauta rufithoraca n. sp.5
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"Sinocurculia shanwangensis n. sp." = Atactogaster shanwangensis2, "Sinocurculia granulata n. sp." = Atactogaster granulata, Hylobius plenus5, Hylobius medianus n. sp.5, Hylobius aff. niitakensis5
"Sinocurculia shanwangensis n. sp." = Atactogaster shanwangensis2 Hong 1985 snout beetle 80128, 78001, 80107, 80134, 80173, 80174
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Longifistulia parva n. gen. n. sp.3
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Cerambycidae "indet. 1"5, Mesosa soteria n. sp.5, Sinocalosoma longicornuta n. sp.2, Mesocacia pulla n. sp.5, Alifasciola daedalea n. sp.5
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"Engonius alviolatus" = Gerstaeckerus alviolatus5
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Lathrobium homorum n. sp.5
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Scarabaeoidea "indet. 1"5 Latreille 1802 scarab beetle 79045 (Magniscarabaeus furvus in Hong & Wang - pl 17, fig 2)
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Anomala sp.5, "Magniscarabaeus brunneus n. gen. n. sp." = Anomala palaeobrunnea5, "Magniscarabaeus furvus" = Anomala furva2, Anomala synemosyna n. sp.5, Anomala lochmocola n. sp.5, Holotrichia spatha n. sp.5
"Magniscarabaeus brunneus n. gen. n. sp." = Anomala palaeobrunnea5 Krell 2000 scarab beetle 79105; seven including 80101, 80120, 80172, 80188, 80184, 80194, mentioned in Hong & Wang 1987
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Geotrupidae indet.5 Latreille 1802 earth-boring dung beetle 79107 (Magniscarabaeus brunneus in Hong, 1985 - pl 20, fig 3)
"Obitiscarabaeus jiaoyanshanense n. gen. n. sp." = Geotrupes jiaoyanshanense Hong 1984 earth-boring dung beetle 79114/5
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Aliscarabaeus granulatus n. sp.
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Scarites mancus n. sp.5, Calosoma cf. maderae5, ? Carabus ovalus n. sp.2, "Shanwangicarabus furvus n. sp." = Chlaenius furvus2, "Shanwangicarabus paucinus n. sp." = Chlaenius furvus2
"Shanwangicarabus furvus n. sp." = Chlaenius furvus2 Hong 1984 ground beetle 80125, 80106, 80126, 480183
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Cupes longus n. sp.4
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"Procarabus longipes n. sp." = Cistelites longipes5
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Tipula insolita n. sp.5
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Sciara brevipetiola n. sp.5
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Bibio ruptineurus n. sp.5, Bibio deformatus n. sp.5, "Fungivora bullata n. sp." = Bibio bullatus5, Bibio brunneus n. sp.5, Bibio shanwangensis1, Bibio ventricosus n. sp.5, Bibio expansus5, "Bibio miocenica n. sp." = Bibio miocenicus1, Bibio cratopodus5, Clothonopsis miocenica n. gen. n. sp.4, Plecia ludongensis n. sp.1, Plecia solaris n. sp.5
Bibio deformatus n. sp.5 Zhang 1989 march fly 79057 (Bibio shanwangensis in Hong, 1985 - pl 4, fig 1)
Bibio ventricosus n. sp.5 Zhang 1989 march fly 79033 (Bibio shanwangensis in Hong, 1985 - pl 7, figs 1, 2, 3)
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