Zambrana: MP 18, Spain
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
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Amphibia
- Temnospondyli
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Anura indet.
(Fischer von Waldheim 1813)
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Minwer-Barakat et al. 2013 | |||||||||
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Reptilia
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Diplocynodon sp.
Pomel 1847
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23 specimens | |||||||||
| isolated teeth and osteodermes | ||||||||||
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Lacertilia indet.
(Owen 1842)
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Minwer-Barakat et al. 2013 | |||||||||
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Reptilia
- Testudines
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Chelonia indet.
(Latreille 1800)
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3 specimens | |||||||||
| fragments of plastron | ||||||||||
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= Cryptodira indet.
Cope 1868
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Minwer-Barakat et al. 2013 | |||||||||
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Mammalia
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cf. Quercygale sp.
Kretzoi 1945
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1 specimen | |||||||||
| isolated teeth | ||||||||||
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= Quercygale sp.
Kretzoi 1945
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Minwer-Barakat et al. 2013 | |||||||||
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Paramiacis sp.
Filhol 1876
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Minwer-Barakat et al. 2013 | |||||||||
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Mammalia
- Perissodactyla
- Palaeotheriidae
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Plagiolophus aff. mazateronensis
Cuesta 1994
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1 specimen | |||||||||
| fragment of mandible with teeth | ||||||||||
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= Leptolophus sp.
Remy 1965
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Badiola et al. 2002 | |||||||||
| four additional specimens described in this reference, all teeth and tooth fragments | ||||||||||
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Leptolophus informal new species
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Minwer-Barakat et al. 2013 | |||||||||
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Leptolophus cuestai n. sp.
Perales-Gogenola et al. 2021
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Perales-Gogenola et al. 2021 | |||||||||
| Holotype: MCNA 9948, right M2; MCNA 9947, right M1; MCNA 9002, incomplete left hemimandible preserving the m1–(m2)–(m3) series; MCNA 9949, incomplete right hemimandible showing the p4– m1–(m2)–(m3) series | ||||||||||
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Leptolophus franzeni n. sp.
Perales-Gogenola et al. 2021
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Perales-Gogenola et al. 2021 | |||||||||
| Holotype: UPV/EHU-Z4.1092, cranium fragment with right and left C–P2–P3–P4–M1–M2–(M3) series. Fragmented right P2 and P4, and left P2; MCNA 11848, left M3; MCNA 11849, right M3; UPV/EHU- Z4.1041 fragment of right hemimandible with a par- tially erupted m3. | ||||||||||
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Pachynolophus zambranensis n. sp.
Badiola et al. 2005
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Badiola et al. 2005 | 26 specimens | ||||||||
| MCNA 10657 (holotype). MCNA 10658-10661 (paratypes). 21 referred specimens | ||||||||||
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Paranchilophus sp.
Casanovas and Santafé 1989
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1 specimen | |||||||||
| fragment of mandible with teeth | ||||||||||
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= Paranchilophus remyi
Casanovas and Santafé 1989
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Minwer-Barakat et al. 2013 | |||||||||
| recombined as Anchilophus (Paranchilophus) remyi | ||||||||||
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Palaeotherium informal new species
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2 specimens | |||||||||
| fragments of mandibles with teeth | ||||||||||
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Iberolophus arabensis n. gen., n. sp.
Badiola and Cuesta 2008
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Badiola and Cuesta 2008 | 15 specimens | ||||||||
| MCNA 11953 (holotype). Paratypes: MCNA 10183, skull fragment, showing the right and left M1–M3 molar series; MCNA 11888, left P3–4; MCNA 11906, left P1–2; MCNA 13372, left p4; and MCNA 13373, right p1. Referred material: —MCNA 11904, right P3–4; MCNA 11887, 11893, left P3–4; MCNA 13375, fragment of the left P3–4; MCNA 11905, fragment of the left P3–4; MCNA 11950, right P1-2 (incomplete lingually); MCNA 13374, fragment of the right upper premolar; MCNA 11951, fragment of the upper premolar; MCNA 11952, fragment of the lower premolar | ||||||||||
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Mammalia
- Perissodactyla
- Pachynolophidae
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Mammalia indet.
Linnaeus 1758
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= Plagiolophinae indet.
Remy 1976
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Astibia et al. 2000 | |||||||||
| isolated tooth | ||||||||||
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Mammalia
- Dichobunidae
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Dichobune leporina
Cuvier 1822
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Minwer-Barakat et al. 2013 | |||||||||
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Mammalia
- Anoplotheriidae
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Anoplotherium sp.
Cuvier 1804
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Minwer-Barakat et al. 2013 | |||||||||
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Anoplotherium laurillardi
Pomel 1851
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Minwer-Barakat et al. 2013 | |||||||||
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Mammalia
- Xiphodontidae
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Haplomeryx sp.
Schlosser 1886
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Minwer-Barakat et al. 2013 | |||||||||
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Xiphodontidae indet.
Flower 1883
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1 specimen | |||||||||
| fragment of mandible with teeth | ||||||||||
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= Xiphodon gracilis
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Minwer-Barakat et al. 2013 | |||||||||
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Mammalia
- Rodentia
- Theridomyidae
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Pseudoltinomys sp.
Lavocat 1951
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Minwer-Barakat et al. 2013 | |||||||||
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Elfomys aff. parvulus
Hartenberger 1971
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7 specimens | |||||||||
| isolated teeth | ||||||||||
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= Elfomys parvulus
Hartenberger 1971
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Minwer-Barakat et al. 2013 | |||||||||
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Theridomys aff. golpei
Hartenberger 1973
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6 specimens | |||||||||
| isolated teeth | ||||||||||
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Pseudosciuridae indet.
Zittel 1893
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1 specimen | |||||||||
| isolated tooth | ||||||||||
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= Pseudosciurinae indet.
Lavocat 1951
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Astibia et al. 2000 | |||||||||
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Remys sp.
Thaler 1966
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Minwer-Barakat et al. 2013 | |||||||||
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Mammalia
- Rodentia
- Gliridae
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Gliravus priscus
Stehlin and Schaub 1951
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1 specimen | |||||||||
| isolated tooth | ||||||||||
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= Glamys priscus
Stehlin and Schaub 1951
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Astibia et al. 2000 | |||||||||
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Mammalia
- Primates
- Omomyidae
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Microchoerus aff. erinaceus
Wood 1844
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Minwer-Barakat et al. 2013 | 2 specimens | ||||||||
| Right mandible fragment with p3-m3 (MCNA 14518); right mandible fragment with p4-m3 (MCNA 14519). | ||||||||||
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Mammalia
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Peratherium cuvieri
Fisher 1829
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Badiola and Cuesta 2006 | 2 specimens | ||||||||
| MCNA 11853, 118854, hemimandibles | ||||||||||
| see common names | ||||||||||
Geography
| Country: | Spain | State/province: | Basque County | County: | Alava |
| Coordinates: | 42.7° North, 2.9° West (view map) | ||||
| Paleocoordinates: | 41.3° North, 6.1° West | ||||
| Basis of coordinate: | based on nearby landmark | ||||
| Altitude: | 512 meters |
Time
| Period: | Paleogene | Epoch: | Eocene |
| Stage: | Priabonian | 10 m.y. bin: | Cenozoic 3 |
| Key time interval: | MP 18 | ||
| Age range of interval: | 37.71000 - 33.90000 m.y. ago |
Stratigraphy
| Stratigraphic resolution: | group of beds | ||||
| Stratigraphy comments: biozone MP 18, early middle Ludian (Priabonian)
European Headonian Land Mammal Age corresponding to the reference level MP 18 (Astibia et al., 2000; Badiola, 2004; Badiola et al., 2009b). | |||||
Lithology and environment
| Primary lithology: | carbonaceous marl |
| Secondary lithology: | "limestone" |
| Lithology description: lacustrine coal-bearing marls and limestones the fossiliferous beds, which consist of palustrine- lacustrine coal-bearing marls and marlstones, are located in one of the oldest lacustrine systems in the Miranda-Treviño Basin (Basque-Cantabrian Region) (Astibia et al., 2000) | |
| Environment: | lacustrine indet. |
| Geology comments: the basin was highly influenced by tectonic activity, and the first compressive stresses related to the Pyrean phase of the Alpine orogenesis that took place during the early Eocene
The depositional setting was a shallow, low-gradient freshwater lake margin with a peripheral swamp (Iriarte et al., 2003; Badiola et al., 2009a). | |
Taphonomy
| Modes of preservation: | body |
| Size of fossils: | macrofossils,mesofossils |
| Disassociated minor elements: | all |
Collection methods and comments
| Collection methods: | field collection |
| Reason for describing collection: | taxonomic analysis |
Metadata
| Database number: | 38817 | ||
| Authorizer: | J. Alroy, P. Mannion, R. Butler | Enterer: | E. Leckey, R. Butler, G. Varnham |
| Modifier: | G. Varnham | Research group: | vertebrate |
| Created: | 2004-05-06 11:12:35 | Last modified: | 2022-04-26 04:25:58 |
| Access level: | the public | Released: | 2004-05-06 11:12:35 |
| Creative Commons license: | CC BY |
Reference information
Primary reference:
| 10455. | A. Badiola, H. Astibia, A. Aranburu, X. Pereda Suberbiola, X. Murelaga, C. Sese, M. A. Cuesta, S. Moya-Sola, J. I. Baceta and M. Kohler. 1999. A new continental vertebrate locality from the Upper Eocene of Zambrana (Miranda-Trevino Basin, Alava, Basque Country). IV European Workshop on vertebrate Paleontology, Albarracin, Spain 19 [J. Alroy/E. Leckey/E. Leckey] |
Secondary references:
| 10459 | H. Astibia, A. Aranburu, X. Pereda Suberbiola, X. Murelaga, C. Sesé, M. A. Cuesta, S. Moyà-Solà, J. E. Baceta, A. Badiola and M. Köhler. 2000. Un nouveau site à vertébrés continentaux de l'Éocène supérieur de Zambrana (Bassin de Maranda-Treviño, Alava, Pays basque). Géobios 32(2):233-248 [J. Alroy/E. Leckey/J. Alroy] | |
| 10407 | A. Badiola, H. Astibia, X. Pereda Suberbiola and X. Murelaga. 2002. First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe. Geodiversitas 24(4):841-848 [J. Alroy/E. Leckey/E. Leckey] | |
| 60655 | A. Badiola, L. Checa, M. A. Cuesta, R. Quer, J. J. Hooker and H. Astibia. 2009. The role of new Iberian finds in understanding European Eocene mammalian paleobiogeography. Geologica Acta 7:243-258 [R. Butler/R. Butler] | |
| 60631 | A. Badiola and M. A. Cuesta. 2006. Los marsupiales del yacimiento del Eoceno Superior de Zambrana (Álava, Región Vasco-Cantábrica). Estudios Geológicos 62:349-358 [R. Butler/R. Butler] | |
| 60598 | A. Badiola and M.-A. Cuesta. 2008. New endemic Eocene equoids from the Iberian Peninsula (western Europe). Journal of Vertebrate Paleontology 28(4):1149-1161 [R. Butler/R. Butler] | |
| 60632 | A. Badiola, X. Pereda-Suberbiola, and M. A. Cuesta. 2005. Una nueva especie de Pachynolophus (Mammalia, Perissodactyla) de Zambrana (Álava, Región Vasco-Cantábrica). Análisis filogenético de Pachynolophus y primera cita en el Eoceno superior de la Península Ibérica. Geobios 38(1):1-16 [R. Butler/R. Butler/R. Butler] | |
| 60647 | R. Minwer-Barakat, A. Badiola, J. Marigo and S. Moya-Sola. 2013. First record of the genus Microchoerus (Omomyidae, Primates) in the western Iberian Peninsula and its palaeobiogeographic implications. Journal of Human Evolution 65(3):313-321 [R. Butler/R. Butler] | |
| 81458 | L. Perales-Gogenola, A. Badiola, A. Gómez-Olivencia and X. Pereda-Suberbiola. 2021. New Leptolophus (Palaeotheriidae) species from the Iberian Peninsula and early evidence of hypsodonty in an Eocene perissodactyl. Journal of Vertebrate Paleontology 41(1):e1912061 [P. Mannion/G. Varnham] |