Muthmannsdorf: Early/Lower Campanian, Austria
collected by Suess & Pawlowitsch 1870

List of taxa
Where & when
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Testudines
Chelonia indet. (Latreille 1800)
Bunzel 1871 2 specimens
synonym of Testudines
"Chersiten und Paludinosen"
Reptilia - Testudines - Dortokidae
Rabi et al. 2013 2 specimens
Reptilia - Testudines
Emys informal sp. 2 Dumeril 1806
Seeley 1881
    = cf. Kallokibotioninae indet. Nopsca 1923
Rabi et al. 2013
Emys informal sp. 3 Dumeril 1806
Seeley 1881
    = cf. Kallokibotioninae indet. Nopsca 1923
Rabi et al. 2013
Emys neumayri n. sp. Seeley 1881
Seeley 1881
    = cf. Kallokibotioninae indet. Nopsca 1923
Rabi et al. 2013
    = Kallokibotion sp. Nopcsa 1923
Joyce 2017
Reptilia - Dinosauria
Dinosauria indet. Owen 1842
Bunzel 1871
Megalosaurus pannoniensis n. sp. Seeley 1881
Seeley 1881 2 specimens
nomen dubium belonging to Tetanurae
PIUW uncat.
Reptilia - Nodosauridae
Struthiosaurus n. gen. sp. Bunzel 1870
    = Struthiosaurus austriacus n. sp. Bunzel 1871
Bunzel 1871
PIWU 2349/6
Danubiosaurus anceps n. gen., n. sp. Bunzel 1871
Bunzel 1871 1 specimen
synonym of Struthiosaurus austriacus
Pleuropeltus suessii n. gen., n. sp. Seeley 1881
Seeley 1881
synonym of Struthiosaurus austriacus
? Hylaeosaurus sp. Mantell 1833
Bunzel 1871 1 specimen
    = Crataeomus pawlowitschii n. gen., n. sp. Seeley 1881
Seeley 1881
synonym of Struthiosaurus austriacus
Scelidosaurus sp. Owen 1861
Bunzel 1871 15 specimens
    = Crataeomus lepidophorus n. sp. Seeley 1881
Seeley 1881
synonym of Struthiosaurus austriacus
Rhadinosaurus alcimus n. gen., n. sp. Seeley 1881
Seeley 1881
nomen dubium belonging to Ankylosauria
(15 measurements)
Reptilia - Rhabdodontidae
Iguanodon sp. Mantell 1825
Bunzel 1871 1 specimen
    = Mochlodon sp. Seeley 1881
Seeley 1881
    = Zalmoxes sp. Weishampel et al. 2003
Sachs and Hornung 2006
Lacerta sp. Linnaeus 1758
Bunzel 1871 4 specimens
    = Zalmoxes sp. Weishampel et al. 2003
Sachs and Hornung 2006
Iguanodon suessi n. sp. (Bunzel 1871)
Bunzel 1871 9 specimens
recombined as Rhabdodon suessii
Lectotype right dentary (PIUW 2349/2); dDentary tooth (PIUW 2349/3); maxillary tooth (PIUW 2349/4); fragmentary parietal (PIUW 2349/54); fragmentary left scapula (PIUW 3518); fragmentary ?radius (PIUW 3517); ?manual ungual (PIUW 2349/38); fragmentary left femur (PIUW 2349/3); fragmentary ?right tibia (PIUW 2348/35) listed by Osi et al. (2012) (3 measurements)
Oligosaurus adelus n. gen., n. sp. Seeley 1881
Seeley 1881
synonym of Rhabdodon priscus
Ornithomerus gracilis n. gen., n. sp. Seeley 1881
Seeley 1881
synonym of Rhabdodon priscus
Pterosauria indet. (Kaup 1834)
Buffetaut et al. 2011 1 specimen
UWPI 2349/103, five larger and six smaller fragments of wing phalanges.
Reptilia - Azhdarchidae
Ornithocheirus sp. Seeley 1869
Wellnhofer 1980 1 specimen
UWPI 2349/102, proximal half of right humerus
    = ? Nyctosauridae indet. Hay 1902
Jianu et al. 1997
    = Azhdarchidae indet. Nessov 1984
Buffetaut et al. 2011
Ornithocheirus bunzeli n. sp. Seeley 1881
Seeley 1881 1 specimen
nomen dubium belonging to Pterodactyloidea
UWPI 2349/101, rear end of left lower jaw
Crocodilus carcharidens n. sp. Bunzel 1871
Bunzel 1871 1 specimen
recombined as Doratodon carcharidens
IPUW 2349/57
Reptilia - Crocodilia
Crocodilia indet. Owen 1842
Bunzel 1871 1 specimen
spelled with current rank as Crocodylia
described as "Crocodili ambigui"
Crocodilus sp. (Laurenti 1768)
Bunzel 1871 12 specimens
    = Crocodilus proavus n. sp. Seeley 1881
Seeley 1881
nomen dubium belonging to Crocodylomorpha
Reptilia - Squamata
Araeosaurus gracilis n. gen., n. sp. Seeley 1881
Seeley 1881
Reptilia - Champsosauridae
Champsosauridae indet. Cope 1876
Buffetaut 1989 2 specimens
2349/55a, 2349/55b (vertebral centra), referred to Iguanodon suessi by Bunzel (1871)
see common names

Country:Austria State/province:Niederosterreich
Coordinates: 47.8° North, 16.2° East (view map)
Paleocoordinates:33.1° North, 17.9° East
Basis of coordinate:based on nearby landmark
Altitude:262 meters
Geographic resolution:outcrop
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
Key time interval:Early/Lower Campanian
Age range of interval:83.50000 - 70.60000 m.y. ago
Geological group:Lower Gosau Formation:Grünbach
Stratigraphic resolution:bed
Lithology and environment
Primary lithology: coal
Secondary lithology: sandstone
Includes fossils?Y
Lithology description: coal-bearing beds, interbedded with sandstones and marls
Modes of preservation:body
Lagerst�tten type:concentrate
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils
Disassociated major elements:many
Disassociated minor elements:many
Collection methods and comments
Collection excludes:species names
Collection methods:selective quarrying,surface (float),surface (in situ),mechanical,field collection
Reason for describing collection:general faunal/floral analysis
Collectors:Suess & Pawlowitsch Collection dates:1870
Also known as:Gosau, Neue Welt, Weiner Neustadt
Database number:48720
Authorizer:M. Carrano, P. Holroyd, R. Butler, P. Mannion, R. Benson Enterer:M. Carrano, P. Holroyd, R. Butler, P. Mannion, R. Benson
Modifier:M. Carrano Research group:vertebrate
Created:2005-03-29 14:12:08 Last modified:2019-02-13 14:54:26
Access level:the public Released:2005-03-29 14:12:08
Creative Commons license:CC BY
Reference information

Primary reference:

12805.ETE E. Bunzel. 1870. Notice of a fragment of a reptilian skull from the Upper Cretaceous of Grünbach. Quarterly Review of the Geological Society of London 26:394 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

37976 E. Buffetaut. 1989. Erster nachweis von Choristodera (Reptilia, Diapsida) in der Oberkreide Europas: Champsosaurierwirbel aus den Gosau-Schichten (Campan) Niederösterreichs. Sitzungsberichten der Österreichs Akademis der Wissenschaften Mathematisch-Naturwissenschaftlichen Klasse, Abteilung I 197:389-394 [R. Benson/R. Benson]
34546 E. Buffetaut, A. Osi, and E. Prondvai. 2011. The pterosaurian remains from the Grünbach Formation (Campanian, Gosau Group) of Austria: a reappraisal of 'Ornithocheirus buenzeli. Geological Magazine 148:334-339 [R. Butler/R. Butler/R. Butler]
12997ETE E. Bunzel. 1871. Die Reptilfauna der Gosauformation in der Neuen Welt bei Weiner-Neustadt. Abhandlungen der Kaiserlich-Königlichen Geologischen Reichsanstalt 5:1-18 [M. Carrano/M. Carrano/M. Carrano]
65036 A. Dinca, M. Tocorjescu, and A. Stilla. 1971. Despre vîrsta depozitelor continentale cu dinozaurieni din bazinele Hateg si Rusca Montana [On the age of continental deposits with dinosaurians from the Hateg and Rusca Montana basins]. Dari de Seama ale Sedintelor 58(4):83-94 [M. Carrano/M. Carrano]
46182 A. Gaudry. 1890. Les Enchainements du Monde Animal dans les Temps Geologiques. Fossiles Secondaires 1-322 [M. Carrano/M. Carrano]
62949 E. Hennig. 1915. In F. Frech (ed.), Fossilium Catalogus. I: Animalia. Pars 9: Stegosauria 1-16 [M. Carrano/M. Carrano]
32135 C.-M. Jianu, D. B. Weishampel, and E. Stiuca. 1997. Old and new pterosaur material from the Hateg Basin (Late Cretaceous) of western Romania, and comments about pterosaur diversity in the Late Cretaceous of Europe. 2nd European Workshop on Vertebrate Paleontology, Espéraza-Quillan, Abstracts. Musée des Dinosaures, Espéraza [R. Butler/R. Butler/R. Butler]
62124 W. G. Joyce. 2017. A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58(1):65-113 [P. Holroyd/P. Holroyd/P. Mannion]
7842 J. Le Loeuff and E. Buffetaut. 1991. Tarascosaurus salluvicus nov. gen., nov. sp., dinosaure théropode du Crétacé supérieur du sud de la France [Tarascosaurus salluvicus nov. gen., nov. sp., a theropod dinosaur from the Upper Cretaceous of southern France]. Géobios 25(5):585-594 [M. Carrano/M. Carrano/M. Carrano]
67934 J. Le Loeuff and E. Buffetaut. 1995. The evolution of Late Cretaceous non-marine vertebrate fauna in Europe. Sixth Symposium on Mesozoic Terrestrial Ecosystems and Biota, Short Papers 181-184 [M. Carrano/M. Carrano]
53030 B. F. Nopcsa. 1897. Vorläufiger Bericht über das Auftreten von oberer Kreide im Hátszeger Thale in Siebenbürgen [Preliminary report on the occurrence of the Upper Cretaceous in the Hátszeg Valley in Transylvania]. Verhandlungen der Kaiserlich-Königlichen Geologischen Reichsanstalt, Wien 1897:273-274 [M. Carrano/M. Carrano]
13876ETE B. F. Nopcsa. 1902. Notizen über cretacische Dinosaurier [Notes on Cretaceous dinosaurs]. Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften 111(1):93-114 [M. Carrano/M. Carrano/M. Carrano]
51605 B. F. Nopcsa. 1929. Dinosaurierreste aus Siebenbürgen V. [Dinosaur remains from Siebenbürgen V]. Geologica Hungarica, Series Palaeontologica 4:1-72 [M. Carrano/M. Carrano]
14501ETE D. B. Norman and D. B. Weishampel. 1990. Iguanodontidae and related ornithopods. In D. B. Weishampel, H. Osmolska, and P. Dodson (eds.), The Dinosauria. University of California Press, Berkeley 510-533 [M. Carrano/M. Carrano/M. Carrano]
43083 A. Osi, E. Prondvai, R. Butler and D. B. Weishampel. 2012. Phylogeny, histology and inferred body size evolution in a new rhabdodontid dinosaur from the Late Cretaceous of Hungary. PLoS ONE 7(9):e44318. [R. Benson/R. Benson]
17519ETE X. Pereda Suberbiola and P. M. Galton. 2001. Reappraisal of the nodosaurid ankylosaur Struthiosaurus austriacus Bunzel from the Upper Cretaceous Gosau Beds of Austria. In K. Carpenter (ed.), The Armored Dinosaurs. Indiana University Press, Bloomington 173-210 [M. Carrano/M. Carrano]
59665 M. Rabi, M. Vremir, and H. Tong. 2013. Preliminary Overview of Late Cretaceous Turtle Diversity in Eastern Central Europe (Austria, Hungary, and Romania). In D. B. Brinkman, P. A. Holroyd, and J. D. Gardner (eds.), Morphology and Evolution of Turtles 307-336 [P. Holroyd/P. Holroyd]
17757ETE S. Sachs and J. J. Hornung. 2006. Juvenile ornithopod (Dinosauria: Rhabdodontidae) remains from the Upper Cretaceous (lower Campanian, Gosau Group) of Muthmannsdorf (Lower Austria). Géobios 39:415-425 [M. Carrano/M. Carrano/M. Carrano]
12806ETE H. G. Seeley. 1881. The reptile fauna of the Gosau Formation preserved in the Geological Museum of the University of Vienna. Quarterly Journal of the Geological Society of London 37(148):620-707 [M. Carrano/M. Carrano/M. Carrano]
17355ETE E. Suess. 1881. Note on the Gosau Beds of the Neue Welt, west of Weiner Neustadt. Quarterly Journal of the Geological Society of London 37(148):702-703 [M. Carrano/M. Carrano]
42854 R. Tabuce, M. T. Antunes, R. Smith and T. Smith. 2006. Dental and tarsal morphology of the European Paleocene/Eocene “condylarth” mammal Microhyus. Acta Palaeontologica Polonica 51(1):37-52 [P. Mannion/P. Mannion/M. Carrano]
19380ETE D. B. Weishampel, D. Grigorescu, and D. B. Norman. 1991. The dinosaurs of Transylvania. National Geographic Research & Exploration 7(2):196-215 [M. Carrano/M. Carrano]
31157 P. Wellnhofer. 1980. Flugsauriereste aus der Gosau-Kreide von Muthmannsdorf (Niederösterreich) - ein Beitrag zur Kiefermechanik der Pterosaurier. Mitteilungen der Bayerischen Staatssammlung für Paläontologie und historische Geologie 20:95-112 [R. Butler/R. Butler]