PBDB

Chorlakki: Lutetian, Pakistan

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

unclassified

	Birbalomys ? vandermeuleni Hussain et al. 1978	Gingerich et al. 1983

	cf. Varohstichthys sp.	Gayet 1987
Reptilia - Testudines

	Chorlakkichelys shahi de Broin 1987	de Broin 1987
	nomen dubium belonging to Carettochelyidae
Reptilia - Testudines - Trionychidae

	Trionychinae indet. Gray 1825	de Broin 1987
Reptilia - Agamidae

	Tinosaurus sp. (Marsh 1872)	Rage 1987
	synonym of Saniwa
Reptilia - Boidae

	Erycinae indet. Bonaparte 1825	Rage 1987

	Boinae indet. Gray 1825	Rage 1987
Mammalia - Rodentia - Chapattimyidae

	Birbalomys ijlsti (Hussain et al. 1978)	Gingerich et al. 1983
	recombined as Basalomys ijlsti

	Chapattimys debruijni Hartenberger 1982	Gingerich et al. 1983

	Chapattimys wilsoni Hussain et al. 1978	Gingerich et al. 1983

	Saykanomys sp. Shevyreva 1972
	= Birbalomys sondaari Hussain et al. 1978	Hartenberger 1982

	Birbalomys sondaari (Hussain et al. 1978)	Gingerich et al. 1983

	Birbalomys ? woodi Sahni and Khare 1973	Gingerich et al. 1983

	Gumbatomys asifi Hartenberger 1982	Gingerich et al. 1983
Mammalia - Rodentia

	cf. Tamquammys sp. Shevyreva 1971
Mammalia - Rodentia - Yuomyidae

	cf. Petrokoslovia sp. (Shevyreva 1972)	Gingerich et al. 1983
	original and current combination Petrokozlovia

	cf. Petrokoslovia informal sp. indet. 2 (Shevyreva 1972)	Hartenberger 1982
	original and current combination Petrokozlovia
Mammalia - Primates - Omomyidae

	Kohatius coppensi n. gen., n. sp. Russell and Gingerich 1980	Russell and Gingerich 1980
Mammalia - Primates - Notharctidae

	cf. Agerinia sp. Crusafont-Pairó and Golpe-Posse 1973	Thewissen et al. 1987

	Panobius afridi n. gen., n. sp. Russell and Gingerich 1987	Thewissen et al. 1987

	Panobius amplior n. sp. Gunnell et al. 2008	Gunnell et al. 2008
	specimen referred to Adapidae indet. by Russell and Gingerich (1987) and to Panobius afridi by Thewissen et al. (2001)
Mammalia - Plesiosoricidae

	Pakilestes lathrius Russell and Gingerich 1981	Russell and Gingerich 1981
Mammalia

	Seia shahi Russell and Gingerich 1981	Russell and Gingerich 1981
Mammalia - Chiroptera

	Chiroptera indet. Blumenbach 1779	Gingerich et al. 1983
Mammalia - Creodonta

	Paratritemnodon indicus Ranga Rao 1973	Gingerich et al. 1983
Mammalia - Perissodactyla - Paraceratheriidae

	Forstercooperia jigniensis Sahni and Khare 1973	Thewissen et al. 1987
	nomen dubium belonging to Paraceratheriidae
Mammalia - Perissodactyla - Isectolophidae

	Isectolophidae indet. Peterson 1919	Thewissen et al. 1987
Mammalia - Perissodactyla - Brontotheriidae

	? Eotitanops dayi Dehm and Oettingen-Spielberg 1958	Gingerich et al. 1983
Mammalia - Perissodactyla - Anthracobunidae

	Anthracobune pinfoldi Pilgrim 1940

	Pilgrimella pilgrimi Dehm and Oettingen-Spielberg 1958	Gingerich et al. 1983
	synonym of Anthracobune pinfoldi

	Lammidhania wardi (Pilgrim 1940)	Gingerich et al. 1983
	original and current combination Anthracobune wardi
Mammalia - Dichobunidae

	Dichobunidae indet. Gill 1872	Thewissen et al. 1987

	Pakibune chorlakkiensis Thewissen et al. 1987	Thewissen et al. 1987

	Dulcidon gandaiensis	Gingerich et al. 1983

	Chorlakkia hassani n. gen., n. sp. Gingerich et al. 1979	Gingerich et al. 1979
Mammalia - Raoellidae

	Khirtharia dayi Pilgrim 1940

	Indohyus cf. indriae

	Indohyus major n. sp. Thewissen et al. 1987	Thewissen et al. 1987
Mammalia - Diacodexeidae

	Diacodexis pakistanensis Thewissen et al. 1983	Gingerich et al. 1983
	recombined as Gujaratia pakistanensis
Mammalia - Cetacea - Pakicetidae

	Pakicetus inachus n. gen., n. sp. Gingerich and Russell 1981	Gingerich and Russell 1981	8 specimens
	(19 measurements)

	Ichthyolestes pinfoldi Dehm and Oettingen-Spielberg 1958	Thewissen and Hussain 1998	7 specimens
Mammalia

	Basalina basalensis Dehm and Oettingen-Spielberg 1958	Gingerich et al. 1983
Actinopteri - Lepisosteiformes - Lepisosteidae

	Lepisosteus sp. Agassiz 1843	Gayet 1987
Actinopteri - Osteoglossiformes - Osteoglossidae

	Osteoglossidae indet. Bonaparte 1832	Gayet 1987
Actinopteri - Siluriformes - Clariidae

	Clariidae indet. Bonaparte 1846	Gayet 1987
Actinopteri - Siluriformes - Bagridae

	Bagridae indet. Bleeker 1852	Gayet 1987
Actinopteri - Siluriformes - Ariidae

	Ariidae indet. Berg 1958	Gayet 1987
Actinopteri - Cypriniformes - Cyprinidae

	Cyprinidae indet. Rafinesque 1815	Gayet 1987
Actinopteri - Cyprinodontiformes - Cyprinodontidae

	Cyprinodontidae indet.	Gayet 1987
Actinopteri - Anabantiformes

	Eochanna chorlakkiensis	Roe 1991
Actinopteri - Percichthyidae

	Macquaria antiquus Hills 1934	Gayet 1987
Gastropoda - Heterostropha - Planorbidae

	Planorbis sp. Muller 1774
see common names

Geography

Country:	Pakistan	State/province:	North-West Frontier
Coordinates:	33.6° North, 71.9° East (view map)
Paleocoordinates:	14.4° North, 68.1° East
Basis of coordinate:	stated in text
Geographic resolution:	outcrop

Time

Period:	Paleogene	Epoch:	Eocene
Stage:	Lutetian	10 m.y. bin:	Cenozoic 2
*Period:	Early/Lower - Middle Tertiary - Early/Lower - Middle Tertiary	*Epoch:	Early/Lower Eocene - Early/Lower Eocene
*International age/stage:	Late/Upper Ypresian - Late/Upper Ypresian
Key time interval:	Lutetian
Age range of interval:	47.80000 - 41.30000 m.y. ago
* legacy (obsolete) database fields

Stratigraphy

Formation:	Kuldana
Stratigraphic resolution:	group of beds
Stratigraphy comments: Gingerich (2003) places this collection in the earliest Lutetian shown by Gunnell et al. (2008) as being close to the top of the formation, spanning at three beds, and falling just over the Early - Middle Eocene boundary; early Lutetian

Lithology and environment

Primary lithology:	lithified calcareous conglomerate
Includes fossils?	Y
Lithology description: hard, calcareous conglomerate
Environment:	fluvial indet.

Taphonomy

Modes of preservation:	body,original phosphate
Size of fossils:	macrofossils
Preservation of anatomical detail:	good
Associated major elements:	some
Disassociated major elements:	some

Collection methods and comments

Collection methods:	bulk,selective quarrying,surface (in situ),chemical,mechanical,sieve
Rock censused:	200 kg
Reason for describing collection:	general faunal/floral analysis

Metadata

Database number:	28978
Authorizer:	M. Uhen, W. Clyde, J. Alroy	Enterer:	M. Uhen, J. Alroy, W. Clyde
Modifier:	P. Mannion	Research group:	vertebrate
Created:	2003-03-08 14:03:39	Last modified:	2016-07-22 04:24:13
Access level:	the public	Released:	2003-03-08 14:03:39
Creative Commons license:	CC BY

Reference information

Primary reference:

12009.

P. D. Gingerich, D. E. Russell, D. Sigogneau-Russell, J.-L. Hartenberger, S. M. Ibrahim Shah, M. Hassan, K. D. Rose and R. H. Ardrey. 1979. Reconnaissance survey and vertebrate paleontology of some Paleocene and Eocene Formations in Pakistan. Contributions from the Museum of Paleontology, University of Michigan 25(5):105-116 [M. Uhen/M. Uhen/M. Uhen]

Secondary references:

14333		F. de Broin. 1987. Lower vertebrates from the early-middle Eocene Kuldana Formation of Kohat (Pakistan): Chelonia. Contributions from the Museum of Paleontology, University of Michigan 27(7):169-185 [W. Clyde/W. Clyde/W. Clyde]
14329		M. Gayet. 1987. Lower vertebrates from the early-middle Eocene Kuldana Formation of Kohat (Pakistan): Holostei and Teleostei. Contributions from the Museum of Paleontology, University of Michigan 27(7):151-168 [W. Clyde/W. Clyde/W. Clyde]
15283		P. D. Gingerich. 2003. Stratigraphic and micropaleontological constraints on the Middle Eocene age of the mammal-bearing Kuldana Formation of Pakistan. Journal of Vertebrate Paleontology 23(3):643-651 [M. Uhen/M. Uhen]
13810		P. D. Gingerich, D. E. Russell, D. Sigogneau-Russell and J.-L. Hartenberger. 1979. Chorlakkia hassani, a new middle Eocene dichobunid (Mammalia, Artiodactyla) from the Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 25(6):117-124 [W. Clyde/W. Clyde/J. Alroy]
8326		P. D. Gingerich and D. E. Russell. 1981. Pakicetus inachus, a new archaeocete (Mammalia, Cetacea) from the early-middle Eocene Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 25(11):235-246 [M. Uhen/M. Uhen/M. Uhen]
12019		P. D. Gingerich, N. A. Wells, D. E. Russell and S. M. I. Shah. 1983. Origin of whales in epicontinental remnant seas: New evidence from the early Eocene of Pakistan. Science 220:403-406 [M. Uhen/M. Uhen/M. Uhen]
29569		G. F. Gunnell, P. D. Gingerich, M. Haq, J. I. Bloch, I. H. Khan and W. C. Clyde. 2008. New primates (Mammalia) from the early and middle Eocene of Pakistan and their paleobiogeographical implications. Contributions from the Museum of Paleontology, University of Michigan 32(1):1-14 [J. Alroy/J. Alroy]
13077		J.L. Hartenberger. 1982. A review of the Eocene rodents of Pakistan. Contributions from the Museum of Paleontology, University of Michigan 26:19-35 [W. Clyde/L. Gannon/J. Alroy]
14334		J. C. Rage. 1987. Lower vertebrates from the early-middle Eocene Kuldana Formation of Kohat (Pakistan): Squamata. Contributions from the Museum of Paleontology, University of Michigan 27(7):187-193 [W. Clyde/W. Clyde/W. Clyde]
13802		L. J. Roe. 1991. Phylogenetic and ecological significance of Channidae (Osteichthyes, Teleostei) from the early Eocene Kuldana Formation of Kohat, Pakistan. Contributions from the Museum of Paleontology, University of Michigan 28(5):93-100 [W. Clyde/W. Clyde/W. Clyde]
29559		D. E. Russell and P. D. Gingerich. 1980. Un nouveau Primate omomyide dans l'Eocène du Pakistan. Comptes Rendus de l'Académie de Paris, Paris, Série D 291:621-624 [J. Alroy/J. Alroy]
13809		D. E. Russell and P. D. Gingerich. 1981. Lipotyphla, Proteutheria(?), and Chiroptera (Mammalia) from the early-middle Eocene Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 25(14):277-287 [W. Clyde/W. Clyde/W. Clyde]
59921		D. E. Russell and P. D. Gingerich. 1987. Nouveaux primates de l’Éocène du Pakistan. Comptes Rendus de l'Academie des Sciences, Paris, Série II 5:209-214 [P. Mannion/P. Mannion]
14336		J. G. M. Thewissen, P. D. Gingerich, and D. E. Russell. 1987. Artiodactyla and Perissodactyla (Mammalia) from the early-middle Eocene Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 27(11):247-274 [W. Clyde/W. Clyde/M. Uhen]
11990		J. G. M. Thewissen and S. T. Hussain. 1998. Systematic review of the Pakicetidae, early and middle Eocene Cetacea (Mammalia) from Pakistan and India. Bulletin of Carnegie Museum of Natural History 34:220-238 [M. Uhen/M. Uhen/M. Uhen]