Bèni Mellal: Astaracian, Morocco

List of taxa
Where & when
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Squamata - Lacertidae
cf. Eremias sp. Wiegmann 1834
Reptilia - Squamata - Anguidae
Anguidae indet. Gray 1825
Reptilia - Squamata - Pythonidae
Python maurus n. sp. Rage 1976
Reptilia - Squamata - Boidae
Erycinae indet. Bonaparte 1825
invalid subgroup of Boidae
"either Eryx sp. or Gongylophis sp."
Reptilia - Squamata - Elapidae
Naja antiqua n. sp. Rage 1976
Reptilia - Squamata - Colubridae
Colubridae informal indet. 1 Oppel 1811
Colubridae informal indet. 2 Oppel 1811
Reptilia - Squamata - Crotalidae
Vipera maghrebiana n. sp. Rage 1976
Reptilia - Squamata
Scolecophidia indet. Duméril and Bibron 1844
Reptilia - Squamata - Gekkonidae
Gekkonidae informal indet. 1 Gray 1825
Gekkonidae informal indet. 2 Gray 1825
Tarentola sp. Gray 1825
Daza et al. 2014
Large form known from BML 915, frontal; BML 914, a dentary; fragments of maxilla and some vertebrae are also present. A dentary and vertebrae represent the smallest form.
Reptilia - Squamata - Amphisbaenidae
Amphisbaenidae indet. Gray 1865
Reptilia - Squamata - Scincidae
Eumeces sp. Wiegmann 1834
Mammalia - Rodentia - Cricetidae
Cricetodon sp. Lartet 1851
Arambourg 1963
Mammalia - Carnivora - Amphicyonidae
Agnotherium cf. antiquum Kaup 1833
Ginsburg 1977
Mammalia - Carnivora - Mustelidae
Mellalictis mellalensis n. gen., n. sp. Ginsburg 1977
Ginsburg 1977
Martes khelifensis n. sp. Ginsburg 1977
Ginsburg 1977
Mammalia - Carnivora - Felidae
Felis sp. Linnaeus 1758
Ginsburg 1977
new species
Mammalia - Carnivora - Viverridae
Genetta sp. Cuvier 1816
Ginsburg 1977
Mammalia - Carnivora - Hyaenidae
Hyaenictis graeca
Ginsburg 1977
Ictitherium cf. arambourgi
Ginsburg 1977
Mammalia - Hyracoidea
Hyracoidea indet. Huxley 1869
Arambourg 1963
see common names

Country:Morocco State/province:Tadla-Azilal
Coordinates: 32.3° North, 6.3° West (view map)
Paleocoordinates:32.1° North, 7.0° West
Basis of coordinate:estimated from map
Period:Neogene Epoch:Miocene
10 m.y. bin:Cenozoic 5
Key time interval:Astaracian
Age range of interval:15.97000 - 11.60800 m.y. ago
Lithology and environment
Primary lithology: not reported
Environment:terrestrial indet.
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:mechanical,survey of museum collection
Reason for describing collection:taxonomic analysis
Taxonomic list comments:Taxa Palaeonaja sp. and Ophisaurus sp. (entered as "Ophiosaurus sp."), originally included in the taxonomic list, have been removed since they are not reported by Rage (1976, ref no. 12976) from Bèni Mellal. Instead, according to Rage (1976, ref no. 12977), these taxa occur in Çalta (Turkey) and are already present in the taxonomic list of that collection (see PBDB coll. no. 34370). (TL)
Database number:48630
Authorizer:J. Head, J. Mueller, P. Barrett, M. Uhen Enterer:J. Head, T. Cleary, T. Liebrecht, M. Uhen
Modifier:T. Liebrecht Research group:vertebrate
Created:2005-03-24 19:14:58 Last modified:2015-04-13 17:10:41
Access level:the public Released:2005-03-24 19:14:58
Creative Commons license:CC BY
Reference information

Primary reference:

12976. J.-C. Rage. 1976. Les Squamates du Miocène de Bèni Mellal, Maroc. Géologie Méditerranéenne 3(2):57-70 [J. Head/J. Head/J. Head]

Secondary references:

43778 C. Arambourg. 1963. Continental vertebrate faunas of the Tertiary of North Africa. In F. C. Howell, F. Bourliere (eds.), African Ecology and Human Evolution 55-64 [M. Uhen/M. Uhen]
59595 J. D. Daza, A. M. Bauer, and E. D. Snively. 2014. On the fossil record of the Gekkota. The Anatomical Record 297:433-462 [P. Barrett/T. Cleary]
16909 L. Ginsburg. 1977. Les carnivores du Miocene de Beni Mellal (Maroc). Geologie Mediterraneene 4(3):225-240 [M. Uhen/M. Uhen]