Nikiti-2: MN 11, Greece

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Testudines - Testudinidae
Testudo cf. marmorum Gaudry 1862
Garcia et al. 2016
complete shell
    = Testudo marmorum Gaudry 1862
Vlachos et al. 2019
Reptilia - Theropoda - Struthionidae
Struthio cf. karatheodoris Forsyth Major 1888
Koufos et al. 2016
tarsometatarsus
Mammalia - Proboscidea - Choerolophodontidae
Choerolophodon pentelici (Gaudry and Lartet 1856)
Mammalia - Perissodactyla - Equidae
Hipparion sp. de Christol 1832
"large-sized"
Hipparion dietrichi (Wehrli 1941)
Hipparion macedonicum Koufos 1984
Mammalia - Giraffidae
Palaeotragus rouenii (Gaudry 1861)
Kostopoulos 2016
Left P2-(part)M1, NIK-1823; distal part of humerus, NIK-1810; radius of young individual, NIK-1809; metacarpal III + IV, NIK-1807; calcaneum, NIK-1912; part of cuboscaphoid, NIK-1914; proximal metatarsal III + IV, NIK-1808; first phalanx, NIK-1913; first, second and third phalanges, NIK- 1915.
Helladotherium duvernoyi Gaudry and Lartet 1856
"Helledotherium duvernoyi"
Palaeotraginae indet. Pilgrim 1911
Kostopoulos 2016
Part of first phalanx lacking proximal epiphysis, NIK-211; proximal epiphysis of first phalanx, NIK-264 (young); second phalanges of young individuals, NIK-1916 (posterior?), NIK-1917 (anterior?).
Mammalia - Bovidae
Gazella sp. Blaineville 1816
    = Gazella cf. capricornis
Kostopoulos 2016
Partially preserved skull, NIK-1812; partially preserved skull with mandible, NIK-1142; frontlet, NIK-699; right horn-core, NIK-1819; basal part of left horn-core, NIK-1820; P2-M3, NIK-607, 1845. Possibly ascribed: right mandibular ramus with p4-m3, NIK-1892; left mandibular ramus with p3-m3, NIK- 1889, 1896; right mandibular ramus with p4-m3, NIK-1897.
Gazella pilgrimi Bohlin 1935
Kostopoulos 2016
Partially preserved skull, NIK-1811; frontlets, NIK-813, 1813, 1814, 1817; left horn-core, NIK-700, 1815; right horn-core, NIK-1816, 1818; palate, NIK-1899; right P2-M3, NIK-607; part of mandible with i1-m3 sin. and i1, p2-m3 dex., NIK- 778; right mandibular ramii, NIK-589, 1888, 1890, 1898, 1902; left mandibular ramii, NIK-13’, 590, 1891, 1893-1895, 1900, 1901.
Palaeoryx cf. pallasi (Wagner 1857)
Kostopoulos 2016
left maxilla with P2-M3, NIK-1063.
Nisidorcas planicornis (Pilgrim 1939)
Tragoportax amalthea (Roth and Wagner 1854)
Kostopoulos 2016
Skull, NIK-1688; partial skull, NIK-1687; frontlets of young individuals, NIK-1135, 1689; partly preserved hornless skull, NIK-1684; palate, NIK-423, 1685; right P2-M3, NIK- 1141; mandible, NIK-1181; left part of mandible, NIK-1675; lower tooth rows, NIK-425, 433, 1676, 1691.
cf. Ouzocerus sp.
    = Palaeoreas lindermayeri Wagner 1848
Kostopoulos 2016
Tragoportax aff. rugosifrons (Schlosser 1904)
"Tragoportax aff. rugosiforns"
    = ? Miotragocerus sp. Stromer 1928
Kostopoulos 2016
Partial skull with associate mandible, NIK- 764; anterior part of skull with palate, NIK-422, 1690; lower dentitions, NIK-1103, 1140, 1680, 1681.
see common names

Geography
Country:Greece
Coordinates: 40.2° North, 23.7° East (view map)
Paleocoordinates:39.9° North, 23.4° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period: Neogene Epoch: Miocene
Stage: Tortonian 10 m.y. bin: Cenozoic 6
Key time interval: MN 11
Age range of interval: 8.7 - 7.75 m.y. ago
Stratigraphy
Formation:Nikiti
Stratigraphic resolution:bed
Stratigraphy comments: about 20 m higher than the Vallesian collection Nikiti 1 (NKT)
"early Turolian... beginning of MN11"
Lithology and environment
Primary lithology:hematitic siltstone
Lithology description: lens of "red beds"; stratigraphic section indicates fine sandstone mixed with clay, presumably meaning siltstone
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Associated major elements:some
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:field collection
Reason for describing collection:taxonomic analysis
Metadata
Also known as:NIK
Database number:73869
Authorizer:J. Alroy, E. Vlachos Enterer:J. Alroy, P. Kampouridis, E. Vlachos
Modifier:J. Alroy Research group:vertebrate
Created:2007-07-09 11:21:58 Last modified:2007-07-09 14:58:45
Access level:the public Released:2007-07-09 11:21:58
Creative Commons license:CC BY
Reference information

Primary reference:

24807. T. Vlachou and G. D. Koufos. 2002. The hipparions (Mammalia, Perissodactyla) from the Turolian locality "Nikiti-2" (NIK), Macedonia, N. Greece. Annales de Paléontologie 88(4):215-263 [J. Alroy/J. Alroy]

Secondary references:

61581 G. Garcia, D. S. Kostopoulos, X. Valentin and G. D. Koufos. 2016. Testudines. Geobios 49:23-27 [E. Vlachos/E. Vlachos]
71311 D. S. Kostopoulos. 2016. Artiodactyla - Palaeontology of the upper Miocene vertebrate localities of Nikiti (Chalkidiki Peninsula, Macedonia, Greece). Geobios 49:119-234 [E. Vlachos/E. Vlachos/E. Vlachos]
67329 G. D. Koufos, D. S. Kostopoulos, and G. E. Konidaris. 2016. Palaeontology of the upper Miocene vertebrate localities of Nikiti (Chalkidiki Peninsula, Macedonia, Greece) Aves. Geobios 49:29-36 [E. Vlachos/P. Kampouridis]
72005 E. Vlachos, G. L. Georgalis, S. Roussiakis, M. Bohme, and G. Theodorou. 2019. The Pikermian tortoises (Testudines, Testudinidae) from the late Miocene of the South Balkans. Journal of Vertebrate Paleontology 39(5):1-23 [E. Vlachos/E. Vlachos]