PBDB Collection

Andrès (MNHN-UL): Late/Upper Kimmeridgian - Early/Lower Tithonian, Portugal
collected by J. Amorim 1988, 2005

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

unclassified

	Plantae indet. Haeckel 1866	Malafaia et al. 2006
	carbonized plant remains
Actinopteri - Semionotiformes

	Semionotiformes indet. Arambourg and Bertin 1958	Malafaia et al. 2006
Actinopteri - Lepisosteiformes

	Lepidotes sp. Agassiz 1832	Malafaia et al. 2006	1 specimen
	nearly complete individual
Reptilia

	Lepidosauria indet. (Haeckel 1866)	Malafaia et al. 2006
	Several individuals, incl. at least one partially complete skeleton
Reptilia - Sphenodontia

	Sphenodontia indet. (Jaekel 1910)	Ortega et al. 2006
	Close relative of Opisthias (ref 25000)
Reptilia - Dromaeosauridae

	Dromaeosauridae indet. Colbert and Russell 1969	Malafaia et al. 2006
	teeth
Reptilia

	Allosaurus fragilis Marsh 1877		2 individuals
	MHNUL/AND.001
	= Allosaurus europaeus Mateus et al. 2006	Mateus et al. 2006

	Ornithopoda indet. Marsh 1881	Malafaia et al. 2006
Reptilia - Camptosauridae

	Camptosauridae indet. Marsh 1885	Malafaia et al. 2006
	teeth
Reptilia - Dryosauridae

	Dryosauridae indet. Milner and Norman 1984	Malafaia et al. 2006
	= aff. Dryosaurus sp. Marsh 1894	Malafaia et al. 2010
Reptilia

	Camarasaurus sp. Cope 1877	Malafaia et al. 2006
	= Neosauropoda indet. Bonaparte 1986	Malafaia et al. 2010

	Diplodocoidea indet. Upchurch 1995	Malafaia et al. 2006	3 specimens
	MNHNUL.P.AN D302, D303, D304 (teeth)
	= Diplodocidae indet. Marsh 1884	Malafaia et al. 2010

	Titanosauriformes indet. Salgado et al. 1997	Malafaia et al. 2006

	? Turiasauria indet. Royo-Torres et al. 2006	Mocho et al. 2017
Reptilia - Rhamphorhynchidae

	Pterosauria indet. Kaup 1834	Malafaia et al. 2006
	= cf. Rhamphorhynchus sp. Meyer 1847	Malafaia et al. 2010
Reptilia

	Theriosuchus sp. Owen 1878	Malafaia et al. 2006
Reptilia - Goniopholididae

	Goniopholis sp. Owen 1842	Malafaia et al. 2006
Amphibia

	Amphibia indet. Linnaeus 1758	Malafaia et al. 2006
Gastropoda

	Gastropoda indet. Cuvier 1795	Malafaia et al. 2006
Bivalvia

	Bivalvia indet. Linnaeus 1758	Malafaia et al. 2006
see common names

Geography

Country:	Portugal	State/province:	Centro	County:	Leiria
Coordinates:	39.9° North, 8.6° West (view map)
Paleocoordinates:	33.8° North, 2.0° East
Basis of coordinate:	based on nearby landmark
Altitude:	212 meters
Geographic resolution:	small collection

Time

Period:	Jurassic	Epoch:	Late/Upper Jurassic

Key time interval:	Late/Upper Kimmeridgian - Early/Lower Tithonian
Age range of interval:	152.20000 - 146.30000 m.y. ago

Stratigraphy

Formation:	Lourinhã
Stratigraphic resolution:	bed
Stratigraphy comments: = Camadas de Alcobaça, could also be from lowermost Bombarral Formation (= Grès superiores); Malafaia et al 2010 consider the site to be ?upper Kimmeridgian to Tithonian in age Previously assigned to Bombarral Formation. Lourinhã Fm is considered late Kimmeridgian-late Tithonian and it incorporates the Alcobaça Fm (Kimmeridgian) and the Bombarral Fm (also known as Grés Superiores; late Tithonian-early Berrisian). Taylor et al. 2013 suggested the Lourinhã Fm be subdivided into members, but Mateus et al. 2017 proposed a revised stratiraphy of the Luisitanian Basin. The Alcobaça Formation is Late Kimmeridigan and overlain by the Lourinha Formation. The Lourinhã Formation is made up of the Praia da Amoreira and Porto Novo (late Kimmeridgian) members, Praia Azul member (latest Kimmeridgian–earliest Tithonian) and the Assenta member (late Tithonian-earliest Berrisian). The Lourinha Formation is overlain by the Porto da Calada Formation (Berrisian).

Lithology and environment

Primary lithology:	paleosol/pedogenic,micaceous,gray carbonaceous sandstone
Secondary lithology:	fine siltstone
Includes fossils?	Y
Lithology description: "mainly represented by micaceous sandstones (as occur in Andrès), fine sands and silts", also described as "sandstones and marls that represent paleosols", also described as "fine-grained mud-sandstones, sometimes micaceous, with intercalations of some levels of marls, silts and clays, sometimes with abundant calcareous or limonitic nodules"
Environment:	fluvial indet.
Geology comments: "deposited in a fluvial meandering environment with subsidiary deltaic and marginal marine environments.", also called a shallow, distal river setting

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils,mesofossils
Spatial orientation:	life position
Preservation of anatomical detail:	good
Articulated whole bodies:	none
Associated major elements:	many
Disassociated major elements:	many
Disassociated minor elements:	many
Fragmentation:	occasional
Bioerosion:	none
Spatial resolution:	parautochthonous

Collection methods and comments

Collection methods:	selective quarrying,mechanical,field collection
Reason for describing collection:	taxonomic analysis
Collectors:	J. Amorim	Collection dates:	1988, 2005

Metadata

Also known as:	Andrés. Andres, Pombal
Database number:	38680
Authorizer:	M. Carrano, P. Mannion	Enterer:	M. Carrano, J. Tennant
Modifier:	M. Carrano	Research group:	vertebrate
Created:	2004-04-28 10:46:44	Last modified:	2023-11-17 15:44:03
Access level:	the public	Released:	2004-04-28 10:46:44
Creative Commons license:	CC BY

Reference information

Primary reference:

10381.

B. P. Pérez-Moreno, D. J. Chure, C. Pires, C. Marques Silva, V. Dos Santos, P. Dantas, L. Póvoas, M. Cachão, J. L. Sanz and A. M. Galopim Carvalho. 1999. On the presence of Allosaurus fragilis (Theropoda: Carnosauria) in the Upper Jurassic of Portugal: first evidence of an intercontinental dinosaur species. Journal of the Geological Society, London 156:449-452 [M. Carrano/M. Carrano/M. Carrano]

Secondary references:

13166	ETE	J. F. Bonaparte and O. Mateus. 1999. A new diplodocid, Dinheirosaurus lourinhanensis gen. et sp. nov., from the Late Jurassic beds of Portugal. Revista del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigación de las Ciencias Naturales, Paleontología 5(2):13-29 [M. Carrano/M. Carrano/M. Carrano]
25000	ETE	E. Malafaia, P. Dantas, F. Ortega, F. Escaso, J. M. Gasulla, B. Ribeiro, F. Barriga, I. Gromicho, M. García-Oliva, G. Ramalheiro, J. Santamaría, N. L. Pimentel, C. Moniz and A. M. G. Carvalho. 2006. Análisis preliminar de la diversidad faunística en el yacimiento de Andrés (Jurásico Superior. Pombal, Portugal) [Preliminary analysis of faunal diversity in the Andrés locality (Upper Jurassic. Pombal, Portugal)]. IV Encuentro de Jóvenes Investigadores en Paleontología, Salamanca. Libro de Resúmenes 91-92 [M. Carrano/M. Carrano/M. Carrano]
24989	ETE	E. Malafaia, P. Dantas, F. Ortega and F. Escaso. 2007. Nuevos restos de Allosaurus fragilis (Theropoda: Carnosauria) del yacimiento de Andrés (Jurásico Superior; centro-oeste de Portugal) [New remains of Allosaurus fragilis (Theropoda: Carnosauria) from the Andres locality (Upper Jurassic; west-central Portugal)]. In Cantera Paleontológica, Diputación Provincial de Cuenca, Cuenca, Spain 255-271 [M. Carrano/M. Carrano]
44312		E. Malafaia, F. Ortega, F. Escaso, P. Dantas, N. Pimentel, J. M. Gasulla, B. Ribeiro, F. Barriga, and J. L. Sanz. 2010. Vertebrate fauna at the Allosaurus fossil-site of Andres (Upper Jurassic), Pombal, Portugal. Journal of Iberian Geology 36(2):193-204 [P. Mannion/J. Tennant/M. Carrano]
42539		P. D. Mannion, P. Upchurch, O. Mateus, R. N. Barnes, and M. E. H. Jones. 2012. New information on the anatomy and systematic position of Dinheirosaurus lourinhanensis (Sauropoda: Diplodocoidea) from the Late Jurassic of Portugal, with a review of European diplodocoids. Journal of Systematic Palaeontology 10(3):521-551 [P. Mannion/P. Mannion/M. Carrano]
79466		O. Mateus, J. Dinis, and P. P. Cunha. 2017. The Lourinhã Formation: the Upper Jurassic to lowermost Cretaceous of the Lusitanian Basin, Portugal – landscapes where dinosaurs walked. Ciências da Terra Earth Sciences Journal 19(1):75-97 [P. Mannion/G. Varnham/G. Varnham]
19890	ETE	O. Mateus, A. Walen, and M. T. Antunes. 2006. The large theropod fauna of the Lourinhã Formation (Portugal) and its similarity to that of the Morrison Formation, with a description of a new species of Allosaurus. In J. R. Foster & S. G. Lucas (ed.), Paleontology and Geology of the Upper Jurassic Morrison Formation. New Mexico Museum of Natural History and Science Bulletin 36:123-129 [M. Carrano/M. Carrano/M. Carrano]
76192		P. Mocho, R. Royo-Torres, F. Escaso, E. Malafaia, C. M. Chaves, I. Narváez, A. Pérez-García, N. Pimentel, B. C. Silva and F. Ortega. 2017. Upper Jurassic sauropod record in the Lusitanian Basin (Portugal): geographical and lithostratigraphical distribution. Palaeontologia Electronica 20(2):27A:1-50 [M. Carrano/M. Carrano]
75880		F. Ortega, P. Dantas, F. Escaso, J. M. Gasulla, E. Malafaia and B. Ribeiro. 2006. Primera cita de reptiles esfenodontos en el Jurásico Superior de la Península Ibérica [First notice of sphenodontian reptiles in the Upper Jurassic of the Iberian Peninsula]. In E. Fernández-Martínez (ed.), Libro de Resúmenes de las XXII Jornadas de la Sociedad Española de Paleontología 152-153 [M. Carrano/M. Carrano]
86510		F. Ortega, E. Malafaia, F. Escaso, A. Pérez García, and P. Dantas. 2009. Faunas de répteis do Jurássico Superior de Portugal: sobre las faunas de reptiles del Jurásico Superior de Portugal [Reptile faunas from the Upper Jurassic of Portugal: on the reptile faunas from the Upper Jurassic of Portugal]. Paleolusitana 1:43-56 [M. Carrano/M. Carrano]
70303		J. L. Sanz. 2000. Yacimientos ibéricos de dinosaurios [Iberian dinosaur localities]. Dinosaurios: Los Señores del Pasado [Dinosaurs: The Lords of the Land] 267-299 [M. Carrano/M. Carrano]