Sud Sacaco, Montemar: Messinian, Peru

List of taxa
Where & when
Taphonomy & methods
Metadata & references
Taxonomic list
Chondrichthyes - Odontaspidida - Carcharodontidae
Isurus hastalis (Agassiz 1843)
recombined as Cosmopolitodus hastalis
Chondrichthyes - Myliobatiformes - Myliobatidae
Myliobatis sp. Cuvier 1816
Muizon and DeVries 1985
Chondrichthyes - Lamniformes - Otodontidae
Procarcharodon megalodon (Agassiz 1843)
Muizon and DeVries 1985
recombined as Otodus megalodon
Also studied in Pimiento and Balk
Actinopteri - Clupeiformes - Clupeoidei
Alosinae indet.
Muizon and DeVries 1985
Reptilia - Avetheropoda - Scolopacidae
Tringinae indet. Gray 1840
Cheneval 1993
Reptilia - Avetheropoda - Cathartidae
Perugyps diazi n. gen., n. sp. Stucchi and Emslie 2005
Stucchi and Emslie 2005
Reptilia - Avetheropoda - Pelagornithidae
Pelagornithidae indet. Fürbringer 1888
Chavez et al. 2007 1 specimen
Reptilia - Avetheropoda - Spheniscidae
Spheniscidae indet. Bonaparte 1831
Muizon and DeVries 1985
form 2
Spheniscus urbinai Stucchi 2002
Stucchi 2002 1 specimen
Spheniscus megaramphus n. sp. Stucchi et al. 2003
Stucchi et al. 2003 5 specimens
Reptilia - Avetheropoda - Phalacrocoracidae
cf. Phalacrocorax sp. Brisson 1760
Muizon and DeVries 1985
Reptilia - Avetheropoda - Sulidae
cf. Sula sp. Brisson 1760
Muizon and DeVries 1985
Sula magna n. sp. Stucchi 2003
Stucchi 2003 1 specimen
Sula aff. variegata (Tschudi 1843)
Stucchi 2003 7 specimens
Sula sulita n. sp. Stucchi 2003
Stucchi 2003 1 specimen
Morus peruvianus n. sp. Stucchi 2003
Stucchi 2003 1 specimen
Eusuchia indet. Huxley 1875
Muizon and DeVries 1985
Piscogavialis jugaliperforatus n. gen., n. sp. Kraus 1998
Kraus 1998
SMNK 1282 PAL - holotype (skull, mandible and postcranial material)
Mammalia - Carnivora
Carnivora indet. Bowditch 1821
Muizon and DeVries 1985
Mammalia - Carnivora - Phocidae
Phocidae indet. Gray 1821
Monachinae indet. Trouessart 1897
Muizon and DeVries 1985
form 2
Acrophoca sp. Muizon 1981
Muizon and DeVries 1985
Mammalia - Megatheriidae
Megatheriidae indet. Gill 1872
Thalassocnus natans n. gen., n. sp. Muizon and McDonald 1995
Muizon and McDonald 1995 1 specimen
Muizon & McDonald 1995 stated this taxon was from the SAS horizon, but this was later changed to the MTM horizon in later publications (McDonald & Muizon, 2002)
Mammalia - Ungulata - Balaenopteridae
Balaenopteridae indet. Gray 1864
Muizon and DeVries 1985
form 2
Mammalia - Ungulata - Cetotheriidae
Cetotheriidae indet. (Brandt 1872)
Mammalia - Ungulata - Physeteridae
Physeteridae indet. Gray 1821
Muizon 1988
form 2
Mammalia - Ungulata
Acrophyseter deinodon n. gen., n. sp. Lambert et al. 2008
Lambert et al. 2017 1 specimen
(1 measurement)
Mammalia - Ungulata - Phocoenidae
Phocoenidae indet. (Gray 1825)
cf. Piscolithax sp. Muizon 1983
Muizon and DeVries 1985
see common names

Coordinates: 15.6° South, 74.7° West (view map)
Paleocoordinates:15.8° South, 73.4° West
Basis of coordinate:estimated from map
Period:Neogene Epoch:Miocene
Stage:Messinian 10 m.y. bin:Cenozoic 6
Key time interval:Messinian
Age range of interval:7.24600 - 5.33300 m.y. ago
Local section:Pisco Local bed:MTM
Local order:no particular order
Stratigraphic resolution:group of beds
Stratigraphy comments: 4.5-6 Ma (Stucchi & Urbina, 2004); 7.3 - 7.0 Ma (Ehret et al 2012)
Lithology and environment
Primary lithology:diatomaceous siltstone
Includes fossils?Y
Environment:paralic indet.
Geology comments: "a littoral paleoenviroment with protected beaches and reefs exposed to marine sediments" (Stucchi and Emslie 2005), high sedimentation rate, shallow-to-deep shelf, upwelling
Modes of preservation:body,original phosphate
Size of fossils:macrofossils
Preservation of anatomical detail:excellent
Collection methods and comments
Collection methods:surface (in situ),field collection
Reason for describing collection:taxonomic analysis
Also known as:MTM
Database number:46084
Authorizer:M. Uhen, J. Alroy Enterer:M. Uhen, J. Alroy
Modifier:B. Shipps Research group:vertebrate
Created:2005-01-01 15:27:52 Last modified:2017-02-26 16:00:19
Access level:the public Released:2005-01-01 15:27:52
Creative Commons license:CC BY
Reference information

Primary reference:

12182. R. Marocco and C. d.e. Muizon. 1988. Los vertebrados del Neogeno de La Costa Sur del Perú: Ambiente sedimentario y condiciones de fosilización. Bulletin de l'Institut Frances d'Etudes Andines 17(2):105-117 [M. Uhen/M. Uhen/M. Uhen]

Secondary references:

34370 M. Chavez, M. Stucchi, and M. Urbina. 2007. El registro de Pelagornithidae (Aves: Pelecaniformes) y la avifauna neogena del Pacifico sudeste. Bulletin de l'Institut Francais d'Etudes Andines 36(2):175-197 [M. Uhen/M. Uhen]
40374 J. Cheneval. 1993. L'avifaune Mio-Pliocene de la Formation Pisco (Perou) etude prliminaire. Document Labratorie Geologique Lyon 125:85-95 [M. Uhen/M. Uhen]
28038 R. Kraus. 1998. The cranium of Piscogavialis jugalipeforatus n. nen., n. sp. (Gavialidae, Crocodylia) from the Miocene of Peru. Palaeontologische Zeischrift 72(3/4):389-406 [M. Uhen/M. Uhen]
60319 O. Lambert, G. Bianucci, and C. Muizon. 2017. Macroraptorial sperm whales (Cetacea, Odontoceti, Physeteroidea) from the Miocene of Peru. Zoological Journal of the Linnean Society 179:404-474 [M. Uhen/M. Uhen/M. Uhen]
11895 C. Muizon. 1988. Les vertebres fossiles de la Formation Pisco (Perou). Troisieme partie: Les Odontocetes (Cetacea, Mammalia) du Miocene. Editions Recherche sur les Civilisations (78)1-244 [M. Uhen/N. Pyenson/M. Uhen]
16797 C. Muizon and T. J. DeVries. 1985. Geology and paleontology of late Cenozoic marine deposits in the Sacaco area (Peru). Geologische Rundschau 74(3):547-563 [M. Uhen/M. Uhen/M. Uhen]
16788 C. Muizon and H. G. McDonald. 1995. An aquatic sloth from the Pliocene of Peru. Nature 375:224-227 [M. Uhen/M. Uhen]
49081 C. Pimiento and M. Balk. 2015. The extinct giant shark Carcharocles megalodon: A model for understanding deep-time body size trends of marine apex predators. [C. Jaramillo/C. Pimiento/C. Pimiento]
34468 M. Stucchi. 2002. Una nueva especie de Spheniscus (Aves: Spheniscidae) de la Formacion Pisco, Peru. Boletim de la Sociedad Geologica del Peru 94:17-24 [M. Uhen/M. Uhen/M. Uhen]
54566 M. Stucchi. 2003. Los Piqueros (Aves: Sulidae) de la Formacion Pisco, Peru. Boletin de la Sociedad Geologica de Peru 95:75-91 [M. Uhen/M. Uhen]
29278 M. Stucchi and S. D. Emslie. 2005. A new condor (Ciconiiformes, Vulturidae) from the late Miocene/early Pliocene Pisco Formation, Peru. Condor 107(1):107-113 [J. Alroy/J. Alroy]
34433 M. Stucchi, M. Urbina, and A. Giraldo. 2003. Una nueva especie de Spheniscidae del Mioceno Tardio de la Formacion Pisco, Peru. Bulletin de la Institute Frances etudes Andines 32(2):361-375 [M. Uhen/M. Uhen]
18990 M. Stucchi and M. Urbina. 2004. Ramphastosula (aves, Sulidae): a new genus from the early Pliocene of the Pisco Formation, Peru. Journal of Vertebrate Paleontology 24(4):974-978 [M. Uhen/M. Uhen]